Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The rungs of the ladder are pairs of 4 types of nitrogen bases  (thymines, adenines, guanines, cytosines).   ---   The pairs are guanine and cytosine (G-C), or adeninâ¦e and thymine  (A-T). The rungs of the dna ladder are made of alternating sugars and  phosophate groups.