Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
On the sides of the double helix, the two strands of polynucleotide, (think of this as the sides of the ladder) are where the phosphates and sugars, arranged in different orders, are located. The bases connect the two sides (think of this as rungs on the ladder) with a weak chemical bond.