The first component of the PCA.acomp, associated with differences in ratios of carbohydrate with both lipid and protein, explained 80.8% of variance in the macronutrient proportions of bear diets (Figures 1 and 2). The remaining second component, associated with lipid and protein ratios, explained the remaining 19.2% of variance. In the PCA.acomp biplot (Figure 2), the link distance between protein and carbohydrate was greatest, indicating that most relative variation occurs between these two macronutrients. Carbohydrate and lipid also share a large amount of relative variation. The shorter link between protein and lipid indicates that their component ratio was relatively more constant.