Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
While the deoxyribose sugar bonds to the phosphate groups on either side of it, as well as to a nitrogen-containing base, the phosphate group only bonds to the two deoxyribose sugars that surround it. A nitrogen-containing base forms half of each rung of the DNA ladder.