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This supports the idea that in embryos from wild-type mothers, dpp H~- mutations cause the transformation of dorsal into ven-tral epidermis. If tissue determined to become dorsal epidermis dies in the dpp rim-null embryos, then one should not see ventral or lateral tissue formed in the double-mutant combination.
Given the similarity in lethal phenotype between homozygotes for the recessive embryonic lethal dpp alleles and monosomics for dpp ~+, Spencer et al. (1982) suggested that these reces-sive alleles are leaky dpp Hm-mutations. The recessive embryonic lethal alleles partially inactivate functions of the dpp ~sk region.