Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
To make the rungs of the ladder, two bases join together, between the sugar molecules on the two handrails. The phosphate molecules do not have any rungs between them. THERE IS ONLY ONE WAY THE BASES CAN PAIR UP ON THE RUNGS OF THE DNA LADDER. An adenine molecule only pairs with a thymine. A cytosine only pairs with a guanine. They can pair in either order on a rung, giving four possible combinations of bases --.