To the south and west of England, William granted lands to groups of Breton nobles who were more fractious than united, whether in Brittany or in England. This group had no one singular leader who could serve a unifying role . Their numbers included Herve of Leon, the viscount of Leon, who was briefly the Earl of Wiltshire. The third group were those nobles associated with the Richmond-Penthièvre family. Odo of Penthièvre was a principal antagonist of Conan II. Under William I, at three of Odo of Penthièvre's sons (Alan, Stephen and Brien) were granted substantial lands in England, including the Honour of Richmond, lands in Suffolk, and the Earldoms of Richmond and Cornwall. Also in 1066, Hawise succeeded her brother Conan II as hereditary Duchess of Brittany. She married Hoël of Cornouaille. Hoel ruled as Hoel II, Duke of Brittany and started the House of Kernev which continued to rule Brittany until 1156 (see below). Hoël inherited a divided Brittany which was split into six largely independent regions: Rennes, Panthièvre, Léon, Cornouaille, the Broërec, and Nantes. Ducal power was non-existent in Panthièvre and Léon, and even in the rest of Brittany the duke's powers did not extend beyond his own personal lands. The barons in Brittany did not feel that they were in any way vassals of the duke, owing him service in exchange for their lands, and only attended the ducal courts when they felt it was in their own interests to do so. However, the historic rivalry between Brittany and Normandy resurfaced at the close of the 11th century. By 1075, Hoèl returned to the traditional Breton policy of opposing Norman expansion with an alliance with the young king Philip I of France. Ralph de Gael, in exile in Brittany after the unsuccessful 1075 rebellion in England, led incursions into Normandy from his base in Dol. In 1076, King William of England retaliated by leading an army into Brittany to eject Ralph, but was met with a rare defeat by an allied army of Bretons and French forces. In the peace negotiations which followed William offered his second daughter Constance in marriage to the Breton heir Alan IV, though nothing came of the betrothal at the time. By 1086, Alan IV was forced to abandon his duchy after an invasion launched by William I of England.