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A plenum space is a part of a building that can facilitate air circulation for heating and air conditioning systems, by providing pathways for either heated/conditioned or return airflows, usually at greater than atmospheric pressure.
The two main types of supply systems are extended plenum and central plenum. A central plenum system is a system where all of the branch ducts start at the same location called a plenum. The extended plenum system consists of a supply air plenum which is connected to the outlet of the hvac equipment. The main supply ducts are attached to this and typically run to the end of the home. The size of them is reduced as necessary to maintain adequate air flow. The branch ducts attach to the main ducts and carry the conditioned air to the individual registers. That is the basics of how a system works.