Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black wildebeest
The black wildebeest can maintain its body temperature within a small range in spite of large fluctuations in external temperatures. It shows well-developed orientation behaviour towards solar radiation, which helps it thrive in hot, and often shadeless, habitats. The erythrocyte count is high at birth and increases till the age of 2-3 months, while in contrast, the leucocyte count is low at birth and falls throughout the animal's life. The neutrophil count is high at all ages. The haematocrit and haemoglobin content decreases till 20-30 days after birth. A peak in the content of all these haemological parameters occurs at the age of 2-3 months, after which the readings gradually decline, reaching their lowest values in the oldest individuals. The presence of fast-twitch fibres and the ability of the muscles to use large amounts of oxygen help explain the rapid running speed of the black wildebeest and its high resistance to fatigue. Individuals may live for about 20 years.