Although numerous studies found that over-expressing TPS and TPP genes could improve abiotic stress tolerance (Holmström et al., 1996; Jang et al., 2003; Schluepmann et al., 2004; Jiang et al., 2010), observed stress tolerance of transgenic lines did not correlate with trehalose amount (van Dijken et al., 2004). Researchers have investigated function of plant trehalose biosynthetic enzymes by identifying mutations in Arabidopsis TPS1 gene and seeing which were still effective in recovering homologous gene function for yeast mutants deficient in TPS functions (Blázquez et al., 1998; Müller et al., 2001; Vogel et al., 2001). Beyond these stress response, recent intriguing evidence has implicated that TPS genes are important modulators in plant development and inflorescence architecture (Satoh-Nagasawa et al., 2006).