One of a line of military commanders who from 1192 until 1867 were generally the actual rulers of Japan, although nominally subordinate to the emperor.
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The H+ in the blood, however, cannot cross the blood-brain barrier, and thus cannot influence the medullary chemoreceptors. Carbon dioxide in the arterial blood can cross the blood-brain barrier and, through the formation of carbonic acid, can lower the pH of cerebrospinal fluid (fig. 16.30).n increase in blood CO2 stimulates breathing indirectly by lowering the pH of blood and cerebrospinal fluid (CSF). This figure illustrates how a rise in blood CO2 increases the H+ concentration (lowers the pH) of CSF and thereby stimulates chemoreceptor neurons in the medulla oblongata.