The Phanerozoic is divided into three eras: the Paleozoic, Mesozoic, and Cenozoic, which are further subdivided into 12 periods. The Paleozoic features the rise of fish, amphibians and reptiles. The Mesozoic is ruled by the reptiles, and features the evolution of mammals, and more famously, dinosaurs, including birds. The Cenozoic is the time of the mammals, and more recently, humans. The Paleozoic is a time in Earth's history when complex life forms evolved, took their first breath of oxygen on dry land, and when the forerunners of all life on Earth began to diversify. There are six periods in the Paleozoic era: Cambrian, Ordovician, Silurian, Devonian, Carboniferous and Permian. The Cambrian is the first period of the Paleozoic Era and ran from . The Cambrian sparked a rapid expansion in evolution in an event known as the Cambrian explosion during which the greatest number of creatures evolved in a single period in the history of Earth. Plants like algae evolved, and the fauna was dominated by armored arthropods, such as trilobites. Almost all marine phyla evolved in this period. During this time, the super-continent Pannotia began to break up, most of which later recombined into the super-continent Gondwana. The Ordovician spans from 485 million years to 444 million years ago. The Ordovician was a time in Earth's history in which many species still prevalent today evolved, such as primitive fish, cephalopods, and coral. The most common forms of life, however, were trilobites, snails and shellfish. More importantly, the first arthropods crept ashore to colonize Gondwana, a continent empty of animal life. By the end of the Ordovician, Gondwana had moved from the equator to the South Pole, and Laurentia had collided with Baltica, closing the Iapetus Ocean. The glaciation of Gondwana resulted in a major drop in sea level, killing off all life that had established along its coast. Glaciation caused a snowball Earth, leading to the Ordovician–Silurian extinction, during which 60% of marine invertebrates and 25% of families became extinct. This is considered the first mass extinction and the second deadliest in the history of Earth. The Silurian spans from 444 million years to 419 million years ago, which saw a warming from snowball Earth.