The PCoA allowed for the visualization of the spatial structure of the population distribution. Thus, the populations Coxilha, Panambi, and Campos Novos were isolated in quadrants two and three, although Panambi and Campos Novos contained individuals that were far from the center of the population cluster (Fig 3). The Guarapuava and Porto Barreiro, and Xanxerê and Marialva populations overlapped by more than 85% of their occupied areas in the quadrants (Fig 3).