Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black wildebeest
The black wildebeest inhabits open plains, grasslands, and Karoo shrublands in both steep mountainous regions and lower undulating hills. The altitudes in these areas vary from . The herds are often migratory or nomadic, otherwise they may have regular home ranges of . Female herds roam in home ranges around in size. In the past, black wildebeest occurred in the highveld temperate grasslands during the dry winter season and the arid Karoo region during the rains. However, as a result of massive hunting of the animal for its hide, they vanished from their historical range, and are now largely limited to game farms and protected reserves in Southern Africa. In most reserves, the black wildebeest shares its habitat with the blesbok and the springbok.