Black wildebeest
Features necessary for defending a territory, such as the horns and broad-based skull of the modern black wildebeest, have been found in their fossil ancestors. The earliest known fossil remains are in sedimentary rock in Cornelia in the Orange Free State and date back about 800,000 years. Fossils have also been reported from the Vaal River deposits, though whether or not they are as ancient as those found in Cornelia is unclear. Horns of the black wildebeest have been found in sand dunes near Hermanus in South Africa. This is far beyond the recorded range of the species and it has been suggested that these animals may have migrated to that region from the Karoo.
Wildebeest
Not all wildebeest are migratory. Black wildebeest herds are often nomadic or may have a regular home range of . Bulls may occupy territories, usually about apart, but this spacing varies according to the quality of the habitat. In favourable conditions, they may be as close as , or they may be as far apart as in poor habitat. Female herds have home ranges of about in size. Herds of nonterritorial bachelor males roam at will and do not seem to have any restrictions on where they wander.