FM designed and performed the research, analyzed the data and wrote the manuscript. KN, YD, and KH performed the research and analyzed the data. FM, SY, YU, MS, MI, NS (Sakon), NS (Shigemoto), RO, and AO contributed samples and analyzed the data; and HK and KK designed and supervised the research, analyzed the data, and wrote the manuscript. All authors contributed, read, and approved the manuscript.