Ponderosa lemon
The ponderosa lemon originated in roughly 1887, and is believed to come from a chance seedling grown in Hagerstown, Maryland. It was later named and introduced to the nursery trade in 1900. Though often referred to as a cross between the lemon and citron, a recent genomic analysis showed it to solely contain pomelo ("Citrus maxima") and citron ("Citrus medica") DNA, perhaps being an F2 hybrid. Thus they cannot derive from true lemons, which have a mandarin orange component.
Citrus
The large citrus fruit of today evolved originally from small, edible berries over millions of years. Citrus plants diverged from a common ancestor about 15 million years ago, which was about when it diverged from the closely related severinia, for example the Chinese box orange. About 7 million years ago, citrus plants diverged into two groups, the main citrus genus and the ancestors of the trifoliate orange (poncirus), which is closely enough related that it can still be hybridized with all other citrus. These estimates are made using genetic mapping of plant chloroplasts. A DNA study published in "Nature" in 2018 concludes that citrus trees originated in the foothills of the Himalayas, in the area of Assam (India), western Yunnan (China), and northern Myanmar.