Monts de Gueret Animal Park
The wolf pack howls on average for 85 seconds. It is usually started by a single wolf, after which other wolves may join in. The pack leader, the alpha wolf, usually uses a lower-pitched howl and howls more frequently than wolves standing on the lower rungs of the pack's hierarchy. Although some functions of the howling are still unknown by scientists, they believe that wolves may howl to scare off other wolf packs from their territory and to gather their own pack, as wolves possess perfect hearing and their howling can be heard for up to in the forest and in open space. Researchers have found a relationship between the number of times a wolf howls and the strength of their relationship with other pack members. According to Friederike Range, co-director of the University of Vienna's Wolf Science Center, "the strength of the relationship between wolves predicted how many times a wolf howled."
Dhole
Dholes are more social than gray wolves, and have less of a dominance hierarchy, as seasonal scarcity of food is not a serious concern for them. In this manner, they closely resemble African wild dogs in social structure. They live in clans rather than packs, as the latter term refers to a group of animals that always hunt together. In contrast, dhole clans frequently break into small packs of 3–5 animals, particularly during the spring season, as this is the optimal number for catching fawns. Dominant dholes are hard to identify, as they do not engage in dominance displays as wolves do, though other clan members will show submissive behaviour toward them. Intragroup fighting is rarely observed. Dholes are far less territorial than wolves, with pups from one clan often joining another without trouble once they mature sexually. Clans typically number 5-12 individuals in India, though clans of 40 have been reported. In Thailand, clans rarely exceed three individuals. Unlike other canids, there is no evidence of dholes using urine to mark their territories or travel routes. When urinating, dholes, especially males, may raise one hind leg or both to result in a handstand. Handstand urination is also seen in bush dogs ("Speothos venaticus"). They may defecate in conspicuous places, though a territorial function is unlikely, as faeces are mostly deposited within the clan's territory rather than the periphery. Faeces are often deposited in what appear to be communal latrines. They do not scrape the earth with their feet, as other canids do, to mark their territories.