A high level of vitamin E was also achieved in transgenic Brassica napus plant seeds by overexpressing Arabidopsis genes encoding hydroxyl phenyl pyruvate dioxygenases, alone or in combination with genes encoding chimeric homogentisate phytyl transferase and tocopherol cyclase . To enhance the amount of carotenoids in Brassica napus plants, seven key enzyme genes that are involved in ketocarotenoid synthesis, isolated from the soil bacterium Pantoea ananatis, and from the marine bacteria Brevundimonas and Paracoccus strain, were expressed in transgenic plants . In another paper, the Arabidopsis regulatory gene Production of Anthocyanin Pigment 1 (AtPAP1) was expressed in Brassica napus, thus obtaining a significant increase in the levels of the phenolic compounds cyanidin, pelargonidin, and quercetin .