Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The nucleotides in the DNA molecule are linked in series from one phosphate, to the next sugar, to the next phosphate and so on. The DNA molecule can be considered as a ladder with the phosphate and the sugar molecules making up the uprights and the bases making up the rungs. The DNA molecule is a twisted ladder with the phosphate and sugar uprights or rails running in opposite orientation to one another.