Ponderosa lemon
The ponderosa lemon originated in roughly 1887, and is believed to come from a chance seedling grown in Hagerstown, Maryland. It was later named and introduced to the nursery trade in 1900. Though often referred to as a cross between the lemon and citron, a recent genomic analysis showed it to solely contain pomelo ("Citrus maxima") and citron ("Citrus medica") DNA, perhaps being an F2 hybrid. Thus they cannot derive from true lemons, which have a mandarin orange component.
Citrus
The most recent research indicates an origin in the Himalayas. Previous research indicated an origin in the part of Southeast Asia bordered by Northeast India, Burma (Myanmar), and the Yunnan province of China, and it is in this region that some commercial species such as oranges, mandarins, and lemons originated. Citrus fruit has been cultivated in an ever-widening area since ancient times.