Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
A DNA molecule is like a ladder because there are two sides (the sugar-phosphate backbones of the two strands) and rungs (each rung being a pair of complementary bases).