Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
These nucleotides are composed of three parts: a phosphate, a sugar (deoxyribose), and a type of compound base. The deoxyribose and phosphate form the backbone of nucleic acid (the side of the ladder) while the base connect the two polynucleotide chains (like the rungs of the ladder).