Huxley's paper was rejected by the Royal Society, and this became one of the many theories to be advanced and dropped by the early exponents of evolution. Despite rejection by Huxley and the science community, the paper is sometimes cited in support of racialism. Along with Darwin, Huxley was a monogenist, the belief that all humans are part of the same species, with morphological variations emerging out of an initial uniformity. (Stepan, p. 44). This view contrasts polygenism, the theory that each race is actually a separate species with separate sites of origin. Despite Huxley's monogenism and his abolitionism on ethical grounds, Huxley assumed a hierarchy of innate abilities, a stance evinced in papers such as "Emancipation Black and White" and his most famous paper, "Evolution and Ethics". In the former, he writes that the "highest places in the hierarchy of civilization will assuredly not be within the reach of our dusky cousins, though it is by no means necessary that they should be restricted to the lowest". (Stepan, p. 79–80). Though Charles Darwin's evolutionary theory was set forth in 1859 upon publication of "On the Origin of Species", this work was largely absent of explicit reference to Darwin's theory applied to man. This application by Darwin would not become explicit until 1871 with the publication of his second great book on evolution, "The Descent of Man, and Selection in Relation to Sex". Darwin's publication of this book occurred within the heated debates between advocates of monogeny, who held that all races came from a common ancestor, and advocates of polygeny, who held that the races were separately created. Darwin, who had come from a family with strong abolitionist ties, had experienced and was disturbed by cultures of slavery during his voyage on the Beagle years earlier. Noted Darwin biographers Adrian Desmond and James Moore argue that Darwin's writings on evolution were not only influenced by his abolitionist tendencies, but also his belief that non-white races were equal in regard to their intellectual capacity as white races, a belief which had been strongly disputed by scientists such as Morton, Agassiz and Broca, all noted polygenists. By the late 1860s, however, Darwin's theory of evolution had been thought to be compatible with the polygenist thesis (Stepan 1982). Darwin thus used "Descent of Man" to disprove the polygenist thesis and end the debate between polygeny and monogeny once and for all.