Caspian Sea
The wide and endorheic Caspian Sea has a north–south orientation and its main freshwater inflow, the Volga River, enters at the shallow north end. Two deep basins occupy its central and southern areas. These facts lead to horizontal differences in temperature, salinity, and ecology. The Caspian Sea spreads out over nearly 750 miles (1,200 km) from north to south, with an average width of 200 miles (320 km). It covers a region of around 149,200 square miles (386,400 square km)—bigger than the region of Japan—and its surface is about 90 feet (27 meters) below sea level. The sea bed in the southern part reaches as low as 1,023 m (3,356 ft) below sea level, which is the second lowest natural depression on earth after Lake Baikal (−1,180 m, −3,871 ft). The ancient inhabitants of its coast perceived the Caspian Sea as an ocean, probably because of its saltiness and large size.
Caspian lowland desert
The ecoregion is approximately contained within the Caspian Depression, and sunken geological region feeding in to the Caspian, the surface of which is itself at below worldwide sea level. The northern section is almost 900 km wide, and stretches up to 300 km inland across Russia and Kazakhstan. The sector along the southeast coast of the Caspian is mostly in Turkmenistan. Elevations throughout the ecoregion range between -28 and +100 meters. The dry areas were once underwater, but as the sea has slowly receded the dry areas have revealed a landscape of sand dunes, ridges, salt domes, salt pans ('shors'), and clay deserts ('takyrs'). 
The climate of the ecoregion is represented by that in the Volga delta, in the Astrakhan Nature Reserve, where the climate is cool semi-arid (Köppen climate classification BSk). This climate is characterized by high variation in temperature, both daily and seasonally; with low precipitation. Average annual precipitation in the Caspian lowlands is 198 mm. Average temperature in January is , and in July is