Seasonally, brown bear displayed significant variation in the proportion of macronutrients consumed, indicating a tolerance for a wide range of dietary macronutrient proportions throughout the active season thereby supporting our third hypothesis. Protein and lipid proportions become less codependent during autumn, which is consistent with the general pattern of decreasing carnivory combined with the consumption of high‐fat autumn food resources in some ecosystems, such as hard mast. The proportion of carbohydrate in bear diets was highest in the autumn, mostly due to the timing of fruit production, while some populations also consumed starchy roots during this period. However, diets of populations receiving anthropogenic subsidies were on average higher in the proportion of carbohydrates and lower in protein, across all seasons. For example, the diets of Greek (Paralikidis et al., 2010), Italian (Ciucci et al., 2014), and Slovakian (Stofik et al., 2013) bears were high in carbohydrate during summer, due to fruit and anthropogenic food consumption. Given that such foods allow bears to consume closer to preferred proportions of macronutrients, it is not surprising that anthropogenic foods are sources of bear–human conflict (Can, D'Cruze, Garshelis, Beecham, & Macdonald, 2014; Coogan & Raubenheimer, 2016; Morehouse & Boyce, 2017). Garbage, which was not considered in this study and seldom reported (e.g., Mattson, Blanchard, & Knight, 1991; Rigg & Gorman, 2005), would likely have a similar effect on diet proportions (Coogan & Raubenheimer, 2016). Conversely, the highly carnivorous natural diet of central Arctic bears (Gau et al., 2002) was highest in protein during the autumn.