Monts de Gueret Animal Park
The wolf pack howls on average for 85 seconds. It is usually started by a single wolf, after which other wolves may join in. The pack leader, the alpha wolf, usually uses a lower-pitched howl and howls more frequently than wolves standing on the lower rungs of the pack's hierarchy. Although some functions of the howling are still unknown by scientists, they believe that wolves may howl to scare off other wolf packs from their territory and to gather their own pack, as wolves possess perfect hearing and their howling can be heard for up to in the forest and in open space. Researchers have found a relationship between the number of times a wolf howls and the strength of their relationship with other pack members. According to Friederike Range, co-director of the University of Vienna's Wolf Science Center, "the strength of the relationship between wolves predicted how many times a wolf howled."
Wolf
Wolves are highly territorial animals, and generally establish territories far larger than they require to survive in order to assure a steady supply of prey. Territory size depends largely on the amount of prey available and the age of the pack's pups, tending to increase in size in areas with low prey populations or when the pups reach the age of 6 months, thus having the same nutritional needs as adults. Wolf packs travel constantly in search of prey, covering roughly 9% of their territory per day (average ). The core of their territory is on average , in which they spend 50% of their time. Prey density tends to be much higher in the territory's surrounding areas, though wolves tend to avoid hunting in the fringes of their range unless desperate, because of the possibility of fatal encounters with neighboring packs. The smallest territory on record was held by a pack of six wolves in northeastern Minnesota, which occupied an estimated , while the largest was held by an Alaskan pack of ten wolves encompassing a area. Wolf packs are typically settled, and usually only leave their accustomed ranges during severe food shortages.