Leading from this, we propose that future research examine spatially explicit factors influencing the macronutrient proportions of diet. Increasing carnivory has been hypothesized as a general adaptation to an increase in latitude in omnivorous mammals (Vulla et al., 2009); however, other works have suggested that dietary patterns are better explained by spatially explicit environmental factors (Gaston, Chown, & Evans, 2008). Patterns in brown bear diet were better explained by climatic rather than geographic factors (Bojarska & Selva, 2012). It would be interesting to examine the relationships between such factors and nutrition. Furthermore, we suggest the effects of anthropogenic food subsidies on brown bears at the levels of individuals, populations, and communities deserve more research.