unusually for sonata form, the development section begins with a quiet restatement of the opening melody in the subdominant (e minor), a tonality usually reserved for near the end of a sonata form movement somewhere in the recapitulation or coda, and rises to a prolonged climax in the same key, starting with a dramatic variant of the opening melody in the full orchestra with prominent trombones. the expected relative major (d) of the tonic minor first appears only at the end of that climax, and then again for the second subject of the recap (in place of the expected tonic b major)—instead of much earlier, in the second subject of the exposition, as customary. the flutes and oboes then resume their melodic role at the end of that dramatic outburst, transitioning to the recapitulation.