Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
These bases are the rungs of the DNA ladder. (It takes two bases to form a rung -- one for each side of the ladder.). A sugar molecule, a base, and a phosphate molecule group together to make up a nucleotide. Nucleotides are abundant in the cell's nucleus.