according to hassan et al. (2008), around 59.4% of fur are carriers of the e1b1b paternal haplogroup. of these, 68.4% bear the v32 subclade. approximately 6.3% also belong to the haplogroup j1. this points to significant patrilineal gene flow from neighbouring afro-asiatic-speaking populations. the remaining fur individuals are primarily carriers of the a3b2 lineage (31.3%), which is instead common among nilotes.