Biophysical analysis of the model was carried out according to standard theory (Jack et al., 1975). The model electrotonic properties were analyzed using NEURON functions yielding the input impedance, Zin, for each model section and the corresponding signal attenuation between e.g., a dendritic sections and the soma, A = Vdend/Vsoma. This allowed to determine the electrotonic distance L = ln(A). This definition coincided with the more classical one (L = anatomical distance/length constant) defined when a neuron can be reduced through the 3/2 power branching rule to an equivalent cylinder. The NEURON definition proves particularly useful since PCs do not follow the 3/2 power branching rule (Hines and Carnevale, 2001; Hines et al., 2007, 2009).