Cichlids are one of the largest vertebrate families in the world. They are most diverse in Africa and South America. Africa alone is estimated to host at least 1,600 species. Central America and Mexico have about 120 species, as far north as the Rio Grande in southern Texas. Madagascar has its own distinctive species ("Katria", "Oxylapia", "Paratilapia", "Paretroplus", "Ptychochromis", and "Ptychochromoides"), only distantly related to those on the African mainland. Native cichlids are largely absent in Asia, except for 9 species in Israel, Lebanon, and Syria ("Astatotilapia flaviijosephi", "Oreochromis aureus", "O. niloticus", "Sarotherodon galilaeus", "Coptodon zillii", and "Tristramella" spp.), two in Iran ("Iranocichla"), and three in India and Sri Lanka ("Etroplus" and "Pseudetroplus"). If disregarding Trinidad and Tobago (where the few native cichlids are members of genera that are widespread in the South American mainland), the three species from the genus "Nandopsis" are the only cichlids from the Antilles in the Caribbean, specifically Cuba and Hispaniola. Europe, Australia, Antarctica, and North America north of the Rio Grande drainage have no native cichlids, although in Florida, Hawaii, Japan, northern Australia and elsewhere, feral populations of cichlids have become established as exotics. Although most cichlids are found at relatively shallow depths, several exceptions do exist. The deepest known occurrence are "Trematocara" at more than below the surface in Lake Tanganyika. Others found in relatively deep waters include species such as "Alticorpus macrocleithrum" and "Pallidochromis tokolosh" down to below the surface in Lake Malawi, and the whitish (nonpigmented) and blind "Lamprologus lethops", which is believed to live as deep as below the surface in the Congo River. Cichlids are less commonly found in brackish and saltwater habitats, though many species tolerate brackish water for extended periods; "Mayaheros urophthalmus", for example, is equally at home in freshwater marshes and mangrove swamps, and lives and breeds in saltwater environments such as the mangrove belts around barrier islands.