Ancestral range estimation (ARE) for Copits BEAST using BioGeoBEARS under the DIVALIKE model. Labeled nodes (1 to 5, as referred to Fig. 2), with 95% highest posterior densities (grey bars), are discussed in the text. The estimated ancestral ranges with the highest ML probability are shown by boxes on each node. Additional file 2: Figure S1 provides all ARE per node with pies. A pie is placed in this figure at the root with the highest probability less than 50%. The depictions of temperature (in red) and sea level (in black) changes are modified from Zachos et al.  and Haq et al. , respectively. Plio., Pliocene; Plt., Pleistocene