While we test a hypothesis generated by previous theory, our study is not simply an extension of previous models. Here we directly contrast multiple SLMs in a relatively complex model with a large parameter space. To facilitate the exploration of parameter space, we use evolved parameter values for behavioural and learning parameters based on van der Post et al. . In this way parameters are optimized relative to foraging success, and the different SLMs are compared using parameters that derive from this standardized criterion. The earlier models did not use evolved parameters.