Analysis of molecular variance (AMOVA) and the genetic differentiation index (FST) were performed using the Arlequin software version 3.11 . Mantel test was used to verify the correlation between of Nei’s genetic distance and geographical distance  with 1,000 permutations using the NTSYS 2.01 software. The principal coordinates analysis (PCoA) for the observation of the graphical distribution of the samples was performed according to the genetic correlation with the NTSYS 2.01 software.