Pack (canine)
Animals which typically predominate over others are associated with the term "alpha". Among pack-living wolves, alpha wolves are the genetic parents of most cubs in the pack. Such access to mating females creates strong selective pressure for intra-sex competition.
Alpha (ethology)
In the past, the prevailing view on grey wolf packs was that they consisted of individuals vying with each other for dominance, with dominant grey wolves being referred to as the "alpha" male and female, and the subordinates as "beta" and "omega" wolves. This terminology was first used in 1947 by Rudolf Schenkel of the University of Basel, who based his findings on researching the behaviour of captive grey wolves. This view on gray wolf pack dynamics was later popularized by the researcher L. David Mech in his 1970 book "The Wolf". He later found additional evidence that the concept of an Alpha male may have been an erroneous interpretation of incomplete data and formally disavowed this terminology in 1999. He explained that it was heavily based on the behavior of captive packs consisting of unrelated individuals, an error reflecting the once prevailing view that wild pack formation occurred in winter among independent gray wolves. Later research on wild gray wolves revealed that the pack is usually a family consisting of a breeding pair and their offspring of the previous 1–3 years.