For flowering time analyses, plants were grown under long day-conditions with 16 hr white light (110–130 μmol m−2 s−1)/8 hr dark in MLR-351 SANYO growth chambers (Ewald, Bad Nenndorf, Germany). Plants were randomly arranged in trays and trays were rearranged every day. Water was supplied by subirrigation. Flowering time was quantified by counting rosette leaf numbers (RLN). Consistent with previous reports, we observed a strong correlation between days to bolting and rosette leaf number of Arabidopsis accessions (Figure 8—figure supplement 2C) (Atwell et al., 2010). Student’s t-tests (normally distributed values), Wilcoxon rank tests (not normally distributed values), and multiple regression models were calculated with R (http://www.r-project.org/).