Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The sides of the DNA ladder is composed of sugar and phosphate. 4  bases that make up the rungs of the DNA ladder are A, T, G, and C.  The shape of the DNA is a double helix â¦ or twisted ladder.