Aldabrachelys gigantea hololissa
Giant tortoises are among the longest-lived animals on the planet. Some individual Aldabra giant tortoises are thought to be over 200 years of age, but this is difficult to verify because they tend to outlive their human observers. Adwaita was reputedly one of four brought by British seamen from the Seychelles Islands as gifts to Robert Clive of the British East India Company in the 18th century, and came to Calcutta Zoo in 1875. At his death in March 2006 at the Kolkata (formerly Calcutta) Zoo in India, Adwaita is reputed to have reached the longest ever measured life span of 255 years (birth year 1750). Today, Jonathan, a -year-old, Saint Helena-dwelling Seychelles giant tortoise, is thought to be the oldest living terrestrial animal on Earth since the 2006 death of Harriet; a 176 year old Galapagos giant tortoise who lived at Australia Zoo. Coming in second to Jonathan is Esmeralda, at years old. Esmeralda is an Aldabra giant tortoise that lives in the Galapagos Islands.
Galápagos tortoise
Phylogenetic analysis may help to "resurrect" the extinct species of Floreana ("C. nigra") – a species known only from subfossil remains. Some tortoises from Isabela were found to be a partial match for the genetic profile of Floreana specimens from museum collections, possibly indicating the presence of hybrids from a population transported by humans from Floreana to Isabela, resulting either from individuals deliberately transported between the islands, or from individuals thrown overboard from ships to lighten the load. Nine Floreana descendants have been identified in the captive population of the Fausto Llerena Breeding Center on Santa Cruz; the genetic footprint was identified in the genomes of hybrid offspring. This allows the possibility of re-establishing a reconstructed species from selective breeding of the hybrid animals. Furthermore, individuals from the species possibly are still extant. Genetic analysis from a sample of tortoises from Volcan Wolf found 84 first-generation "C. nigra" hybrids, some less than 15 years old. The genetic diversity of these individuals is estimated to have required 38 "C. nigra" parents, many of which could still be alive on Isabela Island.