In addition to kidney diseases and dehydration, other conditions that can cause elevated creatinine levels include heart failure, dehydration, excessive blood loss that causes shock, gout, strenuous exercise, muscle injury, etc.
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Male ligers also have the same levels of testosterone on average as an adult male lion, yet are azoospermic in accordance with Haldane's rule. In addition, female ligers may also attain great size, weighing approximately 320 kg (705 lb) and reaching 3.05 m (10 ft) long on average, and are often fertile.