the division of myrtaceae into leptospermoideae and myrtoideae was challenged by a number of authors, including johnson and briggs (1984), who identified 14 tribes or clades within myrtaceae, and found myrtoideae to be polyphyletic. molecular studies by several groups of authors, as of 2008, have confirmed the baccate (fleshy) fruits evolved twice from capsular fruits and, as such, the two-subfamily classification does not accurately portray the phylogenetic history of the family. thus, many workers are now using a recent analysis by wilson et al. (2001) as a starting point for further analyses of the family. this study pronounced both leptospermoideae and myrtoideae invalid, but retained several smaller suballiances shown to be monophyletic through matk analysis.