Biofortification is a sustainable approach that is based on the fortification of crops through the utilization of nutrient-rich fertilizers, breeding or plant engineering strategies in order to produce and/or accumulate nutritionally important molecules . Among these techniques, conventional breeding could show some disadvantages since it requires long time to introduce traits of interest into local varieties, whereas, through genetic engineering, novel genes can be directly introduced into the genome of transgenic plants. Furthermore, genetic engineering allows for combining several traits in the same plants and nutritional traits can be targeted to specific plant organs .