Morphological terminology follows Carpenter and Cumming (1985), Yamane (1990), and Carpenter and Garcete-Barrett (2003). Observations and descriptions were made with an Olympus SZX11 stereomicroscope and fluorescent lamps. Photographic images were made with Keyence VHX-5000 digital microscope (NWUX, Xi’an), Olympus SZX 12 stereomicroscope with analySIS Soft Imaging System software (RMNH, Leiden), and Microptics-USA/Visionary Digital photomicrographic system, developed by Roy Larimer, multiple layers stacked using Helicon Focus (AMNH, New York).