Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
Answer Wiki. Since DNA is thought to be like a twisted ladder the rungs, not including the rails, of DNA are made up of nitrogenous bases: adenine (A), guanine (G), cytosine (C), and thymine (T).