Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
DNA had two sides, or strands, and that these strands were twisted together like a twisted ladder -- the double helix. The sides of the ladder comprise the sugar-phosphate portions of adjacent nucleotides bonded together.