Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
Strands of DNA are made of the sugar and phosphate portions of the nucleotides, while the middle parts are made of the nitrogenous bases. The nitrogenous bases on the two strands of DNA pair up, purine with pyrimidine (A with T, G with C), and are held together by weak hydrogen bonds. Watson and Crick discovered that DNA had two sides, or strands, and that these strands were twisted together like a twisted ladder -- the double helix.