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1. fourth in order. 2. containing four elements or groups. quaternary. 1 adj, pertaining to a chemical compound in which four atoms or groups of elements are bonded to one atom, such as a quaternary ammonium compound in which four organic radicals are substituted for the four hydrogen molecules on an ammonium ion.
In such proteins, quaternary structure refers to the number and arrangement of the individual polypeptide chains. Each polypeptide is referred to as a subunit of the protein. The same forces and bonds that create tertiary structure also hold subunits together in a stable complex to form the complete protein. Individual chains may be identical, somewhat similar, or totally different.