1) My main concern is with the central focus on the comparison between two isoforms of FLM (FLMb and FLMd). A recent study showed that the qPCR assays targeting these isoforms actually pick up other isoforms, many of which produce truncated proteins (Sureshkumar et al. 2016 Nature Plants). Here, the authors note that they can't even find a true FLMd transcript (out of 163 tested). It is not clear what if any conclusions can be drawn from the FLMd assay, and so devoting so much space throughout the article and figures to FLMd is not useful and distracting. On the other hand, this and earlier studies the FLMb assay does seem to be useful for assaying FLM activity, so I am comfortable with these results.