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ATP synthase is a ubiquitous enzyme that is located in the inner membranes of mitochondria, thylakoid membranes of chloroplasts, or the plasma membranes of bacteria.
Chloroplast ATP synthase and the enzyme from some photosynthetic bacteria have 2 different, although similar, b-type subunits in the proton translocating F O portion, namely b and b' , one copy of each.