Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The nucleotides of DNA line up so that the sugar and phosphate molecules make two long backbones like the handrails of a ladder. To make the rungs of the ladder, two bases join together, between the sugar molecules on the two handrails. The phosphate molecules do not have any rungs between them.