Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
DNA also includes sugars and phosphate groups that make the phosphate-deoxyribose backbone. If you think of the structure of DNA as a twisted ladder, the rungs of the ladder are made from the nitrogenous bases. These bases pair up to make each step of the ladder. They also only pair up in a specific way.