Given the high aAD variability of each carabid species among the sample sites, two threshold levels (‘‘central” and ‘‘nuclear”) were introduced with the exclusive intent of facilitating the interpretation of the data table (S1 Table; see ), i.e. a species is central in the site where it shows the maximum aAD, while it is nuclear in the site(s) where it shows an aAD equal to, or greater than, the mean aAD among the sites .