Pteranodon
While the first "Pteranodon" wing bones were collected by Marsh and Cope in the early 1870s, the first "Pteranodon" skull was found on May 2, 1876, along the Smoky Hill River in Wallace County (now Logan County), Kansas, USA, by Samuel Wendell Williston, a fossil collector working for Marsh. A second, smaller skull soon was discovered as well. These skulls showed that the North American pterosaurs were different from any European species, in that they lacked teeth and had bony crests on their skulls. Marsh recognized this major difference, describing the specimens as "distinguished from all previously known genera of the order Pterosauria by the entire absence of teeth." Marsh recognized that this characteristic warranted a new genus, and he coined the name "Pteranodon" ("wing without tooth") in 1876. Marsh reclassified all the previously named North American species from "Pterodactylus" to "Pteranodon". He considered the smaller skull to belong to "Pteranodon occidentalis", based on its size. Marsh classified the larger skull, YPM 1117, in the new species "Pteranodon longiceps", which he thought to be a medium-sized species in between the small "P. occidentalis" and the large "P. ingens". Marsh also named several additional species: Pteranodon comptus and "Pteranodon nanus" were named for fragmentary skeletons of small individuals, while "Pteranodon gracilis" was based on a wing bone that he mistook for a pelvic bone. He soon realized his mistake, and re-classified that specimen again into a separate genus, which he named "Nyctosaurus". "P, nanus" was also later recognized as a "Nyctosaurus" specimen.
Gannat
The most ancient discoveries in Gannat, ancestors of the rhinoceros from the end of the Oligocene and start of the Miocene, date back 23 million years. Gannat seems to have been a veritable cemetery for these creatures, so many of their remains have been discovered. Also found are fossils of fish, reptiles, tortoises, crocodiles, galliform birds, mammals, marsupials, insectivores, rodents and carnivores. The site is particularly rich in rhinoceros. From 1854, Duvernoy has even described a specimin as "Acerotherium gannatense" (the official name is "Diaceratherium lemanense"). The most complete rhinoceros skeleton was discovered in 1993 when significant fossils were uncovered at the Sichaux quarry by paleontologist François Escuillié (notable discoverer of a small proboscidian mammal, ancestor of the elephant from 50 million years ago). Escuillié was behind the establishment of the Rhinopolis Association in 1994 and founder of Eldonia, a society for those specialising in the renovation of fossils and whose activities are linked to Rhinopolis. The Rhinopolis Association is still active in the Gannat quarries and, since the 1990s, has discovered numerous rhinoceros bones.