Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black wildebeest
A dominant male black wildebeest has a harem of females and will not allow other males to mate with them. The breeding season occurs at the end of the rainy season and lasts a few weeks between February and April. When one of his females comes into estrus, the male concentrates on her and mates with her several times. Sexual behaviour by the male at this time includes stretching low, ears down, sniffing of the female's vulva, performing ritual urination, and touching his chin to the female's rump. At the same time, the female keeps her tail upwards (sometimes vertically) or swishes it across the face of the male. The pair usually separates after copulation, but the female occasionally follows her mate afterwards, touching his rump with her snout. During the breeding season, the male loses condition as he spends little time grazing. Males are known to mount other males.