Lumia (citrus)
A recent genomic analysis of several species commonly called 'lemons' or 'limes' revealed that the various individual lumias have different genetic backgrounds. The 'Hybride Fourny' was found to be an F1 hybrid of a citron- pomelo cross, while the 'Jaffa lemon' was a more complex cross between the two species, perhaps an F2 hybrid. The Pomme d'Adam arose from a citron-micrantha cross, while two other lumias, the ‘Borneo’ and ‘Barum’ lemons, were found to be citron-pomelo-micrantha mixes.
Citrus taxonomy
While most citrus hybrids derive from the three core species, hybrids have also been derived from the micrantha, the Ichang papeda and the kumquat. The best known hybrid from micrantha is the Key lime (or Mexican lime), derived from the breeding of a male citron and a female micrantha. Several citrus varieties are Ichang papeda/mandarin crosses (for which Swingle coined the term ichandarin), including Sudachi and Yuzu (which also includes smaller contributions from pomelo and kumquat). Other more exotic citrus have likewise proved hybrids that include papeda. For example, the Indian wild orange, once suggested as a possible ancestor of today's cultivated citrus fruits, yielded conflicting phylogenetic placements in more limited genetic analysis, but study of nuclear markers and chloroplast DNA showed it to be of maternal citron lineage, with further genetic contributions from mandarin and papeda. The wild Mountain citron, native to the Malay peninsula and called "Citrus halimii" at the time of its discovery, has been found to be a complex mix, being half papeda (type not specified), the other half including kumquat, mandarin and pomelo, with only trace amounts of citron.