Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Alcelaphine gammaherpesvirus 2
All species of African wildebeest and hartbeest can be infected or at least carry this virus. There is no direct evidence that other African wild variants like addax can be hosts, but research is ongoing. The alcelaphine antelope affecting form is primarily found in Africa, in the natural habitat of wildebeest, hartebeest, and topi. It has also occurred in zoos and wild animal parks that also kept wildebeest. The increasing popularity in North America of wild animal parks may spread its range further.