Considering the unique markers (the 4 959 bin markers), the highest marker saturation was found in genome A (39.3%), followed by genomes B (33.6%) and D (27.1%). The average distance between adjacent bin markers ranged from 0.6 cM for 6B to 1.5 cM for 4D, with an average of 0.9 cM per marker pair. Gaps greater than 20.0 cM but less than 30.0 cM were present in chromosomes 3 A (24.9 cM), 3B (21.9 cM) and 4D (20.4 cM); gaps greater than 10 cM but less than 20.0 cM were present in chromosomes 1 A (13.9 cM, 11.7 cM), 2 A (14.6 cM), 2B (10.6 cM), 2D (10.5 cM), 3 A (19.9 cM, 19.2 cM), 3D (12.1 cM), 4D (11.5 cM, 11.2 cM, 11.0 cM, 11.0 cM), 5 A (12.1 cM), 5D (13.2 cM), 6 A (19.2 cM), 6B (13.8 cM, 13.2 cM, 12.7 cM), 6D (14.6 cM, 13.6 cM, 10.9 cM, 10.7 cM), 7B (16.3 cM, 10.1 cM), and 7D (18.8 cM) (Fig. 2).