Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The rungs of the DNA ladder are made of complementary nitrogenous base pairs. DNA contains four different nitrogenous bases: adenine, thymine, cytosine and guanine. The nitrogenous bases are divided into the purines and the pyrimidines; each rung of the DNA molecule contains one purine and one pyrimidine bonded together. Keep Learning.