Eurasian eagle-owl
The Eurasian eagle-owl is a member of the genus "Bubo", which may include either 22 or 25 extant species. Almost all the larger owl species in the world today are included in "Bubo". Based on an extensive fossil record and a central distribution of extant species on that continent, the "Bubo" appears to have evolved into existence in Africa, although early radiations seem to branch from southern Asia as well. Two genera belonging to the scops owls complex, the giant scops owls ("Otus gurneyi") found in Asia and the "Ptilopsis" or the white-faced scops owl found in Africa, although firmly ensconced in the scops owl group, appear to share some characteristics with the eagle-owls. The "Strix" is also related to the "Bubo" and is considered a "sister complex", with the "Pulsatrix" possibly being intermediate between the two. The Eurasian eagle-owl appears to represent an expansion of the genus "Bubo" into the Eurasian continent. A few of the other species of "Bubo" seem to have been derived from the Eurasian eagle-owl, making it a "paraspecies", or they at least share a relatively recent common ancestor. The Pharaoh eagle-owl, distributed in the Arabian Peninsula and sections of the Sahara Desert through North Africa where rocky outcrops are found, was until recently considered a subspecies of the Eurasian eagle-owl. It appears that the Pharaoh eagle-owl differs about 3.8% in mitochondrial DNA from the Eurasian eagle-owl, well past the minimum genetic difference to differentiate species of 1.5%. Smaller and paler than Eurasian eagle-owls, the Pharaoh can also be considered a distinct species largely due to its higher pitched and more descending call and the observation that Eurasian eagle-owls formerly found in Morocco ("B. b. hispanus") apparently did not breed with the co-existing Pharaoh eagle-owls. On the contrary, the race still found together with the Pharaoh eagle-owl in the wild ("B. b. interpositus") in the central Middle East has been found to interbreed in the wild with the Pharaoh eagle-owl, although genetical materials have indicated "interpositus" may itself be a distinct species from the Eurasian eagle-owl, as it differs from the nominate subspecies of the Eurasian eagle-owl by 2.8% in mitochondrial DNA. The Indian eagle-owl was also considered a subspecies of Eurasian eagle-owl until recently, but its smaller size, distinct voice (more clipped and high-pitched than Eurasians), and the fact that it is largely allopatric in distribution (filling out the Indian subcontinent) with other Eurasian eagle-owl races has led to it being considered a distinct species. The mitochondrial DNA of the Indian species also appears considerably distinct from the Eurasian species. The Cape eagle-owl ("B. capensis") appears to represent a return of this genetic line back into the African continent, where it leads a lifestyle similar to northern eagle-owls albeit far to the south. Another offshoot of the northern "Bubo" group is the snowy owl. It appears to have separated from other "Bubo" at least 4 million years ago.
White-tailed eagle
The wintering range is less well understood for the white-tailed eagle given the extreme reductions and fluctuations of northern breeding populations over the last few centuries, so that the delineation of regular wintering areas versus areas of mere vagrancy is difficult to ascertain. It is known that a small number winter on Etang de Lindre of Lorraine, France as well as an area on the border of France to Germany around Strasbourg, with vagrants to elsewhere in France, as well as to Spain, Portugal and Malta. A well-defined wintering population may occur in much of the Netherlands, even with infrequent modern breeding in the northern coastal areas. A non-breeding population is known in western Germany from North Rhine-Westphalia to Bonn, as well as far northern Germany into southwestern Denmark. Other established wintering areas are known in Europe in west-central Italy, northern Austria, fairly broadly in southern Slovakia and northern Hungary and a few protected pockets of Southeast Europe apart from the portions in the north and east where they still breed. Intermittent forms of vagrancy and migration (most from eagles that breed in or disperse from Russia) are known to occur in several areas of Turkey, the Levant countries, Azerbaijan and Iran down to even the Persian Gulf, albeit seldom is the species to be found commonly or reliably anywhere in these regions. Further east, rare wintering areas are known in a few small, scarce pockets of Turkmenistan, Afghanistan, Uzbekistan, Tajikistan and Pakistan. It is a very rare winter visitor to India, namely the extreme northwest and along the border of Nepal to Bhutan and extreme northern Bangladesh. Scattered pockets of wintering birds are known to occur too in central and southern China, into northeastern Mynamar, and more broadly and regularly in much of northeastern China. Good numbers winter too in much of South Korea and Japan down to as far as Honshu. White-tailed eagles dispersing from their breeding grounds or natal sites in the Russian Far East are known to occasionally disperse across the Bering Sea to North America in several parts of the Aleutian Islands, the Pribilof Islands and some of mainland coastal Alaska down to Kodiak Island. Some white-tailed eagles even bred in Alaska on Attu Island in the late 1970s to the early 1980s (until 1984 when the last attempts were recorded) but it was not clear whether young were ever successfully fledged.