Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Black wildebeest
The natural populations of black wildebeest, endemic to the southern part of Africa, were almost completely exterminated in the 19th century, due to their reputation as pests and the value of their hides and meat. However, the species has been reintroduced widely from captive specimens, both in private areas and nature reserves throughout most of Lesotho, Swaziland, and South Africa. It has also been introduced outside its natural range in Namibia and Kenya.