Tidal mixing forces exceed river output, resulting in a well-mixed water column and the disappearance of the vertical salinity gradient. The freshwater-seawater boundary is eliminated due to the intense turbulent mixing and eddy effects. The lower reaches of Delaware Bay and the Raritan River in New Jersey are examples of vertically homogenous estuaries. Inverse estuaries occur in dry climates where evaporation greatly exceeds the inflow of freshwater. A salinity maximum zone is formed, and both riverine and oceanic water flow close to the surface towards this zone. This water is pushed downward and spreads along the bottom in both the seaward and landward direction. An example of an inverse estuary is Spencer Gulf, South Australia. Estuary type varies dramatically depending on freshwater input, and is capable of changing from a wholly marine embayment to any of the other estuary types. The most important variable characteristics of estuary water are the concentration of dissolved oxygen, salinity and sediment load. There is extreme spatial variability in salinity, with a range of near-zero at the tidal limit of tributary rivers to 3.4% at the estuary mouth. At any one point, the salinity will vary considerably over time and seasons, making it a harsh environment for organisms. Sediment often settles in intertidal mudflats which are extremely difficult to colonize. No points of attachment exist for algae, so vegetation based habitat is not established. Sediment can also clog feeding and respiratory structures of species, and special adaptations exist within mudflat species to cope with this problem. Lastly, dissolved oxygen variation can cause problems for life forms. Nutrient-rich sediment from man-made sources can promote primary production life cycles, perhaps leading to eventual decay removing the dissolved oxygen from the water; thus hypoxic or anoxic zones can develop. Estuaries are incredibly dynamic systems, where temperature, salinity, turbidity, depth and flow all change daily in response to the tides. This dynamism makes estuaries highly productive habitats, but also make it difficult for many species to survive year-round. As a result, estuaries large and small experience strong seasonal variation in their fish communities. In winter, the fish community is dominated by hardy marine residents, and in summer a variety of marine and anadromous fishes move into and out of estuaries, capitalizing on their high productivity. Estuaries provide critical habitat to a variety of species that rely on estuaries for life-cycle completion.