Pteranodon
While the first "Pteranodon" wing bones were collected by Marsh and Cope in the early 1870s, the first "Pteranodon" skull was found on May 2, 1876, along the Smoky Hill River in Wallace County (now Logan County), Kansas, USA, by Samuel Wendell Williston, a fossil collector working for Marsh. A second, smaller skull soon was discovered as well. These skulls showed that the North American pterosaurs were different from any European species, in that they lacked teeth and had bony crests on their skulls. Marsh recognized this major difference, describing the specimens as "distinguished from all previously known genera of the order Pterosauria by the entire absence of teeth." Marsh recognized that this characteristic warranted a new genus, and he coined the name "Pteranodon" ("wing without tooth") in 1876. Marsh reclassified all the previously named North American species from "Pterodactylus" to "Pteranodon". He considered the smaller skull to belong to "Pteranodon occidentalis", based on its size. Marsh classified the larger skull, YPM 1117, in the new species "Pteranodon longiceps", which he thought to be a medium-sized species in between the small "P. occidentalis" and the large "P. ingens". Marsh also named several additional species: Pteranodon comptus and "Pteranodon nanus" were named for fragmentary skeletons of small individuals, while "Pteranodon gracilis" was based on a wing bone that he mistook for a pelvic bone. He soon realized his mistake, and re-classified that specimen again into a separate genus, which he named "Nyctosaurus". "P, nanus" was also later recognized as a "Nyctosaurus" specimen.
Pararhabdodon
Near the village Abella de la Conca, in the 1990s, palaeontologist Marc Boada discovered a new site in the Tremp Formation, bearing dinosaur fossils. From this site, later named Les Llaus (LL), a right dentary, specimen designation IPS SRA 27, was excavated. In 1997, Casanovas-Cladellas and colleagues stated this dentary was discovered from SRA, the site where the original "Pararhabdodon" remains were found. The following year, they described the specimen and referred it to "P. isonensis", and again stated it was from the same stratigraphic level. In 2006, the stratigraphy of the region was re-evaluated, and the dentary's location was corrected to the LL locality, away from and below the SRA locality.