Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The sugars are deoxyribose (in DNA) and ribose (in RNA). The Phosphate groups plus the sugars form the sides of the ladder as the molecule comes together. The bases are the rungs or steps on the ladder. The entire molecule will form a twisted ladder when fully complete.