Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black mongoose
The black mongoose ("Galerella nigrata") is a species of mongoose found in Namibia and Angola. Although originally described as a separate species by Thomas (1928), it was often been considered a subspecies of the slender mongoose. However, genetic analysis has confirmed its status as a separate species. Evidence suggests the two species diverged around four million years ago, likely due to some populations becoming separated as the habitat in southern Africa was changing. The black mongoose now occupies a distinct habitat in areas with large boulders and rocky outcrops known as inselbergs in the mountainous regions of northwest Namibia and southwest Angola. After remaining in these areas for millions of years, the black mongoose is highly specialized to survive in the harsh arid environment.