Monts de Gueret Animal Park
The wolf pack howls on average for 85 seconds. It is usually started by a single wolf, after which other wolves may join in. The pack leader, the alpha wolf, usually uses a lower-pitched howl and howls more frequently than wolves standing on the lower rungs of the pack's hierarchy. Although some functions of the howling are still unknown by scientists, they believe that wolves may howl to scare off other wolf packs from their territory and to gather their own pack, as wolves possess perfect hearing and their howling can be heard for up to in the forest and in open space. Researchers have found a relationship between the number of times a wolf howls and the strength of their relationship with other pack members. According to Friederike Range, co-director of the University of Vienna's Wolf Science Center, "the strength of the relationship between wolves predicted how many times a wolf howled."
Alpha (ethology)
In the past, the prevailing view on grey wolf packs was that they consisted of individuals vying with each other for dominance, with dominant grey wolves being referred to as the "alpha" male and female, and the subordinates as "beta" and "omega" wolves. This terminology was first used in 1947 by Rudolf Schenkel of the University of Basel, who based his findings on researching the behaviour of captive grey wolves. This view on gray wolf pack dynamics was later popularized by the researcher L. David Mech in his 1970 book "The Wolf". He later found additional evidence that the concept of an Alpha male may have been an erroneous interpretation of incomplete data and formally disavowed this terminology in 1999. He explained that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves. Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and their offspring of the previous 1–3 years.