Pteranodon
While the first "Pteranodon" wing bones were collected by Marsh and Cope in the early 1870s, the first "Pteranodon" skull was found on May 2, 1876, along the Smoky Hill River in Wallace County (now Logan County), Kansas, USA, by Samuel Wendell Williston, a fossil collector working for Marsh. A second, smaller skull soon was discovered as well. These skulls showed that the North American pterosaurs were different from any European species, in that they lacked teeth and had bony crests on their skulls. Marsh recognized this major difference, describing the specimens as "distinguished from all previously known genera of the order Pterosauria by the entire absence of teeth." Marsh recognized that this characteristic warranted a new genus, and he coined the name "Pteranodon" ("wing without tooth") in 1876. Marsh reclassified all the previously named North American species from "Pterodactylus" to "Pteranodon". He considered the smaller skull to belong to "Pteranodon occidentalis", based on its size. Marsh classified the larger skull, YPM 1117, in the new species "Pteranodon longiceps", which he thought to be a medium-sized species in between the small "P. occidentalis" and the large "P. ingens". Marsh also named several additional species: Pteranodon comptus and "Pteranodon nanus" were named for fragmentary skeletons of small individuals, while "Pteranodon gracilis" was based on a wing bone that he mistook for a pelvic bone. He soon realized his mistake, and re-classified that specimen again into a separate genus, which he named "Nyctosaurus". "P, nanus" was also later recognized as a "Nyctosaurus" specimen.
Paleontology in Kansas
Paleontology in Kansas refers to paleontological research occurring within or conducted by people from the U.S. state of Kansas. Kansas has been the source of some of the most spectacular fossil discoveries in US history. The fossil record of Kansas spans from the Cambrian to the Pleistocene. From the Cambrian to the Devonian, Kansas was covered by a shallow sea. During the ensuing Carboniferous the local sea level began to rise and fall. When sea levels were low the state was home to richly vegetated deltaic swamps where early amphibians and reptiles lived. Seas expanded across most of the state again during the Permian, but on land the state was home to thousands of different insect species. The popular pterosaur "Pteranodon" is best known from this state. During the early part of the Cenozoic era Kansas became a savannah environment. Later, during the Ice Age, glaciers briefly entered the state, which was home to camels, mammoths, mastodons, and saber-teeth. Local fossils may have inspired Native Americans to regard some local hills as the homes of sacred spirit animals. Major scientific discoveries in Kansas included the pterosaur "Pteranodon" and a fossil of the fish "Xiphactinus" that died in the act of swallowing another fish.