Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
In this model of a very short section of DNA you can see how the A-T and C-G base pairs make up the rungs of the ladder and the sugars and phosphates make up the two long strands.