EMT of the proportions of macronutrients (protein = P, carbohydrate = C, and lipid = L) in seasonal (spring, summer, and autumn) brown bear diets in populations with natural diets versus those receiving anthropogenic subsidies. The geometric mean for each season is shown by a filled symbol surrounded by 90% and 99% confidence regions. For reference, the blue line represents the preferred optimal proportion of protein (17% ± 4) selected by captive bears