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In the 1980s, Welker voiced many recurring characters in the original Transformers animated series. He voiced several Decepticons, including the leader Megatron, Soundwave, Skywarp, Mixmaster, Laserbeak, Buzzsaw, Rumble, Frenzy, Ravage and Ratbat, as well as Autobots Mirage, Trailbreaker and Sludge.
Other actors have played Optimus Prime in various iterations of Transformers over the years, but Cullen happily returned to the role for the live-action film franchise. Cullen got his big break as the announcer on The Sonny & Cher Comedy Hour in the early 1970s, and famously voiced Eeyore in Winnie the Pooh.