Doug Pederson - wikipedia Doug Pederson Jump to : navigation , search Not to be confused with Doug Peterson . Doug Pederson Pederson in 2016 Philadelphia Eagles Position : Head coach Born : ( 1968 - 01 - 31 ) January 31 , 1968 ( age 50 ) Bellingham , Washington Height : 6 ft 3 in ( 1.91 m ) Weight : 221 lb ( 100 kg ) Career information High school : Ferndale ( WA ) College : Northeast Louisiana Undrafted : 1991 Career history As player : Miami Dolphins ( 1991 ) * New York / New Jersey Knights ( 1992 ) Miami Dolphins ( 1992 -- 1994 ) Carolina Panthers ( 1995 ) * Rhein Fire ( 1995 ) Miami Dolphins ( 1995 ) Green Bay Packers ( 1995 -- 1998 ) Philadelphia Eagles ( 1999 ) Cleveland Browns ( 2000 ) Green Bay Packers ( 2001 -- 2004 ) * Offseason and / or practice squad member only As coach : Calvary Baptist Academy ( 2005 -- 2008 ) Head coach Philadelphia Eagles ( 2009 -- 2010 ) Offensive quality control coordinator Philadelphia Eagles ( 2011 -- 2012 ) Quarterbacks coach Kansas City Chiefs ( 2013 -- 2015 ) Offensive coordinator Philadelphia Eagles ( 2016 -- present ) Head coach Career highlights and awards As player Super Bowl champion ( XXXI ) As coach Super Bowl champion ( LII ) Career NFL statistics Pass attempts : 522 Pass completions : 286 Percentage : 54.7 TD -- INT : 12 -- 19 Passing yards : 2,762 Passer rating : 62.3 Player stats at NFL.com Head coaching record Regular season : 20 -- 12 (. 625 ) Postseason : 3 -- 0 ( 1.000 ) Career : 23 -- 12 (. 657 ) Coaching stats at PFR Douglas Irving Pederson ( born January 31 , 1968 ) is an American football coach who is currently the head coach of the Philadelphia Eagles of the National Football League ( NFL ) . He served as the offensive coordinator of the Kansas City Chiefs from 2013 -- 2015 . He spent most of his playing career as a member of the Green Bay Packers , serving as a backup quarterback to Brett Favre and holder on placekicks , and winning Super Bowl XXXI with the team over the New England Patriots . He was also a backup to Dan Marino as a member of the Miami Dolphins , and a starting quarterback for the Eagles and Cleveland Browns . In his second season as the Eagles ' head coach , Pederson won Super Bowl LII ( also against the Patriots ) , marking the first Super Bowl title in franchise history . He also became just the fourth person , after Mike Ditka , Tom Flores , and Tony Dungy to win a Super Bowl as both a player and head coach . Contents ( hide ) 1 Playing career 1.1 Early years 1.2 Professional 1.2. 1 Miami Dolphins 1.2. 2 First stint with Packers 1.2. 3 Philadelphia Eagles 1.2. 4 Cleveland Browns 1.2. 5 Green Bay Packers ( second stint ) 2 Coaching career 2.1 High school 2.2 NFL assistant coaching positions 2.3 NFL head coach 2.4 Head coaching record 2.5 Coaching tree 3 Personal life 4 References 5 External links Playing career ( edit ) Early years ( edit ) Pederson was born in Bellingham , Washington , in 1968 . He attended Ferndale High School in nearby Ferndale , Washington , and was an All - State selection in football , basketball , and baseball . After high school he graduated from Northeast Louisiana University , where he was quarterback from 1987 through 1990 . He still holds multiple passing records at the school . Professional ( edit ) Miami Dolphins ( edit ) Pederson originally signed as a rookie free agent by the Miami Dolphins on May 1 , 1991 , out of Northeast Louisiana University ( now University of Louisiana at Monroe ) in Monroe , Louisiana . He was waived on August 17 , 1991 , before the start of the regular season . After spending the 1991 season as a free agent , the New York / New Jersey Knights of the World League of American Football ( WLAF ) drafted him in the fifth round for the first pool of draft - eligible players on February 4 , 1992 . The second pool , which was drafted from on February 20 , consisted of players allocated by NFL teams to the league . He was the backup quarterback to Reggie Slack with the Knights from March to May 1992 . After the WLAF season finished , he was re-signed by the Dolphins on June 2 , 1992 . Pederson spent 1992 training camp with the Dolphins , before being released during final roster cuts again . He was subsequently re-signed to the team 's practice squad , where he practiced on the scout team until he was waived on October 8 , 1992 . He was re-signed by the Dolphins after the season on March 3 , 1993 . After his third training camp with the Dolphins , he was waived again on August 31 , 1993 . For the second consecutive season , Pederson was re-signed to the team 's practice squad , on September 1 , 1993 . Dan Marino , the Dolphins ' starting quarterback since 1983 , ruptured his Achilles ' tendon in a week 6 game against the Cleveland Browns on October 10 , 1993 , forcing backup Scott Mitchell to replace him . Pederson replaced Marino on the active roster , and served as Mitchell 's backup for the next four games . Pederson made his NFL debut on October 24 , 1993 , in a week 8 game against the Indianapolis Colts . He helped head coach Don Shula win his NFL - record 325th victory as a coach when Mitchell suffered a separated shoulder in a week 11 game against the Philadelphia Eagles on November 14 , 1993 . In that record breaking game for Coach Shula , Pederson entered in the 3rd quarter of the game and went 3 for 6 for 34 yards , and completed several crucial 3rd downs . Pederson was able to steer the Dolphins to the win . Pederson also served as the backup to recently acquired Steve DeBerg for the three games Mitchell missed with injury . He briefly entered a week 14 game against the New York Giants while DeBerg was receiving stitches on his face . Mitchell returned as the Dolphins ' starter after week 15 , and Pederson was released in favor of backup DeBerg and third - string quarterback Hugh Millen on December 16 , 1993 . Pederson re-signed with the Dolphins on April 16 , 1994 , after the season ended . He spent the entire 1994 season on the Dolphins ' active roster as the third - string quarterback behind Marino and Bernie Kosar . On February 15 , 1995 , Pederson was selected by the Carolina Panthers in the twenty - second round of the NFL Expansion Draft , after being placed on the Dolphins ' available players list on January 19 , but was released on May 24 , 1995 . He returned to the World League after his release , playing with Rhein Fire . Pederson re-signed with the Dolphins again in June 1995 . After competing with Dan McGwire throughout training camp , Pederson was waived on August 22 , 1995 . Marino suffered a knee injury during a week 6 game , so Pederson was re-signed on October 10 to serve as the third quarterback behind Kosar and McGwire for the next two games . He was released again after Marino returned for week 9 on October 24 . First stint with Packers ( edit ) Pederson worked out for the Green Bay Packers following week 10 in 1995 , due to a season - ending injury suffered by backup Ty Detmer and a minor injury sustained by starter Brett Favre . Third - string quarterback T.J. Rubley was forced to play in week 10 and threw a game - ending interception after calling an audible , going against head coach Mike Holmgren 's playcall . The Packers signed Bob Gagliano to serve as the third - stringer quarterback for weeks 11 and 12 . Pederson replaced Gagliano as the third - string quarterback when he signed with the Packers on November 22 , 1995 . The Packers claimed Jim McMahon off waivers from the Browns to serve as Favre 's backup ahead of Pederson and Rubley on November 29 , 1995 . Rubley was waived on December 13 , leaving McMahon and Pederson as Favre 's backups . Favre did not miss any games , so Pederson did not see any game action for the Packers in 1995 . Pederson served as the third quarterback behind Favre and McMahon in 1996 , playing in one game but recorded no statistics . He received a Super Bowl ring following the Packers ' win over the Patriots in Super Bowl XXXI . He re-signed with the Packers with a two - year contract on February 20 , 1997 . Pederson was again the third quarterback throughout 1997 , backing up Favre and Steve Bono . Pederson beat out Rick Mirer for the backup job to Favre , as well as the primary placekick holder job , in 1998 . In a week 5 loss to the Minnesota Vikings , Pederson replaced Favre in the last five minutes of a blowout game , and threw two touchdowns in his place . However , Pederson suffered a broken jaw that knocked him out for the team 's next four games . Philadelphia Eagles ( edit ) Pederson signed a three - year , $4.5 million contract with the Philadelphia Eagles on February 18 , 1999 , to become the team 's starting quarterback under new head coach Andy Reid , who was Pederson 's quarterbacks coach in Green Bay from 1997 -- 1998 . The Eagles drafted Donovan McNabb with the second overall pick in the 1999 NFL Draft in April 1999 , and Reid said Pederson would remain the starter until McNabb was ready to play . In his nine starts for the Eagles , Pederson had a 2 -- 7 record , a 51.6 % completion rate , 1,168 passing yards , six touchdowns , and nine interceptions . In his first career start , a week 1 game against the Arizona Cardinals , Pederson threw two touchdowns in the first quarter to help give the Eagles a 21 -- 0 lead . The Cardinals came back , however , and won the game on a field goal as time expired , 25 -- 24 . Pederson went 12 - for - 25 for 91 yards and two touchdowns in the game . McNabb replaced Pederson , who suffered a bruised throwing shoulder , after one half in a week 2 loss to the Tampa Bay Buccaneers , in which Pederson went 12 - of - 19 for 100 yards and an interception . Pederson started a week 3 shutout loss ( 26 -- 0 ) to the Buffalo Bills , going 14 - of - 26 for 137 yards and two lost fumbles , before being replaced by McNabb again in the fourth quarter . In a week 4 loss to the New York Giants , Pederson went 6 - for - 15 for 75 yards and two interceptions before being replaced by McNabb after halftime . Pederson 's first NFL win came in week 5 in a game against the Dallas Cowboys . He played the entire game , going 11 - of - 29 for 145 yards , one touchdown , and one interception . Pederson played the entirety of the next three games , posting a 1 -- 2 record while throwing three touchdowns and three interceptions . In his final start as an Eagle , Pederson was benched at halftime of a week 9 game against the Carolina Panthers after going 3 - of - 9 for 28 yards and being down 23 -- 0 . He did not see game action at quarterback again until a week 14 game against the Cowboys in which McNabb suffered an injury in the fourth quarter . Pederson went 8 - for - 12 for 108 yards and a touchdown in the loss , and Koy Detmer received the start ahead of him in week 15 with McNabb still injured . After spending the next season 's training camp with the team , the Eagles released Pederson on August 28 , 2000 . Cleveland Browns ( edit ) Pederson considered retirement after being released by the Eagles , but instead signed a two - year contract with the Cleveland Browns on September 2 , 2000 . The Browns ' backup , Ty Detmer , suffered a season - ending injury , and the Browns needed a backup quarterback to starter Tim Couch . This was the second time in Pederson 's career that he was signed to replace an injured Ty Detmer . Pederson started as the third quarterback behind Couch and Spergon Wynn , until Couch suffered a season - ending injury in week 7 . Pederson started the next six games , posting a 1 -- 5 record . In a week 13 game against the Baltimore Ravens , he was knocked out of the game with bruised ribs and replaced with Wynn . Wynn started the next week against the Jacksonville Jaguars , but he suffered a season - ending injury and Pederson replaced him . Pederson returned for the final two games of the season , losing both , including a 35 -- 24 loss to his former team , the Eagles , and a 24 -- 0 shutout loss to the Tennessee Titans . Pederson was released after the season on February 22 , 2001 . Green Bay Packers ( second stint ) ( edit ) The Packers re-signed Pederson to a one - year contract on March 13 , 2001 , to replace backup Matt Hasselbeck , who was traded to the Seattle Seahawks . Pederson was the primary backup to Favre for the entire 2001 season , and was the primary placekick holder in every game . He was re-signed to a one - year , $650,000 contract with the Packers on April 2 , 2002 . Pederson again was the backup quarterback and primary holder in all 16 games in 2002 . In a week 7 game against the Washington Redskins , Favre suffered a sprained knee and Pederson took most of the snaps in the second half , going 9 - for - 15 for 78 yards to help win the game 30 -- 9 . Pederson also played in games against the Miami Dolphins , Detroit Lions , and New York Jets . He re-signed with the Packers to a one - year , $750,000 contract on April 29 , 2003 . For the third consecutive season , Pederson backed up Favre in all 16 games and held placekicks . He completed both of his passes during the regular season for a total of 16 yards . The Packers re-signed Pederson to a one - year contract on April 28 , 2004 . Tim Couch was signed to compete for the backup quarterback job , but lost out to Pederson and was released on September 5 , 2004 . In a week 3 game against the Indianapolis Colts , Pederson replaced Favre in a blowout loss and went 4 - of - 6 for 34 yards and an interception . The next week , a week 4 game against the New York Giants , Favre sustained a concussion in the third quarter , and Pederson replaced him at quarterback . Pederson went 7 - of - 17 for 86 yards and an interception in the loss before he suffered a hit to his side in the third quarter that resulted in a cracked bone in his back , a torn muscle in his side , and a broken rib . He stayed in the game up until the last snap , when he was replaced by third - string quarterback Craig Nall . Pederson was placed on injured reserve on October 7 , ending his season . He retired in March 2005 to become a head coach at Calvary Baptist Academy . Coaching career ( edit ) High school ( edit ) After his retirement , Pederson was hired as head football coach of Calvary Baptist Academy , a private Christian high school in Shreveport , Louisiana . Calvary was going into its second year as a program when Pederson signed on in March 2005 . Pederson was the head coach at Calvary for four years , and held a 33 -- 7 record in the regular season and an 8 -- 3 record in the post-season . The Cavaliers were in the state playoffs all four years with Pederson as head coach . In his first season in 2005 , the Cavaliers went 5 -- 6 and lost in the first round of the state playoffs . In 2007 , he led the Cavaliers to the semi-finals and to their first district title . NFL assistant coaching positions ( edit ) On January 29 , 2009 , Pederson was hired as the offensive quality control coach for the Philadelphia Eagles , reuniting him with his former head coach , Andy Reid . He was promoted to quarterbacks coach on February 8 , 2011 , replacing James Urban , who was promoted to assistant offensive coordinator . On January 11 , 2013 , Pederson followed Andy Reid to the Kansas City Chiefs to serve as offensive coordinator . NFL head coach ( edit ) On January 18 , 2016 , Pederson was hired as head coach of the Eagles replacing Chip Kelly . Despite having Sam Bradford on the roster as the starting quarterback , the Eagles drafted Carson Wentz with the second overall pick in 2016 . Right before the 2016 season began , Bradford was traded to the Minnesota Vikings and Wentz was named the starting quarterback as a rookie . Pederson and Wentz won their first three NFL games together , but finished the season 7 -- 9 , missing the playoffs . Pederson 's second season was much more successful as he led the Eagles to a 13 - 3 record , winning them the NFC East division championship and allotting them home - field advantage throughout the playoffs . Wentz , who was having a career year and was considered a front runner for league MVP , tore his ACL in Week 14 , leaving backup Nick Foles with the starting job for the remainder of the year . Despite becoming major playoff underdogs due to the loss of Wentz , Foles filled in admirably as the starter , allowing Philadelphia to make it to Super Bowl LII , their first Super Bowl appearance since the 2004 season . Eventual Super Bowl MVP Foles led the team in a 41 - 33 win over the New England Patriots , giving them their first Lombardi Trophy in franchise history and their first league championship since 1960 . Head coaching record ( edit ) Team Year Regular season Postseason Won Lost Ties Win % Finish Won Lost Win % Result PHI 2016 7 9 0 . 438 4th in NFC East -- -- -- -- PHI 2017 13 0 . 813 1st in NFC East 0 1.000 Won Super Bowl LII Total 20 12 0 . 625 0 1.000 Coaching tree ( edit ) NFL head coaches under whom Pederson has served : Andy Reid , Philadelphia Eagles ( 2009 -- 2012 ) Kansas City Chiefs ( 2013 - 2015 ) Assistant coaches under Pederson who have become NFL head coaches : Frank Reich , Indianapolis Colts ( 2018 -- present ) Personal life ( edit ) Pederson was born to Teri ( née Boykin ) and Gordon `` Gordy '' Pederson ( 1939 -- 2016 ) on January 31 , 1968 , in Bellingham , Washington . A devout Christian , Pederson and his wife Jeannie have three sons . Pederson has been a resident of Moorestown , New Jersey . References ( edit ) Jump up ^ Cahill , Dan ( January 25 , 2016 ) . `` Ron Rivera could become one of four to win Super Bowl as player and coach '' . chicago.suntimes.com . Retrieved February 19 , 2018 . Jump up ^ McCallister , Laura ( January 11 , 2013 ) . `` Andy Reid fills out coaching staff , including coordinators '' . KCTV - TV website . Retrieved January 12 , 2013 . Jump up ^ `` ULM Football Records . '' 2008 ULM Warhawks Football Media Guide . Retrieved on September 7 , 2008 . Jump up ^ `` Transactions '' . The Dispatch. 1991 - 05 - 01 . Retrieved 2010 - 02 - 28 . Jump up ^ `` Doug Pederson '' . Pro Football Reference . Retrieved 2010 - 02 - 28 . Jump up ^ Mayo , Michael ( August 17 , 1991 ) . `` Brown On Leave But Robbie Is , Too '' . Sun - Sentinel . Retrieved 2017 - 08 - 06 . Jump up ^ Smith , Timothy W. ( February 5 , 1992 ) . `` Football ; World League Kicks Off Its Talent Hunt for 1992 '' . The New York Times . Retrieved 2017 - 08 - 06 . Jump up ^ Smith , Timothy W. ( May 17 , 1992 ) . `` Football ; Knights Do Their Part but Ca n't Gain Playoff Berth '' . The New York Times . Retrieved 2017 - 08 - 06 . Jump up ^ `` Transactions '' . The Dispatch . June 2 , 1992 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Transactions '' . The New York Times . October 8 , 1992 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Sports Today '' . The Buffalo News . March 3 , 1993 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Transactions '' . The New York Times . August 31 , 1993 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Transactions '' . The New York Times . September 1 , 1993 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Doug Pederson '' . NFL.com . Retrieved 2017 - 08 - 06 . Jump up ^ George , Thomas ( November 15 , 1993 ) . `` Pro Football ; No. 325 : Shula Rides the Shoulders '' . The New York Times . Retrieved 2017 - 08 - 06 . Jump up ^ `` Doug Pederson : Game Logs at NFL.com '' . www.nfl.com . Retrieved 21 February 2018 . Jump up ^ `` FINS FLASHBACK : Coach Shula Wins 325 '' . Retrieved 21 February 2018 . Jump up ^ Freeman , Mike ( December 6 , 1993 ) . `` Pro Football ; Finally , Giants Get Some Respect '' . The New York Times . Retrieved 2017 - 08 - 06 . Jump up ^ Cole , Jason ( December 16 , 1993 ) . `` Pederson , Humphrey Are Dolphins No More '' . Sun - Sentinel . Retrieved 2017 - 08 - 06 . Jump up ^ `` Transactions '' . The New York Times . April 16 , 1994 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Expansion Draft '' . panthers.com . Retrieved 2017 - 08 - 06 . Jump up ^ Cole , Jason ( January 19 , 1995 ) . `` No Big Names On Dolphins ' Expansion List '' . Sun - Sentinel . Retrieved 2017 - 08 - 06 . Jump up ^ `` Capers makes first big roster cuts '' . Star - News . May 24 , 1995 . Retrieved 2010 - 02 - 28 . Jump up ^ Calkins , Geoff ( June 7 , 1995 ) . `` Kicker , Safety Contracts Extended '' . Sun - Sentinel . access - date = requires url = ( help ) Jump up ^ Calkins , Geoff ( August 22 , 1995 ) . `` It 's McGwire : Pederson Loses No. 3 QB Spot '' . Sun - Sentinel . Retrieved 2017 - 08 - 06 . Jump up ^ Cole , Jason ( October 10 , 1995 ) . `` Marino To Miss A Game Or 2 '' . Sun - Sentinel . Retrieved 2017 - 08 - 06 . Jump up ^ `` Checkpoint Was Sore Spot to Chief Fans '' . The Los Angeles Times . October 11 , 1995 . Retrieved 2017 - 08 - 06 . ^ Jump up to : `` Transactions '' . The New York Times . February 23 , 2001 . Retrieved 2017 - 08 - 06 . Jump up ^ Plaschke , Bill ( November 11 , 1995 ) . `` Any Old Joe Could Start For Packers This Week '' . The Los Angeles Times . Retrieved 2017 - 08 - 06 . Jump up ^ `` Hostetler Is Doubtful ; Packers Change Backups '' . The Los Angeles Times . November 23 , 1995 . Retrieved 2017 - 08 - 06 . Jump up ^ `` McMahon Joins the Packers '' . The New York Times . November 29 , 1995 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Rubley , Former Ram , Waived by Packers '' . The Los Angeles Times . December 13 , 1995 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Thursday 's Transactions '' . Laurel Leader . February 21 , 1997 . Retrieved 2017 - 08 - 06 . ( subscription required ) Jump up ^ Pierson , Don ( October 6 , 1998 ) . `` Vikings 37 , Packers 24 '' . The Chicago Tribune . Retrieved 2017 - 08 - 06 . Jump up ^ `` Packers trade quarterback Rick Mirer to Jets for draft choice '' . packers.com . August 20 , 1999 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Eagles Sign Backup Passer to Start '' . The New York Times . February 18 , 1999 . Retrieved 2017 - 08 - 06 . Jump up ^ `` McNabb Signed but Not in Uniform '' . The New York Times . August 1 , 1999 . Retrieved 2017 - 08 - 06 . Jump up ^ Kinney , David ( September 12 , 1999 ) . `` Jacke 's 4th Field Goal Gives Cardinals 25 -- 24 Win '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ Berger , Ken ( September 19 , 1999 ) . `` Bucaneers Pound Philadelphia in McNabb 's Debut '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ Porter , Bill ( September 26 , 1999 ) . `` Bills Bury Punchless Eagles , 26 -- 0 '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ Cananvan , Tom ( October 3 , 1999 ) . `` Giants Edge Eagles , 16 -- 15 '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ Moore , David and Jean - Jacques Taylor ( October 11 , 1999 ) . `` Cowboys ' Irvin Suffers Spinal Injury in Loss '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ `` Reeves Releases A Former Top Pick '' . The New York Times . August 28 , 2000 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Reed Suddenly Leaves Denver , And Hopes to Help Another Team '' . The New York Times . September 2 , 2000 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Big Fine Surprises Steeler '' . The New York Times . December 9 , 2000 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Quarterback Doug Pederson officially rejoins Packers , reports Monday ; Horacek also added '' . packers.com . March 13 , 2001 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Packers Re-Sign QB Doug Pederson '' . packers.com. April 29 , 2003 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Giants release Collins , focus search for backup on O'Donnell '' . Buffalo News . April 29 , 2004 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Packers cut former No. 1 pick Tim Couch '' . ESPN.com . September 5 , 2004 . Retrieved 2017 - 08 - 06 . Jump up ^ Stapleton , Arnie ( October 4 , 2004 ) . `` Packers Suffer A Giant Loss '' . The Washington Post . Retrieved 2017 - 08 - 06 . Jump up ^ `` Packers Re-sign Smith , Place Pederson On I.R , Sign Millhouse To Practice Squad '' . packers.com . October 7 , 2004 . Retrieved 2017 - 08 - 06 . ^ Jump up to : Fedotin , Jeff ( 2006 - 08 - 25 ) . `` Using Lessons He Learned Backing Up Favre , Pederson Coaches High Schoolers '' . Green Bay Packers . Archived from the original on August 27 , 2009 . Retrieved 2010 - 02 - 28 . Jump up ^ `` The History of Calvary Baptist Academy '' . cavsfootball.com. 2007 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Eagles bring back ex-QB Pederson '' . ESPN . Associated Press . 2009 - 01 - 30 . Retrieved 2010 - 02 - 28 . Jump up ^ `` 2011 Coaching Staff Complete '' . Philadelphiaeagles.com. 2011 - 02 - 08 . Retrieved 2011 - 02 - 08 . Jump up ^ `` Chiefs hire Doug Pederson and Bob Sutton as Coordinators '' . nfl.com. 2013 - 01 - 11 . Retrieved 2013 - 01 - 11 . Jump up ^ Sheridan , Phil ( January 18 , 2016 ) . `` Doug Pederson announced as Eagles coach '' . ESPN.com . Retrieved 2017 - 08 - 06 . Jump up ^ Wells , Mike ( 11 February 2018 ) . `` Eagles OC Frank Reich named new Colts coach '' . ESPN . Retrieved 11 February 2018 . Jump up ^ `` Gordon `` Gordy '' Pederson `` . Legacy.com . September 29 , 2016 . Retrieved 2017 - 08 - 06 . Jump up ^ `` Eagles head coach Doug Pederson gives glory to God , says his faith keeps him grounded '' . Jump up ^ McLane , Jeff ( September 11 , 2016 ) . `` Pederson goes on without his first mentor -- his late father '' . The Philadelphia Inquirer . Retrieved 2017 - 08 - 06 . Jump up ^ Cuellar , Dann . `` Doug Pedersons ' neighbors ready to welcome him back home '' , WPVI - TV , January 19 , 2016 . Accessed November 29 , 2017 . `` New Philadelphia Eagles head coach Doug Pederson and his family are excited about moving back to the area ... Before moving to Kansas City in 2011 , the Pederson family lived on a cul - de-sac in the unit block of Hamilton Court in Moorestown , New Jersey . '' External links ( edit ) Philadelphia Eagles profile Current head coaches of the National Football League American Football Conference AFC East AFC North AFC South AFC West Sean McDermott ( Buffalo Bills ) Adam Gase ( Miami Dolphins ) Bill Belichick ( New England Patriots ) Todd Bowles ( New York Jets ) John Harbaugh ( Baltimore Ravens ) Marvin Lewis ( Cincinnati Bengals ) Hue Jackson ( Cleveland Browns ) Mike Tomlin ( Pittsburgh Steelers ) Bill O'Brien ( Houston Texans ) Frank Reich ( Indianapolis Colts ) Doug Marrone ( Jacksonville Jaguars ) Mike Vrabel ( Tennessee Titans ) Vance Joseph ( Denver Broncos ) Andy Reid ( Kansas City Chiefs ) Anthony Lynn ( Los Angeles Chargers ) Jon Gruden ( Oakland Raiders ) National Football Conference NFC East NFC North NFC South NFC West Jason Garrett ( Dallas Cowboys ) Pat Shurmur ( New York Giants ) Doug Pederson ( Philadelphia Eagles ) Jay Gruden ( Washington Redskins ) Matt Nagy ( Chicago Bears ) Matt Patricia ( Detroit Lions ) Mike McCarthy ( Green Bay Packers ) Mike Zimmer ( Minnesota Vikings ) Dan Quinn ( Atlanta Falcons ) Ron Rivera ( Carolina Panthers ) Sean Payton ( New Orleans Saints ) Dirk Koetter ( Tampa Bay Buccaneers ) Steve Wilks ( Arizona Cardinals ) Sean McVay ( Los Angeles Rams ) Kyle Shanahan ( San Francisco 49ers ) Pete Carroll ( Seattle Seahawks ) Links to related articles Louisiana -- Monroe Warhawks starting quarterbacks Lloyd Ray ( 1951 ) Mark Boatright ( 1953 ) Pete Chambless ( 1954 ) Everett Reynolds ( 1956 -- 1958 ) Nathan Zametto ( 1957 ) Bob Randel ( 1959 ) Don Wattigny ( 1960 ) Ronnie Myrick ( 1961 -- 1962 ) Bookie Shillcutt ( 1963 ) Johnny Garrison ( 1964 ) Steve Mansur ( 1965 -- 1968 ) Terry Reisig ( 1967 ) Mike Audirsch ( 1969 ) Allen Taylor ( 1970 ) Don Smith ( 1971 -- 1972 ) Scotty Dyer ( 1973 -- 1974 ) Joe Bruner ( 1975 ) Mike Howell ( 1976 ) Bud Cespiva ( 1977 ) Kirby Arceneaux ( 1978 ) John Holman ( 1979 -- 1982 ) Rodney Horn ( 1983 -- 1984 ) Bubby Brister ( 1984 -- 1985 ) Stan Humphries ( 1986 -- 1987 ) Doug Pederson ( 1987 -- 1990 ) Wendal Lowrey ( 1991 -- 1992 ) Robert Cobb ( 1993 ) Raymond Philyaw ( 1993 -- 1996 ) Daniel Jeremiah ( 1997 ) Andre Vige ( 1998 ) Andy Chance ( 1999 -- 2001 ) Steven Jyles ( 2002 -- 2005 ) Kinsmon Lancaster ( 2006 -- 2008 ) Trey Revell ( 2007 -- 2009 ) Cody Wells ( 2009 , 2012 ) Kolton Browning ( 2010 -- 2013 ) Brayle Brown ( 2013 ) Pete Thomas ( 2014 ) Garrett Smith ( 2015 -- 2017 ) Earnest Carrington ( 2015 ) Will Collins ( 2016 ) Caleb Evans ( 2016 -- 2017 ) 1987 Northeast Louisiana Indians football -- NCAA Division I - AA national champions Teddy Garcia Jackie Harris Stan Humphries Doug Pederson Head coach : Pat Collins Assistant coaches : Rickey Bustle Chris Scelfo Green Bay Packers Super Bowl XXXI champions 4 Brett Favre 7 Kyle Wachholtz 9 Jim McMahon 13 Chris Jacke 17 Craig Hentrich 18 Doug Pederson 21 Craig Newsome 25 Dorsey Levens 27 Calvin Jones 28 Roderick Mullen 30 William Henderson 32 Travis Jervey 33 Doug Evans 34 Edgar Bennett 36 LeRoy Butler 37 Tyrone Williams 39 Mike Prior 40 Chris Hayes 41 Eugene Robinson 46 Michael Robinson 51 Brian Williams 52 Frank Winters 53 George Koonce 54 Ron Cox 55 Bernardo Harris 56 Lamont Hollinquest 59 Wayne Simmons 62 Marco Rivera 63 Adam Timmerman 64 Bruce Wilkerson 65 Lindsay Knapp 67 Jeff Dellenbach 68 Gary Brown 71 Santana Dotson 72 Earl Dotson 73 Aaron Taylor 75 Ken Ruettgers 77 John Michels 80 Derrick Mayes 81 Desmond Howard ( MVP ) 82 Don Beebe 83 Jeff Thomason 84 Andre Rison 85 Terry Mickens 86 Antonio Freeman 87 Robert Brooks 88 Keith Jackson 89 Mark Chmura 90 Darius Holland 91 Shannon Clavelle 92 Reggie White 93 Gilbert Brown 94 Bob Kuberski 95 Keith McKenzie 96 Sean Jones 98 Gabe Wilkins Head coach : Mike Holmgren Coaches : Larry Brooks Nolan Cromwell Gil Haskell Johnny Holland Sherman Lewis Jim Lind Tom Lovat Marty Mornhinweg Andy Reid Gary Reynolds Fritz Shurmur Harry Sydney Bob Valesente Philadelphia Eagles Super Bowl LII champions 4 Jake Elliott 6 Caleb Sturgis 7 Nate Sudfeld 8 Donnie Jones 9 Nick Foles ( MVP ) 10 Mack Hollins 11 Carson Wentz 13 Nelson Agholor 14 Marcus Johnson 16 Bryce Treggs 17 Alshon Jeffery 18 Shelton Gibson 19 Rashard Davis 21 Patrick Robinson 22 Sidney Jones 23 Rodney McLeod 24 Corey Graham 26 Jaylen Watkins 27 Malcolm Jenkins 28 Wendell Smallwood 29 LeGarrette Blount 30 Corey Clement 31 Jalen Mills 32 Rasul Douglas 33 De'Vante Bausby 34 Donnel Pumphrey 35 D.J. Killings 36 Jay Ajayi 38 Kenjon Barner 39 Harold Jones - Quartey 41 Ronald Darby 42 Chris Maragos 43 Darren Sproles 45 Rick Lovato 47 Nathan Gerry 49 Tre Sullivan 50 Bryan Braman 51 Steven Means 52 Najee Goode 53 Nigel Bradham 54 Kamu Grugier - Hill 55 Brandon Graham 56 Chris Long 57 Dannell Ellerbe 58 Jordan Hicks 59 Joe Walker 61 Stefen Wisniewski 62 Jason Kelce 65 Lane Johnson 66 Will Beatty 67 Chance Warmack 68 Josh Andrews 71 Jason Peters 72 Halapoulivaati Vaitai 73 Isaac Seumalo 75 Vinny Curry 78 Darrell Greene 79 Brandon Brooks 82 Torrey Smith 85 Billy Brown 86 Zach Ertz 87 Brent Celek 88 Trey Burton 89 Greg Ward Jr . 91 Fletcher Cox 93 Timmy Jernigan 94 Beau Allen 95 Mychal Kendricks 96 Derek Barnett 97 Destiny Vaeao 98 Elijah Qualls Randall Goforth Aziz Shittu Dom Williams Head coach : Doug Pederson Coaches : Eugene Chung Phillip Daniels John DeFilippo Dave Fipp Ken Flajole Keith Gray Mike Groh Matthew Harper Tim Hauck Josh Hingst Shaun Huls Trent Miles Ryan Paganetti T.J. Paganetti Justin Peelle Spencer Phillips Frank Reich Jim Schwartz Duce Staley Jeff Stoutland Press Taylor Cory Undlin Dino Vasso Chris Wilson Philadelphia Eagles starting quarterbacks Red Kirkman ( 1933 -- 1935 ) Jim Leonard ( 1934 , 1936 ) Stumpy Thomason ( 1935 ) Irv Kupcinet ( 1935 ) Glenn Frey ( 1936 -- 1937 ) Jess Dow ( 1938 -- 1939 ) Davey O'Brien ( 1939 -- 1940 ) Len Barnum ( 1941 ) Tommy Thompson ( 1941 -- 1942 , 1946 -- 1950 ) Foster Watkins ( 1941 ) Roy Zimmerman ( 1943 -- 1946 ) Allie Sherman ( 1945 ) Bill Mackrides ( 1948 ) Jack Myers ( 1948 ) Adrian Burk ( 1951 -- 1956 ) Bobby Thomason ( 1952 -- 1957 ) Sonny Jurgensen ( 1957 , 1961 -- 1963 ) Norm Van Brocklin ( 1958 -- 1960 ) King Hill ( 1962 -- 1966 , 1968 ) Norm Snead ( 1964 -- 1970 ) Jack Concannon ( 1964 , 1966 ) John Huarte ( 1968 ) George Mira ( 1969 ) Rick Arrington ( 1970 -- 1971 ) Pete Liske ( 1971 -- 1972 ) John Reaves ( 1972 ) Roman Gabriel ( 1973 -- 1976 ) Mike Boryla ( 1974 -- 1976 ) Ron Jaworski ( 1977 -- 1986 ) Joe Pisarcik ( 1984 ) Randall Cunningham ( 1985 -- 1995 ) Matt Cavanaugh ( 1986 ) Scott Tinsley ( 1987 ) Guido Merkens ( 1987 ) Jim McMahon ( 1991 -- 1992 ) Jeff Kemp ( 1991 ) Brad Goebel ( 1991 ) Bubby Brister ( 1993 -- 1994 ) Ken O'Brien ( 1993 ) Rodney Peete ( 1995 -- 1998 ) Ty Detmer ( 1996 -- 1997 ) Bobby Hoying ( 1997 -- 1998 ) Koy Detmer ( 1998 -- 1999 , 2002 , 2004 ) Doug Pederson ( 1999 ) Donovan McNabb ( 1999 -- 2009 ) A.J. Feeley ( 2002 , 2007 ) Mike McMahon ( 2005 ) Jeff Garcia ( 2006 ) Kevin Kolb ( 2009 -- 2010 ) Michael Vick ( 2010 -- 2013 ) Vince Young ( 2011 ) Nick Foles ( 2012 -- 2014 , 2017 -- present ) Mark Sanchez ( 2014 -- 2015 ) Sam Bradford ( 2015 ) Carson Wentz ( 2016 -- 2017 ) Cleveland Browns starting quarterbacks Cliff Lewis ( 1946 -- 1947 , 1949 ) Otto Graham ( 1946 -- 1955 ) George Ratterman ( 1953 -- 1956 ) Babe Parilli ( 1956 ) Tommy O'Connell ( 1956 -- 1957 ) Milt Plum ( 1957 -- 1961 ) Len Dawson ( 1961 ) Jim Ninowski ( 1962 -- 1963 , 1965 ) Frank Ryan ( 1962 -- 1968 ) Gary Lane ( 1967 ) Bill Nelsen ( 1968 -- 1972 ) Mike Phipps ( 1970 -- 1976 ) Don Gault ( 1970 ) Brian Sipe ( 1974 -- 1983 ) Will Cureton ( 1975 ) Dave Mays ( 1977 ) Terry Luck ( 1977 ) Paul McDonald ( 1982 -- 1984 ) Gary Danielson ( 1985 -- 1988 ) Bernie Kosar ( 1985 -- 1993 ) Jeff Christensen ( 1987 ) Mike Pagel ( 1988 , 1990 ) Don Strock ( 1988 ) Mike Tomczak ( 1992 ) Todd Philcox ( 1992 -- 1993 ) Vinny Testaverde ( 1993 -- 1995 ) Mark Rypien ( 1994 ) Eric Zeier ( 1995 ) Ty Detmer ( 1999 ) Tim Couch ( 1999 -- 2003 ) Doug Pederson ( 2000 ) Spergon Wynn ( 2000 ) Kelly Holcomb ( 2002 -- 2004 ) Jeff Garcia ( 2004 ) Luke McCown ( 2004 ) Trent Dilfer ( 2005 ) Charlie Frye ( 2005 -- 2007 ) Derek Anderson ( 2006 -- 2009 ) Brady Quinn ( 2008 -- 2009 ) Ken Dorsey ( 2008 ) Bruce Gradkowski ( 2008 ) Jake Delhomme ( 2010 ) Seneca Wallace ( 2010 -- 2011 ) Colt McCoy ( 2010 -- 2011 ) Brandon Weeden ( 2012 -- 2013 ) Thad Lewis ( 2012 ) Brian Hoyer ( 2013 -- 2014 ) Jason Campbell ( 2013 ) Johnny Manziel ( 2014 -- 2015 ) Connor Shaw ( 2014 ) Josh McCown ( 2015 -- 2016 ) Austin Davis ( 2015 ) Robert Griffin III ( 2016 ) Cody Kessler ( 2016 ) DeShone Kizer ( 2017 ) Kevin Hogan ( 2017 ) Philadelphia Eagles head coaches Lud Wray ( 1933 -- 1935 ) Bert Bell ( 1936 -- 1940 ) Greasy Neale ( 1941 -- 1950 ) Bo McMillin ( 1951 ) Wayne Millner # ( 1951 ) Jim Trimble ( 1952 -- 1955 ) Hugh Devore ( 1956 -- 1957 ) Buck Shaw ( 1958 -- 1960 ) Nick Skorich ( 1961 -- 1963 ) Joe Kuharich ( 1964 -- 1968 ) Jerry Williams ( 1969 -- 1971 ) Ed Khayat ( 1971 -- 1972 ) Mike McCormack ( 1973 -- 1975 ) Dick Vermeil ( 1976 -- 1982 ) Marion Campbell ( 1983 -- 1985 ) Fred Bruney # ( 1985 ) Buddy Ryan ( 1986 -- 1990 ) Rich Kotite ( 1991 -- 1994 ) Ray Rhodes ( 1995 -- 1998 ) Andy Reid ( 1999 -- 2012 ) Chip Kelly ( 2013 -- 2015 ) Pat Shurmur # ( 2015 ) Doug Pederson ( 2016 -- ) Pound ( # ) sign denotes interim head coach . Retrieved from `` https://en.wikipedia.org/w/index.php?title=Doug_Pederson&oldid=832875504 '' Categories : 1968 births Living people American football quarterbacks Cleveland Browns players Green Bay Packers players High school football coaches in the United States Kansas City Chiefs coaches Louisiana -- Monroe Warhawks football players Miami Dolphins players New York / New Jersey Knights players People from Moorestown , New Jersey Philadelphia Eagles head coaches Philadelphia Eagles players Players of American football from Washington ( state ) Rhein Fire players Sportspeople from Bellingham , Washington Super Bowl champions Super Bowl - winning head coaches Hidden categories : Pages using citations with accessdate and no URL Pages containing links to subscription - only content Philadelphia Eagles currentteam parameter articles NFL player with coaching information Infobox NFL biography articles missing alt text Talk Contents About Wikipedia Italiano Português Suomi Edit links This page was last edited on 28 March 2018 , at 13 : 22 . Text is available under the Creative Commons Attribution - ShareAlike License ; additional terms may apply . By using this site , you agree to the Terms of Use and Privacy Policy . Wikipedia ® is a registered trademark of the Wikimedia Foundation , Inc. , a non-profit organization . About Wikipedia

Question: what position did doug peterson play in the nfl

a member of the Green Bay Packers, serving as a backup quarterback to Brett Favre and holder on placekicks


National legal aid & Defender Association - wikipedia National legal aid & Defender Association Jump to : navigation , search The National Legal Aid & Defender Association ( NLADA ) is the oldest and largest national , nonprofit membership organization devoted to advocating equal justice for all Americans and was established in 1911 . Contents ( hide ) 1 History 2 See also 3 References 4 External links History ( edit ) The Fourteenth Amendment to the United States Constitution and its Equal Protection Clause provides Equal justice under law . Beginning in the late 1800s and throughout the early years of the 20th century , the American legal profession expressed its commitment to the concept of free legal assistance for poor people in the form of legal aid societies and bar association legal aid committees . The first legal aid society , The German Society of New York , was founded in 1876 to protect German immigrants from exploitation . Subsequently , the agency 's protection was extended to others and in 1890 it became the Legal Aid Society of New York . In 1888 , the Ethical Culture Society of Chicago established by the Bureau of Justice was the first agency to offer legal assistance to individuals regardless of nationality , race or gender . Other municipalities followed suit , and in the first decades of the 20th century most major cities had opened legal aid societies . In 1911 , legal aid societies joined together to form the National Alliance of Legal Aid Societies . Arthur von Briesen of the Legal Aid Society of New York was the first president of the organization that became the National Legal Aid & Defender Association ( NLADA ) in 1949 . The concept of free legal assistance for the poor was promoted by the publication of Reginald Heber Smith 's Justice and the Poor in 1919 . Smith challenged the legal profession to consider it an obligation to see that access to justice was available to all , without regard to ability to pay . `` Without equal access to the law , '' he wrote , `` the system not only robs the poor of their only protection , but places in the hands of their oppressors the most powerful and ruthless weapon ever invented . As a result of Smith 's book , the American Bar Association created the Special Committee on Legal Aid Work . By the middle of the 20th century , virtually every major metropolitan area had some kind of legal aid program . However , the system established was not suffice in meeting the needs of the poor and in the early 1960s a new model for legal services programs emerged . This new model was based on the philosophy that legal services should be a component of an overall anti-poverty effort . The Ford Foundation was one of the original supporters of this model . Subsequently , in 1964 came the passage of the Economic Opportunity Act , which launched the war on poverty and mandated federal funds to be allocated for the first time to fund legal services to the poor . However , this law did not specifically provide for legal services and it took many years and attempts to finally devise a federal construct to support legal aid for the low - income community . After years of research and advocacy , the Legal Services Corporation Act was enacted in 1974 by President Richard Nixon . This program flourished and has evolved over the years to become the organization nationally recognized as the Legal Services Corporation . Over the years legal aid has evolved into a comprehensive program that provides legal assistance to low - income people regardless of race , ethnicity or gender . NLADA is the only national membership organization devoted exclusively to ensuring legal representation for individuals who can not afford an attorney . NLADA serves the equal justice community in two major ways : providing products and services and as a national voice in public policy and legislative debates on the many issues affecting the equal justice community . NLADA also serves as a resource for those seeking more information on equal justice in the United States . In recent years , the NLADA has honored individuals and organizations with a variety of awards at its annual conferences . Awards range from the Mary Ellen Hamilton award , which recognizes a client of a legal aid or defender organization , to the Clara Shortridge Foltz Award , which honors organizations that have pioneered important innovations in the field . Notable recipients of NLADA awards include former Attorney General Janet Reno , Bronx Defenders attorney David Feige , and Hillary Clinton . Paul D. Borman was Chief Federal Defender , Legal Aid & Defender Association of Detroit , from 1979 to 1994 . He was the first federal defender to be appointed to the federal bench , in 1994 . See also ( edit ) Human rights portal Law portal Legal aid in the United States References ( edit ) Jump up ^ `` Legal Aid Societies , '' MSN Encantra , ( 1 ) . Archived November 1 , 2009 . Jump up ^ `` The History of the Legal Aid Society , '' The Legal Aid Society , http://www.legal-aid.org/en/aboutus/ourhistory.aspx Jump up ^ `` Ethical Culture and the Ongoing Quest for Social Justice , '' The Ethical Culture Society of Bergen County , NJ , ( 2 ) Jump up ^ `` Would Keep Law and Charity Apart ; Legal Aid Society Refuses to Amalgamate with the Charities Conference '' . The New York Times . November 16 , 1912 . Jump up ^ Justice and the Poor , Reginald Heber Smith , 1919 Jump up ^ `` NLADA : About NLADA -- Awards , '' National Legal Aid & Defender Association , http://www.nlada.org/About/About_Awards Jump up ^ `` Judge Paul D. Borman '' . United States District Court -- Eastern Michigan District . Jump up ^ ( 3 ) External links ( edit ) nlada.org , the organization 's official website Retrieved from `` https://en.wikipedia.org/w/index.php?title=National_Legal_Aid_%26_Defender_Association&oldid=826346364 '' Categories : 1911 establishments in the United States Legal aid Law - related professional associations Legal advocacy organizations in the United States Non-profit organizations based in Washington , D.C. Organizations established in 1911 Criminal justice reform in the United States Criminal defense organizations Hidden categories : Use mdy dates from June 2015 Talk Contents About Wikipedia Add links This page was last edited on 18 February 2018 , at 16 : 48 . About Wikipedia

Question: which organization is the major voice of the legal profession


Geoffrey Palmer ( actor ) - wikipedia Geoffrey Palmer ( actor ) Jump to : navigation , search Geoffrey Palmer OBE Palmer performing at A Breath of Fresh Air , June 2008 ( 1927 - 06 - 04 ) 4 June 1927 ( age 90 ) London , England Occupation Actor Years active 1958 -- present Spouse ( s ) Sally Green ( m . 1963 ) Children 2 , including Charles Palmer Geoffrey Dyson Palmer , OBE ( born 4 June 1927 ) is an English actor known for his roles in British television sitcoms playing Jimmy Anderson in The Fall and Rise of Reginald Perrin ( 1976 -- 79 ) , Ben Parkinson in Butterflies ( 1978 -- 83 ) and Lionel Hardcastle in As Time Goes By ( 1992 -- 2005 ) . His film appearances include A Fish Called Wanda ( 1988 ) , The Madness of King George ( 1994 ) , Mrs. Brown ( 1997 ) , and Tomorrow Never Dies ( 1997 ) . Contents ( hide ) 1 Career 2 Personal life 3 Awards and recognition 4 Appearances 4.1 Stage 4.2 Radio 4.3 Television 4.4 Film 4.5 Recordings ( spoken word ) 5 References 6 External links Career ( edit ) Palmer 's early television appearances included a variety of roles in Granada Television 's The Army Game , two episodes of The Baron and as a property agent in Cathy Come Home . Getting a major break in John Osborne 's West of Suez at the Royal Court with Ralph Richardson , he then acted in major productions at the Royal Court and for the National Theatre Company and was directed by Laurence Olivier in J.B. Priestley 's Eden End . Palmer found the play so boring , however , that it put him off a stage career for good . Two sitcom roles brought him attention in the 1970s : the hapless brother - in - law of Reggie Perrin in The Fall and Rise of Reginald Perrin ( 1976 -- 79 ) , and the phlegmatic Ben Parkinson in Carla Lane 's Butterflies ( 1978 -- 83 ) . He has continued to appear in productions written by Perrin creator David Nobbs , the latest being the radio comedy The Maltby Collection . He starred opposite Judi Dench for over a decade in the BBC situation comedy As Time Goes By ( 1992 -- 2005 ) . During this time he also appeared with Dench in other productions , including the James Bond film Tomorrow Never Dies , in which he portrayed Admiral Roebuck , and Mrs. Brown , playing Sir Henry Ponsonby to Dench 's Queen Victoria . He played Doctor Price in the Fawlty Towers episode `` The Kipper and the Corpse '' , determined to get breakfast amidst the confusion caused by the death of a guest and Basil 's inept way of handling the emergency . His distinctive voice has given him a career in advertising in such commercials as the ' Slam in the Lamb ' ads for the Meat & Livestock Commission ; television voiceovers such as the Audi commercials in which he popularised the phrase `` Vorsprung durch Technik '' , and as the narrator for the BBC series Grumpy Old Men and Grumpy Old Holidays . He narrated the audiobook version of Dickens ' A Christmas Carol , released in 2005 as a podcast by Penguin Books . He narrates Little England . In the 2006 DVD series The Compleat Angler , Palmer partners Rae Borras in a series of episodes based on Izaak Walton 's 1653 The Compleat Angler . In 2007 , he recorded The Diary of a Nobody by George Grossmith and Weedon Grossmith as an online audiobook . In December 2007 , Palmer appeared in the role of the Captain in `` Voyage of the Damned '' , the Christmas special episode of the BBC science - fiction series Doctor Who ; Palmer previously appeared in the classic era of the show as different characters in the Third Doctor serials Doctor Who and the Silurians and The Mutants . In March 2009 , he joined in a sketch with the two double acts `` Armstrong and Miller '' and `` Mitchell and Webb '' for Comic Relief . In 2011 , he played the reactionary father - in - law of the eponymous clergyman of Rev. in its Christmas episode . Personal life ( edit ) Palmer was born in London and attended Highgate School . He is the son of Norah Gwendolen ( née Robins ) and Frederick Charles Palmer , who was a chartered surveyor . He has a daughter , Harriet , and a son , Charles , a television director , who is married to actress Claire Skinner . Awards and recognition ( edit ) In the New Year 's Honours List published 31 December 2004 he was appointed an Officer of the Order of the British Empire ( OBE ) for services to drama . Appearances ( edit ) Stage ( edit ) Kafka 's Dick by Alan Bennett at the Royal Court Theatre ( 1986 ) Eden End by J.B. Priestley at the Royal National Theatre ( 1974 ) Radio ( edit ) At Home With The Snails ( 2001 -- 2002 ) Les Miserables as Inspector Javert ( 2002 ) The Man Who Was Thursday ( 2005 ) High Table , Lower Orders ( 2005 -- 2006 ) The Maltby Collection ( 2007 -- 2009 ) A Murder of Quality ( 2009 ) North by Northamptonshire ( 2011 - 2012 ) Two Pipe Problems : The Case of the Missing Meerschaum as Mortimer Tregennis ( 2011 ) Television ( edit ) The Army Game ( 1958 -- 1960 ) as Various Characters The Avengers : `` Propellant 23 '' ( 1962 ) as Paul Manning `` Man with Two Shadows '' ( 1963 ) as Dr. Terence `` A Surfeit of H2O '' ( 1965 ) as Martin Smythe The Saint : `` The Rough Diamonds '' ( 1963 ) as Pete Ferguson Gideon 's Way ( TV Series ) `` The Alibi Man '' ( 1965 ) as Jeff Grant The Baron : `` Masquerade '' ( 1966 ) as Anstruther `` The Killing '' ( 1966 ) as Anstuther The Wednesday Play : Cathy Come Home ( 1966 ) as Property Agent Mrs Thursday ( 1966 ) as Henry Baxter Doctor Who : Doctor Who and the Silurians ( 1970 ) as Masters The Mutants ( 1972 ) as Administrator `` Voyage of the Damned '' ( 2007 ) as Captain Hardaker Colditz ( 1972 ) as Doc The Fall and Rise of Reginald Perrin ( 1976 -- 1979 ) as Jimmy Anderson Butterflies ( 1978 -- 1983 ) as Ben Parkinson The Professionals ( 1978 ) as Sinclair in `` Where The Jungle Ends '' Fawlty Towers : `` The Kipper and the Corpse '' ( 1979 ) as Dr. Price The Goodies ( 1980 ) as School Headmaster Whoops Apocalypse ( 1982 ) as Foreign Secretary Death of an Expert Witness ( 1983 ) as Dr. Edwin Lorrimer The Professionals ( 1983 ) as Avery in `` The Ojuka Situation '' Fairly Secret Army ( 1984 - 1986 ) as Major Harry Kitchener Wellington Truscott Executive Stress ( 1986 first series only ) as Donald Fairchild # 1 Season 's Greetings ( 1986 ) as Bernard Hot Metal ( 1986 ) as Harold Stringer Christabel ( 1988 ) as Mr. Burton Blackadder Goes Forth : Episode 6 `` Goodbyeee '' ( 1989 ) ; as Field Marshal Douglas Haig Inspector Morse : `` The Infernal Serpent '' ( 1990 ) as Matthew Copley - Barnes Bergerac ( 1990 ) `` Roots of Evil '' as Nigel Carter As Time Goes By ( 1992 -- 2005 ) as Lionel Hardcastle The Legacy of Reginald Perrin ( 1996 ) as Jimmy Anderson Alice Through the Looking Glass ( 1998 ) as White King The Savages ( 2001 ) as Donald The 1940s House ( 2001 ) as Narrator Absolute Power ( 2003 ) as Lord Harcourt Grumpy Old Men ( 2003 -- 2004 , 2006 ) as Narrator He Knew He Was Right ( 2004 ) as Sir Marmaduke Rowley Grumpy Old Holidays ( 2006 ) as Narrator Ashes to Ashes : `` Episode 8 '' ( 2008 ) ; as Lord Scarman The Long Walk to Finchley ( 2008 ) ; as John Crowder Agatha Christie 's Poirot : `` The Clocks '' ( 2009 ) as Vice Admiral Hamling Grandpa in My Pocket `` Captain Dumbletwit 's Toughest Mission Yet ! '' ( 2010 ) as Grandad Gillbert Rev `` Christmas Special '' Series 2 , episode 7 ( 2011 ) as Martin Henry IV , Part II ( 2012 ) ; as Lord Chief Justice Royal Variety Performance ( 2014 ) ; as the announcer ( voice - only ) Film ( edit ) A Prize of Arms ( 1962 ) as Cpl . Myers Incident at Midnight ( 1963 ) as Dr. Tanfield Ring of Spies ( 1964 ) as Police Officer ( uncredited ) Cast a Giant Shadow ( 1966 ) as David ( uncredited ) O Lucky Man ! ( 1973 ) as Examinator Doctor / Basil Keyes The Battle of Billy 's Pond ( 1976 ) - First Policeman The Outsider ( 1979 ) as Colonel Wyndham The Honorary Consul ( 1983 ) as Belfrage : British Ambassador A Zed & Two Noughts ( 1985 ) as Fallast Clockwise ( 1986 ) as Headmaster A Fish Called Wanda ( 1988 ) as Judge Hawks ( 1988 ) as SAAB Salesman The Madness of King George ( 1994 ) as Warren Mrs. Brown ( 1997 ) as Henry Ponsonby Tomorrow Never Dies ( 1997 ) as Admiral Roebuck Stiff Upper Lips ( 1998 ) as His Butler 's Voice Anna and the King ( 1999 ) as Lord John Bradley Rat ( 2000 ) as The Doctor Peter Pan ( 2003 ) as Sir Edward Quiller Couch Picadilly Jim ( 2005 ) - Bayliss The Pink Panther 2 ( 2009 ) as Joubert W.E. ( 2011 ) as Stanley Baldwin Lost Christmas ( 2011 ) as Dr. Clarence Run for Your Wife ( 2012 ) as Man on Bus Bert and Dickie ( 2012 ) as Charles Burnell The Last Sparks of Sundown ( 2014 ) as Sir Buster Sparks ( voice ) Paddington ( 2014 ) as Head Geographer Recordings ( spoken word ) ( edit ) Welcome to the Pleasuredome ( Fruitless Mix ) ( 1985 ) References ( edit ) Jump up ^ `` The Spectator ( 11 June 2011 ) '' . Exacteditions.com. 2011 - 06 - 11 . Retrieved 2015 - 03 - 13 . Jump up ^ `` The Penguin Podcast : A Christmas Carol - Episode 1 '' . 15 December 2005 . Archived from the original on 17 December 2005 . Retrieved 13 March 2015 . Jump up ^ `` Kylie Boards Titanic ! '' . BBC . 11 July 2007 . Archived from the original on 25 November 2007 . Retrieved 11 July 2007 . Jump up ^ `` Geoffrey Palmer Biography ( 1927 - ) '' . filmreference.com . Retrieved 2012 - 01 - 14 . Jump up ^ Loose Women , 12 December 2011 Jump up ^ `` The London Gazette '' . 31 December 2004 : 12 . Retrieved 2015 - 03 - 13 . Jump up ^ `` BBC Radio 4 Extra - G.K. Chesterton - The Man Who Was Thursday '' . Bbc.co.uk. 2016 - 08 - 21 . Retrieved 2016 - 08 - 21 . Jump up ^ `` BBC Radio 4 - Afternoon Drama , Two Pipe Problems , The Case of the Missing Meerschaum '' . Bbc.co.uk. 2013 - 12 - 25 . Retrieved 2015 - 03 - 13 . External links ( edit ) Geoffrey Palmer on IMDb Selected performances in Theatre Archive University of Bristol VIAF : 7732233 ISNI : 0000 0000 7252 9609 GND : 173637132 SUDOC : 114267413 MusicBrainz : b2548661 - d5ba - 4f3c - a731 - 4aec89b0b5a9 Retrieved from `` https://en.wikipedia.org/w/index.php?title=Geoffrey_Palmer_(actor)&oldid=825516396 '' Categories : 1927 births Living people Audiobook narrators English male film actors English male stage actors English male television actors English male voice actors English male radio actors Officers of the Order of the British Empire People educated at Highgate School Male actors from London 20th - century English male actors 20th - century Royal Marines personnel British male comedy actors Hidden categories : Use dmy dates from April 2012 Articles with hCards Pages using div col with deprecated parameters Wikipedia articles with VIAF identifiers Wikipedia articles with ISNI identifiers Wikipedia articles with GND identifiers Wikipedia articles with MusicBrainz identifiers Talk Contents About Wikipedia تۆرکجه Deutsch Español فارسی עברית Simple English Suomi Svenska Edit links This page was last edited on 13 February 2018 , at 21 : 12 . Text is available under the Creative Commons Attribution - ShareAlike License ; additional terms may apply . By using this site , you agree to the Terms of Use and Privacy Policy . Wikipedia ® is a registered trademark of the Wikimedia Foundation , Inc. , a non-profit organization . About Wikipedia

Question: who played lionel in as time goes by


Cellular respiration - wikipedia Cellular respiration Jump to : navigation , search This article needs additional citations for verification . Please help improve this article by adding citations to reliable sources . Unsourced material may be challenged and removed . ( September 2014 ) ( Learn how and when to remove this template message ) Typical eukaryotic cell Cellular respiration is a set of metabolic reactions and processes that take place in the cells of organisms to convert biochemical energy from nutrients into adenosine triphosphate ( ATP ) , and then release waste products . The reactions involved in respiration are catabolic reactions , which break large molecules into smaller ones , releasing energy in the process , as weak so - called `` high - energy '' bonds are replaced by stronger bonds in the products . Respiration is one of the key ways a cell releases chemical energy to fuel cellular activity . Cellular respiration is considered an exothermic redox reaction which releases heat . The overall reaction occurs in a series of biochemical steps , most of which are redox reactions themselves . Although cellular respiration is technically a combustion reaction , it clearly does not resemble one when it occurs in a living cell because of the slow release of energy from the series of reactions . Nutrients that are commonly used by animal and plant cells in respiration include sugar , amino acids and fatty acids , and the most common oxidizing agent ( electron acceptor ) is molecular oxygen ( O ) . The chemical energy stored in ATP ( its third phosphate group is weakly bonded to the rest of the molecule and is cheaply broken allowing stronger bonds to form , thereby transferring energy for use by the cell ) can then be used to drive processes requiring energy , including biosynthesis , locomotion or transportation of molecules across cell membranes . Contents ( hide ) 1 Aerobic respiration 1.1 Aerobic respiration summed up 1.2 Glycolysis 1.3 Oxidative decarboxylation of pyruvate 1.4 Citric acid cycle 1.5 Oxidative phosphorylation 2 Efficiency of ATP production 3 Fermentation 4 Anaerobic respiration 5 See also 6 References 7 External links Aerobic respiration Aerobic respiration ( red arrows ) is the main means by which both fungi and animals utilize chemical energy in the form of organic compounds that were previously created through photosynthesis ( green arrow ) . Aerobic respiration requires oxygen ( O ) in order to create ATP . Although carbohydrates , fats , and proteins are consumed as reactants , it is the preferred method of pyruvate breakdown in glycolysis and requires that pyruvate enter the mitochondria in order to be fully oxidized by the Krebs cycle . The products of this process are carbon dioxide and water , but the energy transferred is used to break bonds in ADP as the third phosphate group is added to form ATP ( adenosine triphosphate ) , by substrate - level phosphorylation , NADH and FADH Simplified reaction : C H O ( s ) + 6 O ( g ) → 6 CO ( g ) + 6 H O ( l ) + heat ΔG = − 2880 kJ per mol of C H O The negative ΔG indicates that the reaction can occur spontaneously . The potential of NADH and FADH is converted to more ATP through an electron transport chain with oxygen as the `` terminal electron acceptor '' . Most of the ATP produced by aerobic cellular respiration is made by oxidative phosphorylation . This works by the energy released in the consumption of pyruvate being used to create a chemiosmotic potential by pumping protons across a membrane . This potential is then used to drive ATP synthase and produce ATP from ADP and a phosphate group . Biology textbooks often state that 38 ATP molecules can be made per oxidised glucose molecule during cellular respiration ( 2 from glycolysis , 2 from the Krebs cycle , and about 34 from the electron transport system ) . However , this maximum yield is never quite reached because of losses due to leaky membranes as well as the cost of moving pyruvate and ADP into the mitochondrial matrix , and current estimates range around 29 to 30 ATP per glucose . Aerobic metabolism is up to 15 times more efficient than anaerobic metabolism ( which yields 2 molecules ATP per 1 molecule glucose ) . However some anaerobic organisms , such as methanogens are able to continue with anaerobic respiration , yielding more ATP by using other inorganic molecules ( not oxygen ) as final electron acceptors in the electron transport chain . They share the initial pathway of glycolysis but aerobic metabolism continues with the Krebs cycle and oxidative phosphorylation . The post-glycolytic reactions take place in the mitochondria in eukaryotic cells , and in the cytoplasm in prokaryotic cells . Aerobic respiration summed up 1 . Glycolysis : -- - 2 ATPs + Glucose → 2 Pyruvic Acid + 4 Hydrogen + 4 ATPs 2 . Formation of Acetyl CoA : -- - 2 Pyruvic Acid + 2 CoA → 2 Acetyl CoA + 2 Carbon Dioxide + 2 Hydrogen 3 . Krebs Cycle : -- - 2 Acetyl CoA + 3 O2 → 6 Hydrogen + 4 Carbon Dioxide + 2 ATPs 4 . Electron Transport System : -- - 12 Hydrogen + 3 O2 → 6 Water + 32 ATPs Overall Reaction : -- - Glucose + 6 O2 → 6 Carbon Dioxide + 6 Water + 36 ATPs Glycolysis Out of the cytoplasm it goes into the Krebs cycle with the acetyl CoA . It then mixes with CO and makes 2 ATP , NADH , and FADH . From there the NADH and FADH go into the NADH reductase , which produces the enzyme . The NADH pulls the enzyme 's electrons to send through the electron transport chain . The electron transport chain pulls H ions through the chain . From the electron transport chain , the released hydrogen ions make ADP for an end result of 32 ATP . O attracts itself to the left over electron to make water . Lastly , ATP leaves through the ATP channel and out of the mitochondria . Main article : Glycolysis Glycolysis is a metabolic pathway that takes place in the cytosol of cells in all living organisms . This pathway can function with or without the presence of oxygen . In humans , aerobic conditions produce pyruvate and anaerobic conditions produce lactate . In aerobic conditions , the process converts one molecule of glucose into two molecules of pyruvate ( pyruvic acid ) , generating energy in the form of two net molecules of ATP . Four molecules of ATP per glucose are actually produced , however , two are consumed as part of the preparatory phase . The initial phosphorylation of glucose is required to increase the reactivity ( decrease its stability ) in order for the molecule to be cleaved into two pyruvate molecules by the enzyme aldolase . During the pay - off phase of glycolysis , four phosphate groups are transferred to ADP by substrate - level phosphorylation to make four ATP , and two NADH are produced when the pyruvate are oxidized . The overall reaction can be expressed this way : Glucose + 2 NAD + 2 P + 2 ADP → 2 pyruvate + 2 NADH + 2 ATP + 2 H + 2 H O + heat Starting with glucose , 1 ATP is used to donate a phosphate to glucose to produce glucose 6 - phosphate . Glycogen can be converted into glucose 6 - phosphate as well with the help of glycogen phosphorylase . During energy metabolism , glucose 6 - phosphate becomes fructose 6 - phosphate . An additional ATP is used to phosphorylate fructose 6 - phosphate into fructose 1 , 6 - disphosphate by the help of phosphofructokinase . Fructose 1 , 6 - diphosphate then splits into two phosphorylated molecules with three carbon chains which later degrades into pyruvate . Glycolysis can be literally translated as `` sugar splitting '' . Oxidative decarboxylation of pyruvate Main article : Pyruvate decarboxylation Pyruvate is oxidized to acetyl - CoA and CO by the pyruvate dehydrogenase complex ( PDC ) . The PDC contains multiple copies of three enzymes and is located in the mitochondria of eukaryotic cells and in the cytosol of prokaryotes . In the conversion of pyruvate to acetyl - CoA , one molecule of NADH and one molecule of CO is formed . Citric acid cycle Main article : Citric acid cycle This is also called the Krebs cycle or the tricarboxylic acid cycle . When oxygen is present , acetyl - CoA is produced from the pyruvate molecules created from glycolysis . Once acetyl - CoA is formed , aerobic or anaerobic respiration can occur . When oxygen is present , the mitochondria will undergo aerobic respiration which leads to the Krebs cycle . However , if oxygen is not present , fermentation of the pyruvate molecule will occur . In the presence of oxygen , when acetyl - CoA is produced , the molecule then enters the citric acid cycle ( Krebs cycle ) inside the mitochondrial matrix , and is oxidized to CO while at the same time reducing NAD to NADH . NADH can be used by the electron transport chain to create further ATP as part of oxidative phosphorylation . To fully oxidize the equivalent of one glucose molecule , two acetyl - CoA must be metabolized by the Krebs cycle . Two waste products , H O and CO , are created during this cycle . The citric acid cycle is an 8 - step process involving 18 different enzymes and co-enzymes . During the cycle , acetyl - CoA ( 2 carbons ) + oxaloacetate ( 4 carbons ) yields citrate ( 6 carbons ) , which is rearranged to a more reactive form called isocitrate ( 6 carbons ) . Isocitrate is modified to become α - ketoglutarate ( 5 carbons ) , succinyl - CoA , succinate , fumarate , malate , and , finally , oxaloacetate . The net gain of high - energy compounds from one cycle is 3 NADH , 1 FADH , and 1 GTP ; the GTP may subsequently be used to produce ATP . Thus , the total yield from 1 glucose molecule ( 2 pyruvate molecules ) is 6 NADH , 2 FADH , and 2 ATP . Oxidative phosphorylation Main articles : Oxidative phosphorylation , Electron transport chain , Electrochemical gradient , and ATP synthase In eukaryotes , oxidative phosphorylation occurs in the mitochondrial cristae . It comprises the electron transport chain that establishes a proton gradient ( chemiosmotic potential ) across the boundary of inner membrane by oxidizing the NADH produced from the Krebs cycle . ATP is synthesized by the ATP synthase enzyme when the chemiosmotic gradient is used to drive the phosphorylation of ADP . The electrons are finally transferred to exogenous oxygen and , with the addition of two protons , water is formed . Efficiency of ATP production The table below describes the reactions involved when one glucose molecule is fully oxidized into carbon dioxide . It is assumed that all the reduced coenzymes are oxidized by the electron transport chain and used for oxidative phosphorylation . Step coenzyme yield ATP yield Source of ATP Glycolysis preparatory phase − 2 Phosphorylation of glucose and fructose 6 - phosphate uses two ATP from the cytoplasm . Glycolysis pay - off phase Substrate - level phosphorylation 2 NADH 3 or 5 Oxidative phosphorylation : Each NADH produces net 1.5 ATP ( instead of usual 2.5 ) due to NADH transport over the mitochondrial membrane Oxidative decarboxylation of pyruvate 2 NADH 5 Oxidative phosphorylation Krebs cycle Substrate - level phosphorylation 6 NADH 15 Oxidative phosphorylation 2 FADH Oxidative phosphorylation Total yield 30 or 32 ATP From the complete oxidation of one glucose molecule to carbon dioxide and oxidation of all the reduced coenzymes . Although there is a theoretical yield of 38 ATP molecules per glucose during cellular respiration , such conditions are generally not realized because of losses such as the cost of moving pyruvate ( from glycolysis ) , phosphate , and ADP ( substrates for ATP synthesis ) into the mitochondria . All are actively transported using carriers that utilize the stored energy in the proton electrochemical gradient . Pyruvate is taken up by a specific , low Km transporter to bring it into the mitochondrial matrix for oxidation by the pyruvate dehydrogenase complex . The phosphate carrier ( PiC ) mediates the electroneutral exchange ( antiport ) of phosphate ( H PO ; P ) for OH or symport of phosphate and protons ( H ) across the inner membrane , and the driving force for moving phosphate ions into the mitochondria is the proton motive force . The ATP - ADP translocase ( also called adenine nucleotide translocase , ANT ) is an antiporter and exchanges ADP and ATP across the inner membrane . The driving force is due to the ATP ( − 4 ) having a more negative charge than the ADP ( − 3 ) , and thus it dissipates some of the electrical component of the proton electrochemical gradient . The outcome of these transport processes using the proton electrochemical gradient is that more than 3 H are needed to make 1 ATP . Obviously this reduces the theoretical efficiency of the whole process and the likely maximum is closer to 28 -- 30 ATP molecules . In practice the efficiency may be even lower because the inner membrane of the mitochondria is slightly leaky to protons . Other factors may also dissipate the proton gradient creating an apparently leaky mitochondria . An uncoupling protein known as thermogenin is expressed in some cell types and is a channel that can transport protons . When this protein is active in the inner membrane it short circuits the coupling between the electron transport chain and ATP synthesis . The potential energy from the proton gradient is not used to make ATP but generates heat . This is particularly important in brown fat thermogenesis of newborn and hibernating mammals . Stoichiometry of aerobic respiration and most known fermentation types in eucaryotic cell . Numbers in circles indicate counts of carbon atoms in molecules , C6 is glucose C H O , C1 carbon dioxide CO . Mitochondrial outer membrane is omitted . According to some of newer sources the ATP yield during aerobic respiration is not 36 -- 38 , but only about 30 -- 32 ATP molecules / 1 molecule of glucose , because : ATP : NADH + H and ATP : FADH ratios during the oxidative phosphorylation appear to be not 3 and 2 , but 2.5 and 1.5 respectively . Unlike in the substrate - level phosphorylation , the stoichiometry here is difficult to establish . ATP synthase produces 1 ATP / 3 H. However the exchange of matrix ATP for cytosolic ADP and Pi ( antiport with OH or symport with H ) mediated by ATP -- ADP translocase and phosphate carrier consumes 1 H / 1 ATP as a result of regeneration of the transmembrane potential changed during this transfer , so the net ratio is 1 ATP : 4 H . The mitochondrial electron transport chain proton pump transfers across the inner membrane 10 H / 1 NADH + H ( 4 + 2 + 4 ) or 6 H / 1 FADH ( 2 + 4 ) . So the final stoichiometry is 1 NADH + H : 10 H : 10 / 4 ATP = 1 NADH + H : 2.5 ATP 1 FADH : 6 H : 6 / 4 ATP = 1 FADH : 1.5 ATP ATP : NADH + H coming from glycolysis ratio during the oxidative phosphorylation is 1.5 , as for FADH , if hydrogen atoms ( 2H + 2e ) are transferred from cytosolic NADH + H to mitochondrial FAD by the glycerol phosphate shuttle located in the inner mitochondrial membrane . 2.5 in case of malate - aspartate shuttle transferring hydrogen atoms from cytosolic NADH + H to mitochondrial NAD So finally we have , per molecule of glucose Substrate - level phosphorylation : 2 ATP from glycolysis + 2 ATP ( directly GTP ) from Krebs cycle Oxidative phosphorylation 2 NADH + H from glycolysis : 2 × 1.5 ATP ( if glycerol phosphate shuttle transfers hydrogen atoms ) or 2 × 2.5 ATP ( malate - aspartate shuttle ) 2 NADH + H from the oxidative decarboxylation of pyruvate and 6 from Krebs cycle : 8 × 2.5 ATP 2 FADH from the Krebs cycle : 2 × 1.5 ATP Altogether this gives 4 + 3 ( or 5 ) + 20 + 3 = 30 ( or 32 ) ATP per molecule of glucose The total ATP yield in ethanol or lactic acid fermentation is only 2 molecules coming from glycolysis , because pyruvate is not transferred to the mitochondrion and finally oxidized to the carbon dioxide ( CO ) , but reduced to ethanol or lactic acid in the cytoplasm . Fermentation Main article : Fermentation Without oxygen , pyruvate ( pyruvic acid ) is not metabolized by cellular respiration but undergoes a process of fermentation . The pyruvate is not transported into the mitochondrion , but remains in the cytoplasm , where it is converted to waste products that may be removed from the cell . This serves the purpose of oxidizing the electron carriers so that they can perform glycolysis again and removing the excess pyruvate . Fermentation oxidizes NADH to NAD+ so it can be re-used in glycolysis . In the absence of oxygen , fermentation prevents the buildup of NADH in the cytoplasm and provides NAD+ for glycolysis . This waste product varies depending on the organism . In skeletal muscles , the waste product is lactic acid . This type of fermentation is called lactic acid fermentation . In strenuous exercise , when energy demands exceed energy supply , the respiratory chain can not process all of the hydrogen atoms joined by NADH . During anaerobic glycolysis , NAD+ regenerates when pairs of hydrogen combine with pyruvate to form lactate . Lactate formation is catalyzed by lactate dehydrogenase in a reversible reaction . Lactate can also be used as an indirect precursor for liver glycogen . During recovery , when oxygen becomes available , NAD+ attaches to hydrogen from lactate to form ATP . In yeast , the waste products are ethanol and carbon dioxide . This type of fermentation is known as alcoholic or ethanol fermentation . The ATP generated in this process is made by substrate - level phosphorylation , which does not require oxygen . Fermentation is less efficient at using the energy from glucose : only 2 ATP are produced per glucose , compared to the 38 ATP per glucose nominally produced by aerobic respiration . This is because the waste products of fermentation still contain chemical potential energy that can be released by oxidation . Ethanol , for example , can be burned in an internal combustion engine like gasoline . Glycolytic ATP , however , is created more quickly . For prokaryotes to continue a rapid growth rate when they are shifted from an aerobic environment to an anaerobic environment , they must increase the rate of the glycolytic reactions . For multicellular organisms , during short bursts of strenuous activity , muscle cells use fermentation to supplement the ATP production from the slower aerobic respiration , so fermentation may be used by a cell even before the oxygen levels are depleted , as is the case in sports that do not require athletes to pace themselves , such as sprinting . Anaerobic respiration Main article : Anaerobic respiration Cellular respiration is the process by which biological fuels are oxidised in the presence of an inorganic electron acceptor ( such as oxygen ) to produce large amounts of energy , to drive the bulk production of ATP . Anaerobic respiration is used by some microorganisms in which neither oxygen ( aerobic respiration ) nor pyruvate derivatives ( fermentation ) is the final electron acceptor . Rather , an inorganic acceptor such as sulfate or nitrate is used . Such organisms are typically found in unusual places such as underwater caves or near hydrothermal vents at the bottom of the ocean . See also Maintenance respiration : maintenance as a functional component of cellular respiration Pasteur point Respirometry : research tool to explore cellular respiration Tetrazolium chloride : cellular respiration indicator Complex 1 : NADH : ubiquinone oxidoreductes References Jump up ^ Bailey , Regina . `` Cellular Respiration '' . Archived from the original on 2012 - 05 - 05 . ^ Jump up to : Rich , P.R. ( 2003 ) . `` The molecular machinery of Keilin 's respiratory chain '' . Biochemical Society Transactions . 31 ( Pt 6 ) : 1095 -- 1105 . doi : 10.1042 / BST0311095 . PMID 14641005 . Jump up ^ Reece1 Urry2 Cain3 Wasserman4 Minorsky5 Jackson6 , Jane1 Lisa2 Michael3 Steven4 Peter5 Robert6 ( 2010 ) . Campbell Biology Ninth Edition . Pearson Education , Inc. p. 168 . ^ Jump up to : `` Cellular Respiration '' ( PDF ) . Archived ( PDF ) from the original on 2017 - 05 - 10 . Jump up ^ Porter , R. ; Brand , M. ( 1 September 1995 ) . `` Mitochondrial proton conductance and H+ / O ratio are independent of electron transport rate in isolated hepatocytes '' . The Biochemical Journal ( Free full text ) . 310 ( Pt 2 ) : 379 -- 382 . doi : 10.1042 / bj3100379 . ISSN 0264 - 6021 . PMC 1135905 . PMID 7654171 . ^ Jump up to : Stryer , Lubert ( 1995 ) . Biochemistry ( fourth ed . ) . New York -- Basingstoke : W.H. Freeman and Company . ISBN 978 - 0716720096 . External links A detailed description of respiration vs. fermentation Kimball 's online resource for cellular respiration Cellular Respiration and Fermentation at Clermont College Library resources about Cellular respiration Resources in your library Metabolism , catabolism , anabolism General Metabolic pathway Metabolic network Primary nutritional groups Energy metabolism Aerobic respiration Glycolysis → Pyruvate decarboxylation → Citric acid cycle → Oxidative phosphorylation ( electron transport chain + ATP synthase ) Anaerobic respiration Electron acceptors are other than oxygen Fermentation Glycolysis → Substrate - level phosphorylation ABE Ethanol Lactic acid Specific paths Protein metabolism Protein synthesis Catabolism Carbohydrate metabolism ( carbohydrate catabolism and anabolism ) Human Glycolysis ⇄ Gluconeogenesis Glycogenolysis ⇄ Glycogenesis Pentose phosphate pathway Fructolysis Galactolysis Glycosylation N - linked O - linked Nonhuman Photosynthesis Anoxygenic photosynthesis Chemosynthesis Carbon fixation Xylose metabolism Radiotrophism Lipid metabolism ( lipolysis , lipogenesis ) Fatty acid metabolism Fatty acid degradation ( Beta oxidation ) Fatty acid synthesis Other Steroid metabolism Sphingolipid metabolism Eicosanoid metabolism Ketosis Reverse cholesterol transport Amino acid Amino acid synthesis Urea cycle Nucleotide metabolism Purine metabolism Nucleotide salvage Pyrimidine metabolism Other Metal metabolism Iron metabolism Ethanol metabolism Metabolism map Carbon Fixation Photo - respiration Pentose Phosphate Pathway Citric Acid Cycle Glyoxylate Cycle Urea Cycle Fatty Acid Synthesis Fatty Acid Elongation Beta Oxidation Peroxisomal Beta Oxidation Glyco - genolysis Glyco - genesis Glyco - lysis Gluconeo - genesis Decarb - oxylation Fermentation Keto - lysis Keto - genesis feeders to Gluconeo - genesis Direct / C4 / CAM Carbon Intake Light Reaction Oxidative Phosphorylation Amino Acid Deamination Citrate Shuttle Lipogenesis Lipolysis Steroidogenesis MVA Pathway MEP Pathway Shikimate Pathway Transcription & Replication Translation Proteolysis Glycosy - lation Sugar Acids Double / Multiple Sugars & Glycans Simple Sugars Inositol - P Amino Sugars & Sialic Acids Nucleotide Sugars Hexose - P Triose - P Glycerol P - glycerates Pentose - P Tetrose - P Propionyl - CoA Succinate Acetyl - CoA Pentose - P P - glycerates Glyoxylate Photosystems Pyruvate Lactate Acetyl - CoA Citrate Oxalo - acetate Malate Succinyl - CoA α - Keto - glutarate Ketone Bodies Respiratory Chain Serine Group Alanine Branched - chain Amino Acids Aspartate Group Homoserine Group & Lysine Glutamate Group & Proline Arginine Creatine & Polyamines Ketogenic & Glucogenic Amino Acids Amino Acids Shikimate Aromatic Amino Acids & Histidine Ascorbate ( Vitamin C ) δ - ALA Bile Pigments Hemes Cobalamins ( Vitamin B ) Various Vitamin B 's Calciferols ( Vitamin D ) Retinoids ( Vitamin A ) Quinones ( Vitamin K ) & Carotenoids ( Vitamin E ) Cofactors Vitamins & Minerals Antioxidants PRPP Nucleotides Nucleic Acids Proteins Glycoproteins & Proteoglycans Chlorophylls MEP MVA Acetyl - CoA Polyketides Terpenoid Backbones Terpenoids & Carotenoids ( Vitamin A ) Cholesterol Bile Acids Glycero - phospholipids Glycerolipids Acyl - CoA Fatty Acids Glyco - sphingolipids Sphingolipids Waxes Polyunsaturated Fatty Acids Neurotransmitters & Thyroid Hormones Steroids Endo - cannabinoids Eicosanoids Major metabolic pathways in metro - style map . Click any text ( name of pathway or metabolites ) to link to the corresponding article . Single lines : pathways common to most lifeforms . Double lines : pathways not in humans ( occurs in e.g. plants , fungi , prokaryotes ) . Orange nodes : carbohydrate metabolism . Violet nodes : photosynthesis . Red nodes : cellular respiration . Pink nodes : cell signaling . Blue nodes : amino acid metabolism . Grey nodes : vitamin and cofactor metabolism . Brown nodes : nucleotide and protein metabolism . Green nodes : lipid metabolism . Metabolism portal Retrieved from `` https://en.wikipedia.org/w/index.php?title=Cellular_respiration&oldid=830663802 '' Categories : Cellular respiration Metabolism Plant physiology Hidden categories : Wikipedia indefinitely move - protected pages Articles needing additional references from September 2014 All articles needing additional references Talk View source Contents About Wikipedia বাংলা Bân - lâm - gú Беларуская Български Bosanski Brezhoneg Català Čeština Cymraeg Dansk Deutsch Eesti Ελληνικά Español Euskara فارسی Français Galego 한국어 Հայերեն हिन्दी Hrvatski Bahasa Indonesia Íslenska Italiano עברית Қазақша Kreyòl ayisyen Lietuvių Македонски Bahasa Melayu Nederlands Norsk Occitan Polski Português Română Русский සිංහල Simple English Slovenščina کوردی Српски / srpski Srpskohrvatski / српскохрватски Basa Sunda Suomi Svenska தமிழ் ไทย Türkçe Українська 粵語 中文 45 more Edit links This page was last edited on 16 March 2018 , at 06 : 02 . Text is available under the Creative Commons Attribution - ShareAlike License ; additional terms may apply . By using this site , you agree to the Terms of Use and Privacy Policy . Wikipedia ® is a registered trademark of the Wikimedia Foundation , Inc. , a non-profit organization . About Wikipedia

Question: how many co2 molecules are produced in aerobic respiration