What was the first animal to be domesticated by humans?
Domestication
The beginnings of animal domestication involved a protracted coevolutionary process with multiple stages along different pathways. It is proposed that there were three major pathways that most animal domesticates followed into domestication: (1) commensals, adapted to a human niche (e.g., dogs, cats, fowl, possibly pigs); (2) prey animals sought for food (e.g., sheep, goats, cattle, water buffalo, yak, pig, reindeer, llama and alpaca); and (3) targeted animals for draft and nonfood resources (e.g., horse, donkey, camel). The dog was the first domesticant, and was established across Eurasia before the end of the Late Pleistocene era, well before cultivation and before the domestication of other animals. Humans did not intend to domesticate animals from, or at least they did not envision a domesticated animal resulting from, either the commensal or prey pathways. In both of these cases, humans became entangled with these species as the relationship between them, and the human role in their survival and reproduction, intensified, leading eventually to a formalised animal husbandry. Although the directed pathway proceeded from capture to taming, the other two pathways are not as goal-oriented and archaeological records suggest that they took place over much longer time frames.
Cat
It was long thought that cat domestication was initiated in Egypt, because cats in ancient Egypt were venerated since around 3100 BC.
However, the earliest indication for the taming of an African wildcat ("F. lybica") was found in Cyprus, where a cat skeleton was excavated close by a human Neolithic grave dating to around 7500 BC. African wildcats were probably first domesticated in the Near East. The leopard cat ("Prionailurus bengalensis") was tamed independently in China around 5500 BC, though this line of partially domesticated cats leaves no trace in the domestic cat populations of today.
As of 2007, the domestic cat was the second-most popular pet in the U.S. by number of pets owned, after freshwater fish. As of 2010, it was ranked the third-most popular pet in the UK, after fish and dogs, with around 8 million being owned.
Maya civilization
The Maya had few domestic animals; dogs were domesticated by 3000 BC, and the Muscovy duck by the Late Postclassic. Ocellated turkeys were unsuitable for domestication, but were rounded up in the wild and penned for fattening. All of these were used as food animals; dogs were additionally used for hunting. It is possible that deer were also penned and fattened.
Domestication of animals
Humans were already reliant on domestic plants and animals when they imagined the domestic versions of wild animals. Although horses, donkeys, and Old World camels were sometimes hunted as prey species, they were each deliberately brought into the human niche for sources of transport. Domestication was still a multi-generational adaptation to human selection pressures, including tameness, but without a suitable evolutionary response then domestication was not achieved. For example, despite the fact that hunters of the Near Eastern gazelle in the Epipaleolithic avoided culling reproductive females to promote population balance, neither gazelles nor zebras possessed the necessary prerequisites and were never domesticated. There is no clear evidence for the domestication of any herded prey animal originating in Africa.
Animal testing
Animals have repeatedly been used through the history of biomedical research. In 1831, the founders of the Dublin Zoo were members of the medical profession who were interested in studying animals while they were alive and when they were dead. In the 1880s, Louis Pasteur convincingly demonstrated the germ theory of medicine by inducing anthrax in sheep. In the 1880s, Robert Koch infected mice and guinea pigs with anthrax and tuberculosis. In the 1890s, Ivan Pavlov famously used dogs to describe classical conditioning. In World War I, German agents infected sheep bound for Russia with anthrax, and inoculated mules and horses of the French cavalry with the equine glanders disease. Between 1917 and 1918, the Germans infected mules in Argentina bound for American forces, resulting in the death of 200 mules. Insulin was first isolated from dogs in 1922, and revolutionized the treatment of diabetes. On 3 November 1957, a Soviet dog, Laika, became the first of many animals to orbit the earth. In the 1970s, antibiotic treatments and vaccines for leprosy were developed using armadillos, then given to humans. The ability of humans to change the genetics of animals took a large step forwards in 1974 when Rudolf Jaenisch was able to produce the first transgenic mammal, by integrating DNA from the SV40 virus into the genome of mice. This genetic research progressed rapidly and, in 1996, Dolly the sheep was born, the first mammal to be cloned from an adult cell.
Bovidae
The earliest evidence of cattle domestication is from 8000 BC, suggesting that the process began in Cyprus and the Euphrates basin.
Origin of the domestic dog
The origin of the domestic dog is not clear. The domestic dog is a member of the genus "Canis", which forms part of the wolf-like canids, and is the most widely abundant terrestrial carnivore. The closest living relative of the dog is the extant gray wolf, and there is no evidence of any other canine contributing to its genetic lineage. The dog and the extant gray wolf are sister taxa, as modern wolves are not closely related to the wolves that were first domesticated. The archaeological record and genetic analysis show the remains of the Bonn–Oberkassel dog buried beside humans 14,200 years ago to be the first undisputed dog, with disputed remains occurring 36,000 years ago. These dates imply that the earliest dogs arose in the time of human hunter-gatherers and not agriculturists. The dog was the first species to be domesticated.
Domesticated plants and animals of Austronesia
The candlenut ("Aleurites moluccanus") was first domesticated in Island Southeast Asia. Remains of harvested candlenuts have been recovered from archaeological sites in Timor and Morotai in eastern Indonesia, dated to around 13,000 BP and 11,000 BP respectively.
Neolithic
Domestication of sheep and goats reached Egypt from the Near East possibly as early as 6000 BC. Graeme Barker states "The first indisputable evidence for domestic plants and animals in the Nile valley is not until the early fifth millennium BC in northern Egypt and a thousand years later further south, in both cases as part of strategies that still relied heavily on fishing, hunting, and the gathering of wild plants" and suggests that these subsistence changes were not due to farmers migrating from the Near East but was an indigenous development, with cereals either indigenous or obtained through exchange. Other scholars argue that the primary stimulus for agriculture and domesticated animals (as well as mud-brick architecture and other Neolithic cultural features) in Egypt was from the Middle East.
Even-toed ungulate
Artiodactyls have been hunted by primitive humans for various reasons: for meat or fur, as well as to use their forehead weapons, bones, and teeth as weapons or tools. Their domestication began around 8000 BCE. To date, humans have domesticated goats, sheep, cattle, camels, llamas, alpacas, and pigs. Initially, livestock was used primarily for food, but they began being used for work activities around 3000 BCE. Clear evidence exists of antelope being used for food 2 million years ago in the Olduvai Gorge, part of the Great Rift Valley. Cro-Magnons relied heavily on reindeer for food, skins, tools, and weapons; with dropping temperatures and increased reindeer numbers at the end of the Pleistocene, they became the prey of choice. Reindeer remains accounted for 94% of bones and teeth found in a cave above the Céou River that was inhabited around 12,500 years ago.
Greylag goose
"Anser anser", the greylag goose, is a member of the waterfowl family Anatidae. It was first described by the Swedish naturalist Carl Linnaeus in 1758 as "Anas anser", but was transferred two years later to the new genus "Anser", erected by the French zoologist Mathurin Jacques Brisson, where it is the type species. Two subspecies are recognised; "A. a. anser", the western greylag goose, breeds in Iceland and north and central Europe; "A. a. rubrirostris", the eastern greylag goose, breeds in Romania, Turkey and Russia eastwards to northeastern China. The two subspecies intergrade where their ranges meet. The greylag goose sometimes hybridises with other species of goose including the barnacle goose ("Branta leucopsis") and the Canada goose ("Branta canadensis"), and occasionally with the mute swan ("Cygnus olor"). The greylag goose was one of the first animals to be domesticated; this happened at least 3000 years ago in Ancient Egypt, the domestic breed being known as "A. a. domesticus". As the domestic goose is a subspecies of the greylag goose they are able to interbreed, with the offspring sharing characteristics of both the wild and tame birds.
Canidae
One canid, the domestic dog, entered into a partnership with humans a long time ago. The archaeological record shows the first undisputed dog remains buried beside humans 14,700 years ago, with disputed remains occurring 36,000 years ago. These dates imply that the earliest dogs arose in the time of human hunter-gatherers and not agriculturists. The dog was the first domesticated species. The fact that wolves are pack animals with cooperative social structures may have been the reason that the relationship developed. Humans benefited from the canid's loyalty, cooperation, teamwork, alertness and tracking abilities, while the wolf may have benefited from the use of weapons to tackle larger prey and the sharing of food. Humans and dogs may have evolved together.
Dairy farming
While cattle were domesticated as early as 11,000 years ago as a food source and as beasts of burden, the earliest evidence of using domesticated cows for dairy production is the seventh millennium BC - the early Neolithic era - in northwestern Anatolia. Dairy farming developed elsewhere in the world in subsequent centuries: the sixth millennium BC in eastern Europe, the fifth millennium BC in Africa, and the fourth millennium BC in Britain and Northern Europe.
Origin of the domestic dog
Ancient DNA supports the hypothesis that dog domestication preceded the emergence of agriculture and was initiated close to the Last Glacial Maximum 27,000 YBP when hunter-gatherers preyed on megafauna, and when proto-dogs might have taken advantage of carcasses left on site by early hunters, assisted in the capture of prey, or provided defense from large competing predators at kill-sites. Wolves were probably attracted to human campfires by the smell of meat being cooked and discarded refuse in the vicinity, first loosely attaching themselves and then considering these as part of their home territory where their warning growls would alert humans to the approach of outsiders. The wolves most likely drawn to human camps were the less-aggressive, subdominant pack members with lowered flight response, higher stress thresholds, less wary around humans, and therefore better candidates for domestication. The earliest sign of domestication in dogs was the neotonization of skull morphology and the shortening of snout length that results in tooth crowding, reduction in tooth size, and a reduction in the number of teeth, which has been attributed to the strong selection for reduced aggression. This process may have begun during the initial commensal stage of dog domestication, even before humans began to be active partners in the process.
Self-domestication
Self-domestication is described by biological anthropologist Richard Wrangham as being in an environment where lessening of aggression was beneficial for survival. As grain plants and livestock became domesticated 9,000 years ago in the Fertile Crescent, wild cats adapted to living with humans, hunting rodents in grain stores and "abandoning their aggressive wild-born behaviors", which led to today's house cats.
Equus (genus)
Archaeological, biogeographical, and linguistic evidence suggests that the donkey was first domesticated by nomadic pastoral people in North Africa over 5,000 years ago. The animals were used to help cope with the increased aridity of the Sahara and the Horn of Africa. Genetic evidence finds that the donkey was domesticated twice based on two distinct mitochondrial DNA haplogroups. It also points to a single ancestor, the Nubian wild ass. Attempts to domesticate zebras were largely unsuccessful, though Walter Rothschild trained some to draw a carriage in England.
Paleolithic dog
The debate centres around "Homo sapiens" and if they had entered into cooperation with wolves soon after they moved into Eurasia, and if so when and where did these wolves change into domesticated dogs. In arguing that domestication leads to reduction in size, the late domestication theory ignores that modern horses and pigs are larger than their wild ancestors. It also ignores that if hunter-gathers entered into a hunting relationship with wolves then there would be no need of selection for a reduction in size. A reduction in size would have occurred much later when humans moved into agricultural villages. The late domestication theory does not consider the possibility that humans may have formed a relationship with non-domesticated wolves and that dogs in the early stages of domestication might be indistinguishable from wolves. According to indigenous North Americans, over the past 20,000 years the canids living with them were wolves that could not be distinguished as dogs.
Bovini
Bovins and humans have had a long and complex relationship. Five of seven species have been successfully domesticated, with one species (cattle) being the most successful member of their lineage. Domesticated shortly after the last Ice Age, there are at least 1.4 billion cattle in the world. Domestic bovins have been selectively bred for beef, dairy products, leather and serve as working animals. However many species of wild cattle are threatened by extinction due to habitat loss to make room for cattle farming as well as unregulated hunting. Some are already extinct like the aurochs, two subspecies of European bison and perhaps the kouprey.
Domestication of animals
Prey pathway animals include sheep, goats, cattle, water buffalo, yak, pig, reindeer, llama and alpaca. The right conditions for the domestication for some of them appear to have been in place in the central and eastern Fertile Crescent at the end of the Younger Dryas climatic downturn and the beginning of the Early Holocene about 11,700 YBP, and by 10,000 YBP people were preferentially killing young males of a variety of species and allowed the females to live in order to produce more offspring. By measuring the size, sex ratios, and mortality profiles of zooarchaeological specimens, archeologists have been able to document changes in the management strategies of hunted sheep, goats, pigs, and cows in the Fertile Crescent starting 11,700 YBP. A recent demographic and metrical study of cow and pig remains at Sha’ar Hagolan, Israel, demonstrated that both species were severely overhunted before domestication, suggesting that the intensive exploitation led to management strategies adopted throughout the region that ultimately led to the domestication of these populations following the prey pathway. This pattern of overhunting before domestication suggests that the prey pathway was as accidental and unintentional as the commensal pathway.
Domestic goose
Domestic geese ("Anser anser domesticus" or "Anser cygnoides domesticus") are domesticated grey geese (either greylag geese or swan geese) that are kept by humans as poultry for their meat, eggs, and down feathers since ancient times.
Musa acuminata
"Musa acuminata" is one of the earliest plants to be domesticated by humans for agriculture. They were first domesticated in Southeast Asia and surrounding areas (possibly New Guinea, eastern Indonesia, and the Philippines) around 10 kya (8000 BCE). It has been suggested that "M. acuminata" may have originally been domesticated for parts other than the fruit. Either for fiber, for construction materials, or for its edible male bud. They were selected early for parthenocarpy and seed sterility in their fruits, a process that might have taken thousands of years. This initially led to the first 'human-edible' banana diploid clones (modern AA cultivars). Diploid clones are still able to produce viable seeds when pollinated by wild species. This resulted in the development of triploid clones which were conserved for their larger fruit.
Bovidae
The domestication of bovids has contributed in shifting the dependence of human beings from hunting and gathering to agriculture. The Bovidae include three of the five domesticated mammals whose use has spread outside their original ranges, namely cattle, sheep, and goats; all are from Eurasia, and are now found across the world. The other two species are the horse and pig. Other large bovids that have been domesticated but which remain within the ranges of their wild ancestors are the water buffalo (from the Indian water buffalo), domestic yak (from the wild yak), zebu (from the Indian aurochs), gayal (from the gaur) and Bali cattle (from the banteng).
Some antelopes have been domesticated including the scimitar oryx, addax, elands and the extinct bubal hartebeest. In Ancient Egypt oryxes, addaxes and bubal hartebeests are depicted in carved walls.
History of agriculture
Jujube was domesticated in the Indian subcontinent by 9000 BC. Barley and wheat cultivation – along with the domestication of cattle, primarily sheep and goats – followed in Mehrgarh culture by 8000–6000 BC. This period also saw the first domestication of the elephant. Pastoral farming in India included threshing, planting crops in rows – either of two or of six – and storing grain in granaries. Cotton was cultivated by the 5th–4th millennium BC. By the 5th millennium BC, agricultural communities became widespread in Kashmir. Irrigation was developed in the Indus Valley Civilization by around 4500 BC. The size and prosperity of the Indus civilization grew as a result of this innovation, leading to more thoroughly planned settlements which used drainage and sewers. Archeological evidence of an animal-drawn plough dates back to 2500 BC in the Indus Valley Civilization.
Origin of the domestic dog
The domestic dog is the most widely abundant large carnivore. The dog was the first species and the only large carnivore to have been domesticated. Over the past 200 years, dogs have undergone rapid phenotypic change and were formed into today's modern dog breeds due to artificial selection imposed by humans. These breeds can vary in size and weight from a teacup poodle to a giant mastiff. The skull, body, and limb proportions vary significantly between breeds, with dogs displaying more phenotypic diversity than can be found within the entire order of carnivores. Some breeds demonstrate outstanding skills in herding, retrieving, scent detection, and guarding, which demonstrates the functional and behavioral diversity of dogs. There have been major advances in understanding the genes that gave rise to the phenotypic traits of dogs. The first dogs were certainly wolflike, however the phenotypic changes that coincided with the dog–wolf genetic divergence are not known.
Origin of the domestic dog
The hypothesis is that two genetically different, and possibly now extinct, wolf populations were domesticated independently in eastern and western Eurasia to produce paleolithic dogs. The eastern Eurasian dogs then dispersed westward alongside humans, reaching western Europe 6,400–14,000 years ago where they partially replaced the western paleolithic dogs. A single domestication is thought to be due to chance, however dual domestication on different sides of the world is unlikely to have happened randomly and it suggests that external factors - an environmental driver - may have forced wolves to work together with humans for survival. It is possible that wolves took advantage of resources that humans had, or humans may have been introduced to wolves in an area in which they didn't previously live.
Animal husbandry
A cow was a great advantage to a villager as she produced more milk than her calf needed, and her strength could be put to use as a working animal, pulling a plough to increase production of crops, and drawing a sledge, and later a cart, to bring the produce home from the field. Draught animals were first used about 4,000 BC in the Middle East, increasing agricultural production immeasurably. In southern Asia, the elephant was domesticated by 6,000 BC.
African wildcat
Results of genetic research indicate that the African wildcat diverged into three clades about 173,000 years ago, namely the Near Eastern wildcat, Southern African wildcat and Asiatic wildcat. African wildcats were first domesticated about 10,000 years ago in the Near East, and are the ancestors of the domestic cat. In Cyprus, an African wildcat was found in a burial site next to a human skeleton in the Pre-Pottery Neolithic B settlement Shillourokambos. The graves are estimated to have been established by Neolithic farmers about 9,500 years ago and are the earliest known evidence for a close association between a human and a cat. Their proximity indicates that the cat may have been tamed or domesticated.
Crossings between domestic cats and African wildcats are still common today.
History of agriculture
Wild grains were collected and eaten from at least 20,000 BC. From around 9500 BC, the eight Neolithic founder crops – emmer wheat, einkorn wheat, hulled barley, peas, lentils, bitter vetch, chick peas, and flax – were cultivated in the Levant. Rye may have been cultivated earlier but this remains controversial. Rice was domesticated in China by 6200 BC with earliest known cultivation from 5700 BC, followed by mung, soy and azuki beans. Pigs were domesticated in Mesopotamia around 11,000 BC, followed by sheep between 11,000 BC and 9000 BC. Cattle were domesticated from the wild aurochs in the areas of modern Turkey and Pakistan around 8500 BC. Sugarcane and some root vegetables were domesticated in New Guinea around 7000 BC. Sorghum was domesticated in the Sahel region of Africa by 5000 BC. In the Andes of South America, the potato was domesticated between 8000 BC and 5000 BC, along with beans, coca, llamas, alpacas, and guinea pigs. Bananas were cultivated and hybridized in the same period in Papua New Guinea. In Mesoamerica, wild teosinte was domesticated to maize by 4000 BC. Cotton was domesticated in Peru by 3600 BC. Camels were domesticated late, perhaps around 3000 BC.
Uruk period
The Uruk period also saw important developments in the realm of pastoralism. First of all, it is in this period that the wild onager was finally domesticated as the donkey. It was the first domesticated equid in the region and became the most important beast of burden in the Near East (the dromedary was only domesticated in the 3rd millennium BC, in Arabia). With its high transport capacity (about double that of a human), it enabled the further development of trade over short and long distances. Pastoralism of animals which had already been domesticated (sheep, horses, cattle) also developed further. Previously these animals had been raised mainly as sources of meat, but they now became more important for the products which they provided (wool, fur, hides, milk) and as beasts of burden. This final aspect was especially connected with the cattle, which became essential for work in the fields with the appearance of the ard, and the donkey which assumed a major role in the transportation of goods.
Cat behavior
Humans and cats have a long history starting back with the ancient Egyptians, who were the first to domesticate cats.
Raw milk
Humans first learned to regularly consume the milk of other mammals following the domestication of animals during the Neolithic Revolution or the development of agriculture. This development occurred independently in several places around the world from as early as 9000–7000 BC in Mesopotamia to 3500–3000 BC in the Americas. The most important dairy animals—cattle, sheep and goats—were first domesticated in Mesopotamia, although domestic cattle had been independently derived from wild aurochs populations several times since. From there dairy animals spread to Europe (beginning around 7000 BC but not reaching Britain and Scandinavia until after 4000 BC), and South Asia (7000–5500 BC).
History of agriculture
Agriculture began independently in different parts of the globe, and included a diverse range of taxa. At least 11 separate regions of the Old and New World were involved as independent centers of origin. Some of the earliest known domestications were of animals. Domestic pigs had multiple centres of origin in Eurasia, including Europe, East Asia and Southwest Asia, where wild boar were first domesticated about 10,500 years ago. Sheep were domesticated in Mesopotamia between 11,000 BC and 9000 BC. Cattle were domesticated from the wild aurochs in the areas of modern Turkey and Pakistan around 8500 BC. Camels were domesticated late, perhaps around 3000 BC.
Livestock
The dog was domesticated early; dogs appear in Europe and the Far East from about 15,000 years ago. Goats and sheep were domesticated in multiple events sometime between 11,000 and 5,000 years ago in Southwest Asia. Pigs were domesticated by 8,500 BC in the Near East and 6,000 BC in China. Domestication of the horse dates to around 4000 BC. Cattle have been domesticated since approximately 10,500 years ago. Chickens and other poultry may have been domesticated around 7000 BC.
Domestication
The domestication of animals and plants began with the wolf ("Canis lupus") at least 15,000 years before present (YBP), which then led to a rapid shift in the evolution, ecology, and demography of both humans and numerous species of animals and plants. The sudden appearance of the domestic dog ("Canis lupus familiaris") in the archaeological record was followed by livestock and crop domestication, and the transition of humans from foraging to farming in different places and times across the planet. Around 10,000 YBP, a new way of life emerged for humans through the management and exploitation of plant and animal species, leading to higher-density populations in the centers of domestication, the expansion of agricultural economies, and the development of urban communities.
Rodent
Rodents are used widely as model organisms in animal testing. Albino mutant rats were first used for research in 1828 and later became the first animal domesticated for purely scientific purposes. Nowadays, the house mouse is the most commonly used laboratory rodent, and in 1979 it was estimated that fifty million were used annually worldwide. They are favored because of their small size, fertility, short gestation period and ease of handling and because they are susceptible to many of the conditions and infections that afflict humans. They are used in research into genetics, developmental biology, cell biology, oncology and immunology. Guinea pigs were popular laboratory animals until the late 20th century; about 2.5 million guinea pigs were used annually in the United States for research in the 1960s, but that total decreased to about 375,000 by the mid-1990s. In 2007, they constituted about 2% of all laboratory animals. Guinea pigs played a major role in the establishment of germ theory in the late 19th century, through the experiments of Louis Pasteur, Émile Roux, and Robert Koch. They have been launched into orbital space flight several times—first by the USSR on the Sputnik 9 biosatellite of March 9, 1961, with a successful recovery. The naked mole rat is the only known mammal that is poikilothermic; it is used in studies on thermoregulation. It is also unusual in not producing the neurotransmitter substance P, a fact which researchers find useful in studies on pain.
Laboratory rat
Rat-baiting was a popular sport which involved filling a pit with rats and timing how long it took for a terrier to kill them all. Over time, breeding the rats for these contests may have produced variations in color, notably the albino and hooded varieties. The first time one of these albino mutants was brought into a laboratory for a study was in 1828, in an experiment on fasting. Over the next 30 years rats were used for several more experiments and eventually the laboratory rat became the first animal domesticated for purely scientific reasons.
Interspecies friendship
In the previous 11,000 years, humans have brought a wide range of species into domestication to use as livestock, working animals, household pets, and companions. The influence of human behaviour on domesticated animals has led to many species having learned to co-exist - sometimes leading to the formation of an interspecies friendship. For example, interspecies friendships are often observed in humans with their domesticated pets and in pets that live in the same household such as cats and dogs.
History of the domestic sheep
Sheep husbandry spread quickly in Europe. Excavations show that in about 6000 BC, during the Neolithic period of prehistory, the Castelnovien people, living around Châteauneuf-les-Martigues near present-day Marseille in the south of France, were among the first in Europe to keep domestic sheep. Practically from its inception, ancient Greek civilization relied on sheep as primary livestock, and were even said to name individual animals. Scandinavian sheep of a type seen today — with short tails and multi-colored fleece — were also present early on. Later, the Roman Empire kept sheep on a wide scale, and the Romans were an important agent in the spread of sheep raising through much of Europe. Pliny the Elder, in his Natural History ("Naturalis Historia"), speaks at length about sheep and wool. Declaring "Many thanks, too, do we owe to the sheep, both for appeasing the gods, and for giving us the use of its fleece.", he goes on to detail the breeds of ancient sheep and the many colors, lengths and qualities of wool. Romans also pioneered the practice of blanketing sheep, in which a fitted coat (today usually of nylon) is placed over the sheep to improve the cleanliness and luster of its wool.
Domestication of the horse
How and when horses became domesticated is disputed. The clearest evidence of early use of the horse as a means of transport is from chariot burials dated c. 2000 BCE. However, an increasing amount of evidence supports the hypothesis that horses were domesticated in the Eurasian Steppes approximately 3500 BCE; recent discoveries in the context of the Botai culture suggest that Botai settlements in the Akmola Province of Kazakhstan are the location of the earliest domestication of the horse.
History of the domestic sheep
No ovine species native to the Americas has ever been domesticated, despite being closer genetically to domestic sheep than many Asian and European species. The first domestic sheep in North America—most likely of the Churra breed—arrived with Christopher Columbus' second voyage in 1493. The next transatlantic shipment to arrive was with Hernán Cortés in 1519, landing in Mexico. No export of wool or animals is known to have occurred from these populations, but flocks did disseminate throughout what is now Mexico and the Southwest United States with Spanish colonists. Churras were also introduced to the Navajo tribe of Native Americans, and became a key part of their livelihood and culture. The modern presence of the Navajo-Churro breed is a result of this heritage.
Horse
The horse ("Equus ferus caballus") is one of two extant subspecies of "Equus ferus". It is an odd-toed ungulate mammal belonging to the taxonomic family Equidae. The horse has evolved over the past 45 to 55 million years from a small multi-toed creature, "Eohippus", into the large, single-toed animal of today. Humans began domesticating horses around 4000 BC, and their domestication is believed to have been widespread by 3000 BC. Horses in the subspecies "caballus" are domesticated, although some domesticated populations live in the wild as feral horses. These feral populations are not true wild horses, as this term is used to describe horses that have never been domesticated, such as the endangered Przewalski's horse, a separate subspecies, and the only remaining true wild horse. There is an extensive, specialized vocabulary used to describe equine-related concepts, covering everything from anatomy to life stages, size, colors, markings, breeds, locomotion, and behavior.
Origin of the domestic dog
The debate centres around "Homo sapiens" and if they had entered into cooperation with wolves soon after they moved into Eurasia, and if so when and where did these wolves change into domesticated dogs. In arguing that domestication leads to reduction in size, the late domestication theory ignores that modern horses and pigs are larger than their wild ancestors. It also ignores that if hunter-gathers entered into a hunting relationship with wolves then there would be no need of selection for a reduction in size. A reduction in size would have occurred much later when humans moved into agricultural villages. The late domestication theory does not consider the possibility that humans may have formed a relationship with non-domesticated wolves and that dogs in the early stages of domestication might be indistinguishable from wolves. According to indigenous North Americans, over the past 20,000 years the canids living with them were wolves that could not be distinguished as dogs.
Animal husbandry
Fossilised chicken bones dated to 5040 BC have been found in northeastern China, far from where their wild ancestors lived in the jungles of tropical Asia, but archaeologists believe that the original purpose of domestication was for the sport of cockfighting.
Homo mermanus
The "Homo mermanus" are often referred to as "Atlanteans", as it is in the city of Atlantis that their first complex society emerged. Like humans they learned to domesticate animals, such as dolphins, while humans domesticated horses, dogs and camels.
Rabbit
Rabbits have long been domesticated. Beginning in the Middle Ages, the European rabbit has been widely kept as livestock, starting in ancient Rome. Selective breeding has generated a wide variety of rabbit breeds, many of which (since the early 19th century) are also kept as pets. Some strains of rabbit have been bred specifically as research subjects.
Domestic pig
In August 2015, a study looked at over 100 pig genome sequences to ascertain their process of domestication, which was assumed to have been initiated by humans, involved few individuals, and relied on reproductive isolation between wild and domestic forms. The study found that the assumption of reproductive isolation with population bottlenecks was not supported. The study indicated that pigs were domesticated separately in Western Asia and China, with Western Asian pigs introduced into Europe, where they crossed with wild boar. A model that fitted the data included a mixture with a now extinct ghost population of wild pigs during the Pleistocene. The study also found that despite back-crossing with wild pigs, the genomes of domestic pigs have strong signatures of selection at DNA loci that affect behavior and morphology. The study concluded that human selection for domestic traits likely counteracted the homogenizing effect of gene flow from wild boars and created domestication islands in the genome. The same process may also apply to other domesticated animals.
Animal husbandry
The domestication of livestock was driven by the need to have food on hand when hunting was unproductive. The desirable characteristics of a domestic animal are that it should be useful to man, should be able to thrive in his company, should breed freely and be easy to tend.
Domestication was not a single event, but a process repeated at various periods in different places. Sheep and goats were the animals that accompanied the nomads in the Middle East, while cattle and pigs were associated with more settled communities.
Animal husbandry
Meanwhile, in South America, the llama and the alpaca had been domesticated, probably before 3,000 BC, as beasts of burden and for their wool. Neither was strong enough to pull a plough which limited the development of agriculture in the New World.
Archaeogenetics
Archaeogenetics has been used to trace the domestication of pigs throughout the old world. These studies also reveal evidence about the details of early farmers. Methods of Archaeogenetics have also been used to further understand the development of domestication of dogs. Genetic studies have shown that all dogs are descendants from the gray wolf, however, it is currently unknown when, where, and how many times dogs were domesticated. Some genetic studies have indicated multiple domestications while others have not. Archaeological findings help better understand this complicated past by providing solid evidence about the progression of the domestication of dogs. As early humans domesticated dogs the archaeological remains of buried dogs became increasingly more abundant. Not only does this provide more opportunities for archaeologists to study the remains, it also provides clues about early human culture.
Milk
Humans first learned to consume the milk of other mammals regularly following the domestication of animals during the Neolithic Revolution or the development of agriculture. This development occurred independently in several global locations from as early as 9000–7000 BC in Mesopotamia to 3500–3000 BC in the Americas. People first domesticated the most important dairy animals – cattle, sheep and goats – in Southwest Asia, although domestic cattle had been independently derived from wild aurochs populations several times since. Initially animals were kept for meat, and archaeologist Andrew Sherratt has suggested that dairying, along with the exploitation of domestic animals for hair and labor, began much later in a separate secondary products revolution in the fourth millennium BC. Sherratt's model is not supported by recent findings, based on the analysis of lipid residue in prehistoric pottery, that shows that dairying was practiced in the early phases of agriculture in Southwest Asia, by at least the seventh millennium BC.
Cowpea
While it may be uncertain when cultivation began it is still considered one of the oldest domesticated crops. Remains of charred cowpeas from rock shelters in Central Ghana have been dated to the second millennium BCE. In 2300 BC the cowpea is believed to have made its way into South East Asia where secondary domestication events may have occurred. From there they traveled north to the Mediterranean, where they were used by the Greeks and Romans. The first written references to the cowpea were in 300 BC and they probably reached Central and North America during the slave trade through the 17th to early 19th centuries.
Origin of the domestic dog
When and where dogs were first domesticated has taxed geneticists for the past 20 years and archaeologists for many decades longer. Identifying the earliest dogs is difficult because the key morphological characters that are used by zooarchaeologists to differentiate domestic dogs from their wild wolf ancestors (size and position of teeth, dental pathologies, and size and proportion of cranial and postcranial elements) were not yet fixed during the initial phases of the domestication process. The range of natural variation among these characters that may have existed in ancient wolf populations, and the time it took for these traits to appear in dogs, are unknown.
Africa
The domestication of cattle in Africa preceded agriculture and seems to have existed alongside hunter-gatherer cultures. It is speculated that by 6000 BC, cattle were domesticated in North Africa. In the Sahara-Nile complex, people domesticated many animals, including the donkey and a small screw-horned goat which was common from Algeria to Nubia.
Eurasian Steppe
The primary domesticated animals raised were sheep and goats with fewer cattle than one might expect. Camels were used in the drier areas for transport as far west as Astrakhan. There were some yaks along the edge of Tibet. The horse was used for transportation and warfare. The horse was first domesticated on the Pontic–Caspian or Kazakh steppe sometime before 3000 BC, but it took a long time for mounted archery to develop and the process is not fully understood. The stirrup does not seem to have been completely developed until 300 AD (see Stirrup, Saddle, Composite bow, Domestication of the horse and related articles).
Domestic pig
Archaeological evidence suggests that pigs were domesticated from wild boar as early as 13,000–12,700 BC in the Near East in the Tigris Basin, Çayönü, Cafer Höyük, Nevalı Çori being managed in the wild in a way similar to the way they are managed by some modern New Guineans. Remains of pigs have been dated to earlier than 11,400 BC in Cyprus. Those animals must have been introduced from the mainland, which suggests domestication in the adjacent mainland by then. There was also a separate domestication in China which took place about 8000 years ago.
War elephant
The first elephant species to be tamed was the Asian elephant, for use in agriculture. Elephant taming – not full domestication, as they are still captured in the wild, rather than being bred in captivity – may have begun in any of three different places. The oldest evidence comes from the Indus Valley Civilization, around roughly 4500 BC. Archaeological evidence for the presence of wild elephants in the Yellow River valley in Shang China (1600–1100 BC) may suggest that they also used elephants in warfare. The wild elephant populations of Mesopotamia and China declined quickly because of deforestation and human population growth: by c. 850 BC the Mesopotamian elephants were extinct, and by c. 500 BC the Chinese elephants were seriously reduced in numbers and limited to areas well south of the Yellow River.
Wildcat
The earliest evidence of wildcat taming comes from 9,500-year-old Neolithic graves excavated in Shillourokambos, Cyprus, that contained the skeletons of a human and a cat, buried close to one another. As no records of native cats in Cyprus exist, this discovery indicates that Neolithic farmers brought cats to Cyprus from the Middle East, most likely to control rodents. Wildcats were probably domesticated in the Fertile Crescent around the time of the introduction of agriculture.
Neolithic Revolution
The Middle East served as the source for many animals that could be domesticated, such as sheep, goats and pigs. This area was also the first region to domesticate the dromedary. Henri Fleisch discovered and termed the Shepherd Neolithic flint industry from the Bekaa Valley in Lebanon and suggested that it could have been used by the earliest nomadic shepherds. He dated this industry to the Epipaleolithic or Pre-Pottery Neolithic as it is evidently not Paleolithic, Mesolithic or even Pottery Neolithic. The presence of these animals gave the region a large advantage in cultural and economic development. As the climate in the Middle East changed and became drier, many of the farmers were forced to leave, taking their domesticated animals with them. It was this massive emigration from the Middle East that would later help distribute these animals to the rest of Afroeurasia. This emigration was mainly on an east-west axis of similar climates, as crops usually have a narrow optimal climatic range outside of which they cannot grow for reasons of light or rain changes. For instance, wheat does not normally grow in tropical climates, just like tropical crops such as bananas do not grow in colder climates. Some authors, like Jared Diamond, have postulated that this East-West axis is the main reason why plant and animal domestication spread so quickly from the Fertile Crescent to the rest of Eurasia and North Africa, while it did not reach through the North-South axis of Africa to reach the Mediterranean climates of South Africa, where temperate crops were successfully imported by ships in the last 500 years. Similarly, the African Zebu of central Africa and the domesticated bovines of the fertile-crescent — separated by the dry sahara desert — were not introduced into each other's region.
Milk
From Southwest Asia domestic dairy animals spread to Europe (beginning around 7000 BC but did not reach Britain and Scandinavia until after 4000 BC), and South Asia (7000–5500 BC). The first farmers in central Europe and Britain milked their animals. Pastoral and pastoral nomadic economies, which rely predominantly or exclusively on domestic animals and their products rather than crop farming, were developed as European farmers moved into the Pontic-Caspian steppe in the fourth millennium BC, and subsequently spread across much of the Eurasian steppe. Sheep and goats were introduced to Africa from Southwest Asia, but African cattle may have been independently domesticated around 7000–6000 BC. Camels, domesticated in central Arabia in the fourth millennium BC, have also been used as dairy animals in North Africa and the Arabian Peninsula. The earliest Egyptian records of burn treatments describe burn dressings using milk from mothers of male babies. In the rest of the world (i.e., East and Southeast Asia, the Americas and Australia) milk and dairy products were historically not a large part of the diet, either because they remained populated by hunter-gatherers who did not keep animals or the local agricultural economies did not include domesticated dairy species. Milk consumption became common in these regions comparatively recently, as a consequence of European colonialism and political domination over much of the world in the last 500 years.
Chicken
The traditional view is that chickens were first domesticated for cockfighting in Asia, Africa, and Europe. In the last decade, there have been a number of genetic studies to clarify the origins. According to one early study, a single domestication event which took place in what now is the country of Thailand gave rise to the modern chicken with minor transitions separating the modern breeds. However, that study was later found to be based on incomplete data, and recent studies point to multiple maternal origins, with the clade found in the Americas, Europe, Middle East, and Africa, originating from the Indian subcontinent, where a large number of unique haplotypes occur. It is postulated that the red junglefowl, known as the bamboo fowl in many Southeast Asian languages, is a special bird well-adapted to take advantage of the large amounts of fruits that are produced during the end of the 50-year bamboo seeding cycle, to boost its own reproduction. In domesticating the chicken, humans took advantage of this predisposition for prolific reproduction of the red junglefowl when exposed to large amounts of food.
Camel
Martin Heide's 2010 work on the domestication of the camel tentatively concludes that humans had domesticated the Bactrian camel by at least the middle of the third millennium somewhere east of the Zagros Mountains, with the practice then moving into Mesopotamia. Heide suggests that mentions of camels "in the patriarchal narratives may refer, at least in some places, to the Bactrian camel", while noting that the camel is not mentioned in relationship to Canaan.
Dog breed
The domestic dog is the first species, and the only large carnivore, to have been domesticated. Over the past 200 years, dogs have undergone rapid phenotypic change and were formed into today's modern breeds due to artificial selection imposed by humans. These breeds can vary in size and weight from a teacup poodle to a giant mastiff. The skull, body, and limb proportions vary significantly between breeds, with dogs displaying more phenotypic diversity than can be found within the entire order of carnivores. Some breeds demonstrate outstanding skills in herding, retrieving, scent detection, and guarding, which demonstrates the functional and behavioral diversity of dogs. The first dogs were certainly wolflike, however the phenotypic changes that coincided with the dog–wolf genetic divergence away from each other are not known.
Domestication of animals
Domestication should not be confused with taming. Taming is the conditioned behavioral modification of a wild-born animal when its natural avoidance of humans is reduced and it accepts the presence of humans, but domestication is the permanent genetic modification of a bred lineage that leads to an inherited predisposition toward humans. Human selection included tameness, but without a suitable evolutionary response then domestication was not achieved. Domestic animals need not be tame in the behavioral sense, such as the Spanish fighting bull. Wild animals can be tame, such as a hand-raised cheetah. A domestic animal's breeding is controlled by humans and its tameness and tolerance of humans is genetically determined. However, an animal merely bred in captivity is not necessarily domesticated. Tigers, gorillas, and polar bears breed readily in captivity but are not domesticated. Asian elephants are wild animals that with taming manifest outward signs of domestication, yet their breeding is not human controlled and thus they are not true domesticates.
Domestication of the horse
The oldest possible archaeological indicator of a changed relationship between horses and humans is the appearance about 4800-4400 BCE of horse bones and carved images of horses in Chalcolithic graves of the early Khvalynsk culture and the Samara culture in the middle Volga region of Russia. At the Khvalynsk cemetery near the town of Khvalynsk, 158 graves of this period were excavated. Of these, 26 graves contained parts of sacrificed domestic animals, and additional sacrifices occurred in ritual deposits on the original ground surface above the graves. Ten graves contained parts of lower horse legs; two of these also contained the bones of domesticated cattle and sheep. At least 52 domesticated sheep or goats, 23 domesticated cattle, and 11 horses were sacrificed at Khvalynsk. The inclusion of horses with cattle and sheep and the exclusion of obviously wild animals together suggest that horses were categorized symbolically with domesticated animals.
Self-domestication
While humans may have intentionally domesticated wolves into dogs, an alternate hypothesis is that wolves effectively domesticated themselves by establishing a mutually beneficial relationship with prehistoric humans. They scavenged on the remains of the prey animals left by the prehistoric people at the human settlements or the kill sites. Those wolves that were less anxious and aggressive thrived, continued to follow the prehistoric humans, and colonized the human-dominated environments, generation after generation. Gradually, the first primitive dogs emerged from this group.
Goat
Goats are among the earliest animals domesticated by humans. The most recent genetic analysis confirms the archaeological evidence that the wild Bezoar ibex of the Zagros Mountains is the likely original ancestor of probably all domestic goats today.
Origin of the domestic dog
Humans and wolves both exist in complex social groups. How humans and wolves got together remains unknown. One view holds that domestication as a process that is difficult to define. The term was developed by anthropologists with a human-centric view in which humans took wild animals (ungulates) and bred them to be "domestic", usually in order to provide improved food or materials for human consumption. That term may not be appropriate for a large carnivore such as the dog. This alternate view regards dogs as being either socialized and able to live among humans, or unsocialized. There exists today dogs that live with their human families but are unsocialized and will threaten strangers defensively and aggressively no different from a wild wolf. There also exists a number of cases where wild wolves have approached people in remote places, attempting to initiate play and to form companionship. One such notable wolf was Romeo, a gentle black wolf that formed relationships with the people and dogs of Juneau, Alaska. This view holds that before there could have been domestication of the wolf, there had to have been its socialization.
Red junglefowl
In 2012, a study examined mitochondrial DNA recovered from ancient bones from Europe, Thailand, the Pacific, and Chile, and from Spanish colonial sites in Florida and the Dominican Republic, in directly dated samples originating in Europe at 1,000 BP and in the Pacific at 3,000 BP. The study showed that chickens were most likely domesticated from wild red junglefowl, though some have suggested possible genetic contributions from other junglefowl species. Domestication occurred at least 7,400 years ago from a common ancestor flock in the bird's natural range, then proceeded in waves both east and west. The earliest undisputed domestic chicken remains are bones associated with a date around 7,400 BP from the Chishan site, in the Hebei province of China. In the Ganges region of India, red junglefowl were being used by humans as early as 7,000 years ago. No domestic chicken remains older than 4,000 years have been identified in the Indus Valley, and the antiquity of chickens recovered from excavations at Mohenjodaro is still debated.
Dromedary
The dromedary was probably first domesticated in Somalia or the Arabian Peninsula about 4,000 years ago. In the ninth or tenth century BC, the dromedary became popular in the Near East. The Persian invasion of Egypt under Cambyses in 525 BC introduced domesticated camels to the area. The Persian camels were not well-suited to trading or travel over the Sahara; journeys across the desert were made on chariots pulled by horses. The dromedary was introduced into Egypt from south-western Asia (Arabia and Persia). The popularity of dromedaries increased after the Islamic conquest of North Africa. While the invasion was accomplished largely on horseback, new links to the Middle East allowed camels to be imported "en masse". These camels were well-suited to long desert journeys and could carry a great deal of cargo, allowing substantial trans-Saharan trade for the first time. In Libya, dromedaries were used for transport and their milk and meat constituted the local diet.
History of genetic engineering
Genetic recombination caused by human activity has been occurring since around 12,000 BC, when humans first began to domesticate organisms. Genetic engineering as the direct transfer of DNA from one organism to another was first accomplished by Herbert Boyer and Stanley Cohen in 1972. It was the result of a series of advancements in techniques that allowed the direct modification of the genome. Important advances included the discovery of restriction enzymes and DNA ligases, the ability to design plasmids and technologies like polymerase chain reaction and sequencing. Transformation of the DNA into a host organism was accomplished with the invention of biolistics, "Agrobacterium"-mediated recombination and microinjection.
The first genetically modified animal was a mouse created in 1974 by Rudolf Jaenisch. In 1976 the technology was commercialised, with the advent of genetically modified bacteria that produced somatostatin, followed by insulin in 1978. In 1983 an antibiotic resistant gene was inserted into tobacco, leading to the first genetically engineered plant. Advances followed that allowed scientists to manipulate and add genes to a variety of different organisms and induce a range of different effects. Plants were first commercialized with virus resistant tobacco released in China in 1992. The first genetically modified food was the Flavr Savr tomato marketed in 1994. By 2010, 29 countries had planted commercialized biotech crops. In 2000 a paper published in "Science" introduced golden rice, the first food developed with increased nutrient value.
History of the domestic sheep
Sheep were among the first animals to be domesticated by humankind (although the domestication of dogs may be over 20,000 years earlier); the domestication date is estimated to fall between 11,000 and 9,000 B.C in Mesopotamia. Their wild relatives have several characteristics, such as a relative lack of aggression, a manageable size, early sexual maturity, a social nature, and high reproduction rates, which made them particularly suitable for domestication. Today, "Ovis aries" is an entirely domesticated animal that is largely dependent on humans for its health and survival. Feral sheep do exist, but exclusively in areas devoid of large predators (usually islands) and not on the scale of feral horses, goats, pigs, or dogs, although some feral populations have remained isolated long enough to be recognized as distinct breeds.
Red junglefowl
The red junglefowl ("Gallus gallus") is a tropical member of the family Phasianidae. It is the primary progenitor of the domestic chicken (though genetic evidence strongly suggests some past hybridisation with the grey junglefowl, as well). The red junglefowl was first domesticated at least 5000 years ago in Asia. Since then, its domestic form has spread around the world and is kept globally as a very productive food source of both meat and eggs.
House sparrow
The house sparrow was among the first animals to be given a scientific name in the modern system of biological classification, since it was described by Carl Linnaeus, in the 1758 10th edition of "Systema Naturae". It was described from a type specimen collected in Sweden, with the name "Fringilla domestica". Later, the genus name "Fringilla" came to be used only for the common chaffinch and its relatives, and the house sparrow has usually been placed in the genus "Passer" created by French zoologist Mathurin Jacques Brisson in 1760.
Domestic rabbit
Rabbits were first exploited by the Romans as sources of food and fur, and have been kept as pets in Western nations since the 19th century. Beginning in the 1980s, the idea of the domestic rabbit as a house companion, or house rabbit, was promoted. Rabbits can be litter box trained and may come when called, but they need exercise and can damage a house that is not "rabbit proof". Especially following the Easter season, unwanted rabbits end up in animal shelters. Because they have become invasive in Australia, pet rabbits are banned in Queensland.
Origin of the domestic dog
In 2002, a study proposed that immediate human ancestors and wolves may have domesticated each other through a strategic alliance that would change both respectively into humans and dogs. The effects of human psychology, hunting practices, territoriality and social behavior would have been profound. Wolves and dogs mark their territory with scent; however, humans do not have a keen sense of smell and may have begun to mark their territories with symbols, which became the first art and may have been generative of human culture. Wolves hunt large game; however, there is no evidence of pre-"sapiens" hunting large game. Early humans moved from scavenging and small-game hunting to big-game hunting by living in larger, socially more-complex groups, learning to hunt in packs, and developing powers of cooperation and negotiation in complex situations. As these are characteristics of wolves, dogs and humans, it can be argued that these behaviors were enhanced once wolves and humans began to cohabit. Communal hunting led to communal defense. Wolves actively patrol and defend their scent-marked territory, and perhaps humans had their sense of territoriality enhanced by living with wolves. One of the keys to recent human survival has been the forming of partnerships. Strong bonds exist between same-sex wolves, dogs and humans and these bonds are stronger than exist between other same-sex animal pairs. Today, the most widespread form of inter-species bonding occurs between humans and dogs. The concept of friendship has ancient origins but it may have been enhanced through the inter-species relationship to give a survival advantage.
Animal husbandry
The first wild animal to be domesticated was the dog. Half-wild dogs, perhaps starting with young individuals, may have been tolerated as scavengers and killers of vermin, and being naturally pack hunters, were predisposed to become part of the human pack and join in the hunt. Prey animals, sheep, goats, pigs and cattle, were progressively domesticated early in the history of agriculture.
Domestic pig
DNA evidence from subfossil remains of teeth and jawbones of Neolithic pigs shows that the first domestic pigs in Europe had been brought from the Near East. This stimulated the domestication of local European wild boar, resulting in a third domestication event with the Near Eastern genes dying out in European pig stock. Modern domesticated pigs have involved complex exchanges, with European domesticated lines being exported, in turn, to the ancient Near East. Historical records indicate that Asian pigs were introduced into Europe during the 18th and early 19th centuries.
Guns, Germs, and Steel
As early Western Asian civilizations began to trade, they found additional useful animals in adjacent territories, most notably horses and donkeys for use in transport. Diamond identifies 13 species of large animals over domesticated in Eurasia, compared with just one in South America (counting the llama and alpaca as breeds within the same species) and none at all in the rest of the world. Australia and North America suffered from a lack of useful animals due to extinction, probably by human hunting, shortly after the end of the Pleistocene, whilst the only domesticated animals in New Guinea came from the East Asian mainland during the Austronesian settlement some 4,000–5,000 years ago. Sub-Saharan biological relatives of the horse including zebras and onagers proved untameable; and although African elephants can be tamed, it is very difficult to breed them in captivity; Diamond describes the small number of domesticated species (14 out of 148 "candidates") as an instance of the Anna Karenina principle: many promising species have just one of several significant difficulties that prevent domestication. He also makes the intriguing argument that all large mammals that could be domesticated, have been.
Mallard
Mallards have often been ubiquitous in their regions among the ponds, rivers, and streams of human parks, farms, and other man-made waterwayseven to the point of visiting water features in human courtyards.
Mallards have had a long relationship with humans. Almost all domestic duck breeds derive from the mallard, with the exception of a few Muscovy breeds. Mallards are generally monogamous while domestic ducks are mostly polygamous. Domestic ducks have no territorial behaviour and are less aggressive than mallards. Domestic ducks are mostly kept for meat, as their eggs have a strong flavor. They were first domesticated in Southeast Asia at least 4,000 years ago, during the Neolithic Age, and were also farmed by the Romans in Europe, and the Malays in Asia. It is common for mallards to mate with domestic ducks and produce hybrid offspring that are fully fertile. Because of this, mallards have been found to be contaminated with the genes of the domestic duck.
House mouse
The first written reference to mice kept as pets occurs in the "Erya", the oldest extant Chinese dictionary, from a mention in an 1100 B.C. version. Human domestication led to numerous strains of "fancy" or hobby mice with a variety of colours and a docile temperament. Domestic varieties of the house mouse are bred as a food source for some carnivorous pet reptiles, birds, arthropods, and fish. The effects of domestication can be rapid, with captive-reared mice differing in boldness and activity patterns compared to wild-caught mice after 4-5 generations in recent research.
Dog behavior
The origin of the domestic dog ("Canis lupus familiaris" or "Canis familiaris") is not clear. Whole genome sequencing indicates that the dog, the gray wolf and the extinct Taymyr wolf diverged at around the same time 27,000-40,000 years ago. How dogs became domesticated is not clear, however the two main hypotheses are self-domestication or human domestication. There exists evidence of human-canine behavioral coevolution.
Domestication
Domesticated birds principally mean poultry, raised for meat and eggs: some Galliformes (chicken, turkey, guineafowl) and Anseriformes (waterfowl: duck, goose, swan). Also widely domesticated are cagebirds such as songbirds and parrots; these are kept both for pleasure and for use in research.
Domestic pigeon is known as a messenger, research suggests it was domesticated as early as 10,000 years ago.
Chickens were domesticated at least 7,000 years ago, with fossils in China from c. 5400 BCE. The chicken's wild ancestor is "Gallus gallus", the red junglefowl of Southeast Asia, and another species, probably the grey junglefowl of India. It appears to have been kept initially for cockfighting rather than for food.
Origin of the domestic dog
Where the genetic divergence of dog and wolf took place remains controversial, with the most plausible proposals spanning Western Europe, Central Asia, and East Asia. This has been made more complicated by the most recent proposal that fits the available evidence, which is that an initial wolf population split into East and West Eurasian wolves, these were then domesticated independently before going extinct into two distinct dog populations between 14,000-6,400 years ago, and then the Western Eurasian dog population was partially and gradually replaced by East Asian dogs that were brought by humans at least 6,400 years ago.
Meat
The domestication of animals, of which we have evidence dating back to the end of the last glacial period (c. 10,000 BCE), allowed the systematic production of meat and the breeding of animals with a view to improving meat production. The animals which are now the principal sources of meat were domesticated in conjunction with the development of early civilizations:Other animals are or have been raised or hunted for their flesh. The type of meat consumed varies much between different cultures, changes over time, depending on factors such as tradition and the availability of the animals. The amount and kind of meat consumed also varies by income, both between countries and within a given country.
Origin of the domestic dog
Where the genetic divergence of dog and wolf took place remains controversial, with the most plausible proposals spanning Western Europe, Central Asia and East Asia. This has been made more complicated by the most recent proposal that fits the available evidence, which is that an initial wolf population split into East and West Eurasian groups; these, before going extinct, were domesticated independently into two distinct dog populations between 14,000 and 6,400 years ago. The Western Eurasian dog population was partially and gradually replaced by East Asian dogs introduced by humans at least 6,400 years ago.
Animal husbandry
Husbandry has a long history, starting with the Neolithic revolution when animals were first domesticated, from around 13,000 BC onwards, antedating farming of the first crops. By the time of early civilisations such as ancient Egypt, cattle, sheep, goats and pigs were being raised on farms.
Megan and Morag
Megan and Morag, two domestic sheep, were the first mammals to have been successfully cloned from differentiated cells. They are not to be confused with Dolly the sheep which was the first animal to be successfully cloned from an adult somatic cell or Polly the sheep which was the first cloned and transgenic animal. Megan and Morag, like Dolly and Polly, were cloned at the Roslin Institute in Edinburgh, Scotland in 1995.
Quagga
The quagga was long regarded a suitable candidate for domestication, as it counted as the most docile of the striped horses. The earliest Dutch colonists in South Africa had already fantasized about this possibility, because their imported work horses did not perform very well in the extreme climate and regularly fell prey to the feared African horse sickness. In 1843, the English naturalist Charles Hamilton Smith wrote that the quagga was 'unquestionably best calculated for domestication, both as regards strength and docility'. Only a few descriptions have been given of tame or domesticated quaggas in South Africa. In Europe, the only confirmed cases are two stallions driven in a phaeton by Joseph Wilfred Parkins, sheriff of London in 1819-1820, and the quaggas and their hybrid offspring of London Zoo, which were used to pull a cart and transport vegetables from the market to the zoo. Nevertheless, the reveries continued long after the death of the last quagga in 1883. In 1889, the naturalist Henry Bryden wrote: "That an animal so beautiful, so capable of domestication and use, and to be found not long since in so great abundance, should have been allowed to be swept from the face of the earth, is surely a disgrace to our latter-day civilization." 
The specimen in London died in 1872 and the one in Berlin in 1875. The last captive quagga, a female in Amsterdam's Natura Artis Magistra zoo, lived there from 9 May 1867 until it died on 12 August 1883, but its origin and cause of death are unclear. Its death was not recognised as signifying the extinction of its kind at the time, and the zoo requested another specimen; hunters believed it could still be found "closer to the interior" in the Cape Colony. Since locals used the term quagga to refer to all zebras, this may have led to the confusion. The extinction of the quagga was internationally accepted by the 1900 Convention for the Preservation of Wild Animals, Birds and Fish in Africa. The last specimen was featured on a Dutch stamp in 1988. There are 23 known stuffed and mounted quagga specimens throughout the world, including a juvenile, two foals, and a foetus. In addition, a mounted head and neck, a foot, seven complete skeletons, and samples of various tissues remain. A 24th mounted specimen was destroyed in Königsberg, Germany, during World War II, and various skeletons and bones have also been lost.
Domestication of Ficus carica
Ficus carica, known as the common fig, is one of the earliest plants to have ever undergone domestication. "Ficus" "carica" played many important roles in history and has become a symbol in many different cultures and religions. They continue to play a significant part in the societies and economies of today, as they have proven to serve a multitude of uses.
Capra (genus)
Capra is a genus of mammals, the goats, composed of up to nine species, including the wild goat, the markhor, and several species known as ibexes. The domestic goat ("Capra aegagrus hircus") is a domesticated subspecies of the wild goat ("Capra aegagrus"). Evidence of goat domestication dates back more than 8,500 years.
Origin of the domestic dog
There is a genetic difference between domestic and wild populations. There is also such a difference between the domestication traits that researchers believe to have been essential at the early stages of domestication, and the improvement traits that have appeared since the split between wild and domestic populations. Domestication traits are generally fixed within all domesticates, and were selected during the initial episode of domestication of that animal or plant, whereas improvement traits are present only in a proportion of domesticates, though they may be fixed in individual breeds or regional populations. The dog was the first domesticant and was established across Eurasia before the end of the Late Pleistocene era, well before cultivation and before the domestication of other animals.
Domestic duck
Mallard ducks were first domesticated in Southeast Asia at least 4000 years ago, during the Neolithic Age, and were also farmed by the Romans in Europe, and the Malays in Asia. In ancient Egypt, ducks were captured in nets and then bred in captivity. During the Ming Dynasty, the Peking duck—mallards force-fed on grains, making them larger— was known to have good genetic characteristics.
Equus (genus)
The earliest archaeological evidence for the domestication of the horse comes from sites in Ukraine and Kazakhstan, dating to around 4000-3500 BC. By 3000 BC, the horse was completely domesticated, and by 2000 BC, a sharp increase occurred in the number of horse bones found in human settlements in northwestern Europe, indicating the spread of domesticated horses throughout the continent. The most recent, but most irrefutable, evidence of domestication comes from sites where horse remains were buried with chariots in graves of the Sintashta and Petrovka cultures c. 2100 BC. Studies of variation in genetic material shows that very few wild stallions, possibly all from a single haplotype, contributed to the domestic horse, while many mares were part of early domesticated herds.
Domestication of animals
The commensal pathway was traveled by vertebrates that fed on refuse around human habitats or by animals that preyed on other animals drawn to human camps. Those animals established a commensal relationship with humans in which the animals benefited but the humans received no harm but little benefit. Those animals that were most capable of taking advantage of the resources associated with human camps would have been the tamer, less aggressive individuals with shorter fight or flight distances. Later, these animals developed closer social or economic bonds with humans that led to a domestic relationship. The leap from a synanthropic population to a domestic one could only have taken place after the animals had progressed from anthropophily to habituation, to commensalism and partnership, when the relationship between animal and human would have laid the foundation for domestication, including captivity and human-controlled breeding. From this perspective, animal domestication is a coevolutionary process in which a population responds to selective pressure while adapting to a novel niche that included another species with evolving behaviors.
Archaeology and the Book of Mormon
There are six references to "cattle" made in the Book of Mormon, including verbiage suggesting they were domesticated. While the Book of Mormon may follow the common biblical precedent of referring to all domesticated animals as "cattle", there is no evidence that Old World cattle (members of the genus "Bos") inhabited the New World prior to European contact in the 16th century AD. Further, there is currently no archeological evidence of American bison having been domesticated. It is widely accepted that the only large mammal to be domesticated in the Americas was the llama and that no species of goats, deer, or sheep were domesticated before the arrival of the Europeans to the continent.
Texas A&amp;M College of Veterinary Medicine &amp; Biomedical Sciences
The College's research into animal cloning is one of the more publicized ventures. Texas A&M scientists created the first cloned domestic animal, a cat named "CC (cat)", on December 22, 2001. Texas A&M was also the first academic institution to clone each of six different species: cattle, a goat, pigs, a cat, a deer and a horse.
Turkey (bird)
The species "Meleagris gallopavo" is used by humans for their meat. They were first domesticated by the indigenous people of Mexico from at least 800 BC onwards. These domesticates were then either introduced into what is now the US Southwest or independently domesticated a second time by the indigenous people of that region by 200 BC, at first being used for their feathers, which were used in ceremonies and to make robes and blankets. Turkeys were first used for meat by Native Americans by about AD 1100. Compared to wild turkeys, domestic turkeys are selectively bred to grow larger in size for their meat. Americans often eat turkey on special occasions such as at Thanksgiving or Christmas.
Shillourokambos
Historians previously accounted Egypt as the earliest site of cat domestication due to the clear depictions of house cats in ancient Egyptian paintings about 3,600 years old. However, in 2004, a Neolithic grave was excavated in Shillourokambos, Cyprus that contained skeletons, laid close to one another, of both a human and a cat. The grave is estimated to be 9,500 years old, pushing back the earliest known feline-human association significantly. The cat specimen is large and closely resembles the African wildcat ("Felis silvestris lybica"), rather than present-day domestic cats.
Western honey bee
Humans have been collecting honey from bees for thousands of years, with evidence in the form of rock art found in France and Spain, dating to around 7000 BCE. The honey bee is one of the few invertebrate animals to have been domesticated. Bees were likely first domesticated in ancient Egypt, where tomb paintings depict bee-keeping. Europeans brought bees to North America in 1622.
Working animal
The history of working animals may predate agriculture, with dogs used by our hunter-gatherer ancestors. Around the world, millions of animals work in relationship with their owners. Domesticated species are often bred for different uses and conditions, especially horses and working dogs. Working animals are usually raised on farms, though some are still captured from the wild, such as dolphins and some Asian elephants.
History of biology
The earliest humans must have had and passed on knowledge about plants and animals to increase their chances of survival. This may have included knowledge of human and animal anatomy and aspects of animal behavior (such as migration patterns). However, the first major turning point in biological knowledge came with the Neolithic Revolution about 10,000 years ago. Humans first domesticated plants for farming, then livestock animals to accompany the resulting sedentary societies.