Question mark
In the early 13th century, when the growth of communities of scholars (universities) in Paris and other major cities led to an expansion and streamlining of the book-production trade, punctuation was rationalized by assigning Alcuin's stroke-over-dot specifically to interrogatives; by this time the stroke was more sharply curved and can easily be recognized as the modern question mark.
Exclamation mark
The exclamation mark is sometimes used in conjunction with the question mark. This can be in protest or astonishment ("Out of all places, the squatter-camp?!"); a few writers replace this with a single, nonstandard punctuation mark, the interrobang, which is the combination of a question mark and an exclamation mark.