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This supports the idea that in embryos from wild-type mothers, dpp H~- mutations cause the transformation of dorsal into ven-tral epidermis. If tissue determined to become dorsal epidermis dies in the dpp rim-null embryos, then one should not see ventral or lateral tissue formed in the double-mutant combination.
The idea that the recessive embryonic lethal alleles are leaky dpp ni~ mutations is consistent with ob-servations of interactions with mutations elsewhere in dpp. trans combinations of recessive embryonic lethal or haplo-lethal dpp n~ alleles with either dpp ~sk or dpp shy lesions elicit a variety of phenotypes.