Ponderosa lemon
The ponderosa lemon originated in roughly 1887, and is believed to come from a chance seedling grown in Hagerstown, Maryland. It was later named and introduced to the nursery trade in 1900. Though often referred to as a cross between the lemon and citron, a recent genomic analysis showed it to solely contain pomelo ("Citrus maxima") and citron ("Citrus medica") DNA, perhaps being an F2 hybrid. Thus they cannot derive from true lemons, which have a mandarin orange component.
Citrus taxonomy
The trifoliate orange is a cold hardy plant distinguishable by its compound leaves with three leaflets and its deciduous nature, but is close enough to the genus "Citrus" to be used as a rootstock. Swingle moved the trifoliate orange from "Citrus" to its own genus, "Poncirus". It was long thought to be the only member of this genus, until a second species, "Poncirus polyandra", was discovered in Yunnan (China) in the 1980s. Early phylogenetic analysis nested "Poncirus" within the citrus, suggesting they all belong to a single genus, but the genomic sequencing of Wu, "et al.", placed it outside the cluster representing "Citrus", and the authors retained a separate "Poncirus" genus.