The learning and foraging parameters δ i, ϕ i, σ i, ε i, λ i, γ i, ω i, τ i, are specific to forager i. Parameter combinations that lead to greater energy levels lead to faster rates of reproduction. An offspring inherits its parent’s parameters, with a chance of mutation (0.05). In case of mutation, a new parameter value is drawn from a normal distribution centered on the parent’s parameter value, and with a standard deviation that is one fifth of the maximum value of the parameter (see Table 1). Thus parameters can vary between individuals and can evolve over time via inheritance to offspring, mutation and natural selection. The mutation rate was selected operationally such that parameters evolve consistently within a reasonable time frame.