Ponderosa lemon
The ponderosa lemon originated in roughly 1887, and is believed to come from a chance seedling grown in Hagerstown, Maryland. It was later named and introduced to the nursery trade in 1900. Though often referred to as a cross between the lemon and citron, a recent genomic analysis showed it to solely contain pomelo ("Citrus maxima") and citron ("Citrus medica") DNA, perhaps being an F2 hybrid. Thus they cannot derive from true lemons, which have a mandarin orange component.
Citron
The citron is thought to have been native to India, in valleys at the foothills of the eastern Himalayas. It is thought that by the 4th century BC, when Theophrastus mentions the "Persian or Median apple", the citron was mostly cultivated in the Persian Gulf on its way to the Mediterranean basin, where it was cultivated during the later centuries in different areas as described by Erich Isaac. Many mention the role of Alexander the Great and his armies as they attacked Persia and what is today Pakistan, as being responsible for the spread of the citron westward, reaching the European countries such as Greece and Italy.