Digestion
Digestion begins in the mouth with the secretion of saliva and its digestive enzymes. Food is formed into a bolus by the mechanical mastication and swallowed into the esophagus from where it enters the stomach through the action of peristalsis. Gastric juice contains hydrochloric acid and pepsin which would damage the walls of the stomach and mucus is secreted for protection. In the stomach further release of enzymes break down the food further and this is combined with the churning action of the stomach. The partially digested food enters the duodenum as a thick semi-liquid chyme. In the small intestine, the larger part of digestion takes place and this is helped by the secretions of bile, pancreatic juice and intestinal juice. The intestinal walls are lined with villi, and their epithelial cells is covered with numerous microvilli to improve the absorption of nutrients by increasing the surface area of the intestine.
Parasitic nutrition
Endoparasites have various anatomical and biochemical adaptations, typically at the host-parasite interface, to maximise nutrient acquisition. One such adaptation is the tegument, a metabolically active external cover that plays an important role in nutrient extraction from the host. The parasite tegument is permeable to various organic solutes and has transporters for the facilitated or active uptake of nutrients. Various studies have attempted to characterise these transporters in a number of parasites e.g. the amino acid transporter molecules in protozoa. Cestodes do not have a gut so the tegument is therefore critical for nutrient uptake. In cestodes the tegument is highly efficient with spine-like microtriches, similar to microvilli, to increase the surface area available for nutrient acquisition. In many parasites the tegument structure has folds or microvilli to maximise the surface area available for diffusion and uptake of nutrients. The tegument also commonly has additional organelles and features with important functions in metabolism including the glycocalyx. The glycocalyx is a carbohydrate-rich layer that enhances nutrient absorption and secretes enzymes to aid primary digestion.