the tarsometatarsus had a small hypotarsus with a single simple crest and devoid of foramina for tendons. the ankle joint consists of a sizable midward and an even larger outward groove. the conspicuous ridge on the backside of the tarsometatarsus is formed by the third toe's midfoot bones, which are set to the plantar side in respect to their neighbors, with which they are fused. this is particularly noticeable in the lower part of the bone, where a corresponding groove is present on the dorsal side. this smooths out proximad, with the proximal half of the bone—which is not known from an unflattened specimen to date—probably oval or triangular (with the point backwards) in cross section.