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ATP synthase is a ubiquitous enzyme that is located in the inner membranes of mitochondria, thylakoid membranes of chloroplasts, or the plasma membranes of bacteria.
ATP synthase complexes of bacteria, mitochondria and chloroplasts are all very similar, with only minor differences. Mitochondria are believed to have evolved from symbiotic aerobic bacteria ingested by an anaerobic host cell. The limiting membrane of the bacterium became the inner mitochondrial membrane.