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Plymouth was colder, whereas Jamestown was warmer and more humid, and therefore more prone to harbor diseases. I'm sure that there were a lot more differences, but those are the big ones that leapt into my mind.
Plymouth, the shire town of the county and capital of the colony. This was the original 1620 settlement of the Mayflower Pilgrims, and continued as the largest and most significant settlement in the colony until its dissolution in 1691.