Wolves as pets and working animals
Though wolves are trainable, they lack the same degree of tractability seen in dogs. They are generally not as responsive as dogs to coercive techniques involving fear, aversion to stimuli, and force. Generally, far more work is required to obtain the same degree of reliability seen in most dogs. Even then, once a certain behavior has been repeated several times, wolves may get bored and ignore subsequent commands. Wolves are most responsive toward positive conditioning and rewards, though simple praise is not sufficient as in most dogs. Unlike dogs, wolves tend to respond more to hand signals than voice. Although they are more difficult to control than dogs, they can be easier to teach if the motivation exists.
Wolves are less suitable than dogs for working. Swedish wolf biologist Erik Zimen once tried to form a dog sled team composed entirely of wolves. The experiment failed as the wolves ignored most commands and were far more prone to fighting than sled dogs. North American wolves and wolf hybrids were used as experimental attack dogs by the South African Defence Force in Apartheid South Africa in an attempt to breed animals capable of tracking guerrillas. Because the wolves would not follow even basic commands, the experiment was abandoned. However, their sense of smell can rival that of established scent hounds. According to reports published in 2006, ongoing research at the Perm Institute of Internal Troops in Russia demonstrated that wolf/dog cross-breeds took 15–20 seconds to track down a target in training sessions, whereas ordinary police dogs took 3–4 minutes. Their success has led to plans to use them as police dogs for the Russian police.
Wolf
The age of first breeding in gray wolves depends largely on environmental factors: when food is plentiful, or when wolf populations are heavily managed, wolves can rear pups at younger ages in order to better exploit abundant resources. This is further demonstrated by the fact that captive wolves have been known to breed as soon as they reach 9–10 months, while the youngest recorded breeding wolves in the wild were 2 years old. Females are capable of producing pups every year, with one litter annually being the average. Unlike the coyote, the gray wolf never reaches reproductive senescence. Estrus typically occurs in late winter, with older, multiparous females entering estrus 2–3 weeks earlier than younger females. During pregnancy, female wolves remain in a den located away from the peripheral zone of their territories, where violent encounters with other packs are more likely. Old females usually whelp in the den of their previous litter, while younger females typically den near their birthplace. The gestation period lasts 62–75 days, with pups usually being born in the summer period.
Wolves bear relatively large pups in small litters compared to other canid species. The average litter consists of 5–6 pups, with litter sizes tending to increase in areas where prey is abundant, though exceptionally large litters of 14–17 pups occur only 1% of the time. Pups are usually born in spring, coinciding with a corresponding increase in prey populations. Pups are born blind and deaf, and are covered in short soft grayish-brown fur. They weigh at birth, and begin to see after 9–12 days. The milk canines erupt after one month. Pups first leave the den after 3 weeks. At 1.5 months of age, they are agile enough to flee from danger. Mother wolves do not leave the den for the first few weeks, relying on the fathers to provide food for them and their young. Pups begin to eat solid food at the age of 3–4 weeks. Pups have a fast growth rate during their first four months of life: during this period, a pup's weight can increase nearly 30 times. Wolf pups begin play fighting at the age of 3 weeks, though unlike young foxes and coyotes, their bites are inhibited. Actual fights to establish hierarchy usually occur at 5–8 weeks of age. This is in contrast to young foxes and coyotes, which may begin fighting even before the onset of play behavior. By autumn, the pups are mature enough to accompany adults on hunts for large prey.