Besides petal shape and size, floral symmetry significantly affects the ornamental value of flowers. So far, three transcription factors (TFs) that determine the floral symmetry are identified, including CYCLOIDEA (CYC) from the TCP family (TEOSINTE BRANCHED1/CYCLOIDEA/PROLIFERATING CELL NUCLEAR ANTIGEN FACTOR), DIVARICATA (DIV) and RADIALIS (RAD) from the MYB family (Luo et al. 1996; Doebley et al. 1997; Almeida et al. 1997; Galego and Almeida 2002; Corley et al. 2005; Costa et al. 2005). Of these genes, CYC and its orthologues establish the floral monosymmetry through specifying dorsal flower identity (Luo et al. 1996, 1999; Feng et al. 2006; Wang et al. 2008). Studies on floral symmetry genes mainly focused on species with highly derived morphologies in eudicots, whereas only a few studies are available for monocots (Luo et al. 1996, 1999; Feng et al. 2006; Wang et al. 2008; Yuan et al. 2009; Bartlett and Specht 2011; Preston and Hileman 2012; Hoshino et al. 2014). Orchidaceae is characterized by highly specialized zygomorphic flowers. Studies on orchid flower symmetry is very limited (Paolo et al. 2015). Overall, the detailed molecular mechanism or the genetic elements in the regulation of floral organ specification and development in C. faberi remains elusive.