Furthermore, there is implicit evidence that proportions and amounts of dietary macronutrients interact to affect brown bear population dynamics, as local population density has been related to spatial patterns in the amounts of both ungulates (source of protein and lipid) and fruit (source of carbohydrate) together (Nielsen, Larsen, Stenhouse, & Coogan, 2017). Following the above example, it is important to note that both the proportions and amounts of macronutrients interact to produce biological outcomes; therefore, investigating the relationships between the amounts and proportions of dietary macronutrients, and their possible population‐level effects, is an important area of future research. On the other hand, in many animals, dietary macronutrient proportions predict absolute amounts eaten (Raubenheimer, Machovsky‐Capuska, Gosby, & Simpson, 2014).