Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
DNA is often said to have a sugar and phosphate backbone.. Each rung of the ladder is made of two nitrogenous bases linked together in the middle. The length of a DNA molecule is often measured in base pairs, or bpthat is, the number of rungs in the ladder.