Dietary biology of the Eurasian eagle-owl
Generally speaking, there appears to be rather strictly adhered size hierarchy among birds of prey and the very largest raptorial prey attacked by Eurasian eagle-owl tend to roughly equal to their own size. Larger raptors, such as eagles and vultures, seem to be largely invulnerable to eagle-owls. However, in Scandinavia, there are two cases of eagle-owls preying on “fairly large” nestlings of white-tailed eagles ("Haliaeetus albicilla"). Given the size, average adult weight of , and formidable character of this eagle, it is possible that the young of other raptors larger than themselves may too be attacked. Eastern imperial eagles ("Aquila heliaca"), which are nearly as large as the white-tailed eagle at an average weight of , have been killed by eagle-owls in Kazakhstan. Remarkable, considering the greater threat that the eagle normally poses to the eagle-owl than vica versa, it appeared a golden eagle was amongst the prey remains found around a nest in the French Pyrenees. Cases in Spain where large, formidable birds of prey, the Egyptian vulture ("Neophron percnopterus") and Bonelli's eagle, that are roughly equal in size to the eagle-owls, both weighing just over , have been predated, these seem to be taken from the nest or shortly after independence and rare occurrences. However, in Turkey, the eagle-owl is apparently a regular and serious predator of Bonelli's eagles. The Eurasian eagle-owl predates the largest members of the most species-rich diurnal raptor genera: the northern goshawks from the "Accipiter" genus, the upland buzzards from the "Buteo" genus and the gyrfalcons ("F. rusticolus") of the "Falco" genus. Additional large raptors, i.e. those that can average or more in body mass, taken as prey include osprey ("Pandion haliaetus"), the red kites ("Milvus milvus"), short-toed eagles ("Circaetus gallicus"), the greater spotted eagle ("Clanga clanga"), the lesser spotted eagle ("Clanga pomarina"), and the rough-legged buzzards ("Buteo lagopus") and saker falcons ("Falco cherrug"), in both of which females average more than . The taking of rough-legged buzzards and snowy owls must be confined to full-grown victims, since they nest further north than Eurasian eagle-owls. Eurasian eagle-owls also hunt the smallest raptors available, including those such as lesser kestrels ("Falco naumanni") and Japanese sparrowhawks ("Accipiter gularis") that weigh less than . Some of these are cases of eagle-owls killing nestlings or fledglings but they can easily overtake adult buzzards, goshawks and falcons of any size.
Northern goshawk
Outside of the accipitrid group, heavy predation on different varieties of raptorial birds by northern goshawks can continue unabated. Many types of owl are taken and in Europe, the northern goshawk is the second most prolific predator of owls behind the Eurasian eagle owl ("Bubo bubo"). In Bavaria, Germany, the long-eared owl ("Asio otus") was the second most common prey species for nesting goshawks. In the Białowieża Forest of Poland, fairly high numbers of the tawny owl ("Strix alucco") were taken. In all, some 18 species of owl have been recorded in the diet, ranging in size from the Eurasian ("Glaucidium passerinum") and northern pygmy owls ("Glaucidium gnoma") at and , respectively, to all the large northern "Strix" owls including adults and even the great horned owl. Whether adults have ever been killed as prey though is unknown but goshawks have been known to kill great horned owls that they've found near their nests. In addition, about eight species of falcon have been identified in the foods of goshawks. Adult falcons of small species such as kestrels and merlins ("Falco columbarius") can be overpowered quite easily if they can manage to surprise the prey. Larger falcons have turned up in the diet as well, including the prairie falcon ("Falco mexicanus") and the saker falcon ("Falco cherrug"), although possible only nestlings of these species. Brief aerial skirmishes between goshawks and peregrine falcons ("Falco peregrinus") have been described but neither species is known to have killed one another in the wild. In Schleswig-Holstein, Germany, at least four small passerines species were recorded as nesting close to active goshawk nest, due to the incidental shelter that the fierce goshawks inadvertently provides from smaller raptors which are their main predators. Such raptors, including Eurasian kestrels, Eurasian sparrowhawks and long-eared owls, not only avoid goshawk activity where possible but also were found to have lower nest productivity any time they nested relatively close to goshawks per the study. A similar phenomenon, with goshawks inadvertently providing shelter to small passerines, has been recorded in North America as well.
Competition for northern goshawks can also come from mammalian carnivores. Martens, and to a lesser extent other weasels, are presumably one of their more major competitors as their diet often consists of similar prey primarily during spring and summer, tree squirrels and woodland birds, but little has been studied in terms of how the two types of predator effect each other. Most recorded interactions have been predatory, as the goshawk has been recorded preying on a dozen species, from the least weasel ("Mustela nivalis") to the stone marten ("Martes foina"). Northern goshawks have also been recorded as feeding on much bigger predators such as the red fox ("Vulpes vulpes"), the raccoon dog ("Nyctereutes procyonoides") and the striped skunk ("Mephitis mephitis"), but it is not clear whether these were actual kills, as many may be encountered as already dead carrion. Domestic carnivores are sometimes eaten, including dogs and cats, the latter of which has reportedly been taken alive by goshawks. The red fox is a surprisingly considerable competitor for resources with northern goshawks. It was found in Norway that goshawk numbers were higher when voles were at peak numbers, not due to voles as a food source but because foxes were more likely to eat the rodents and ignore grouse, whereas during low vole numbers the foxes are more likely to compete with goshawks over grouse as prey. A decrease of the fox population of Norway due to sarcoptic mange was found result in an increase of grouse numbers and, in turn, northern goshawks. In some areas, red foxes have been found to steal up to half of the goshawks’ kills.