Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black wildebeest
The black wildebeest is currently included in the same genus as the blue wildebeest ("Connochaetes taurinus"). This has not always been the case, and at one time the latter was placed under a separate genus of its own, "Gorgon". The black wildebeest lineage seems to have diverged from the blue wildebeest in the mid- to late Pleistocene, and became a distinct species around a million years ago. This evolution is quite recent on a geologic time scale.