this still leaves open questions relating to the origin of the orb web. did it evolve only once, with araneids later losing the cribellum (and hence cribellate silk), or did it evolve separately in uloborids and araneids? if it evolved only once, how early did this happen in the evolutionary history of entelegyne spiders? if late, then uloborids and araneids may form a monophyletic group, orbiculariae. if very early, then uloborids and ananeids may not be closely related, and many more araneoids that do not spin webs would have lost this ability secondarily.