Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The 'steps' on the 'DNA Ladder' are made up of the four nitrogenous bases, Cytosine, Guanine, Thymine, and Adenine, while the pairing bases (Adenine & Thymine, Cytosine & Guan â¦ ine) are bonded together with a hydrogen bond. ---   The pairs are guanine and cytosine (G-C), or adenin â¦ e and thymine  (A-T). The rungs of the dna ladder are made of alternating sugars and  phosophate groups.