Spatial organization  Spatial organization can be observed when components of an abiotic or biological group are arranged non-randomly in space. Abiotic patterns, such as the ripple formations in sand dunes or the oscillating wave patterns of the Belousov–Zhabotinsky reaction emerge after thousands of particles interact millions of times. On the other hand, individuals in biological groups may be arranged non-randomly due to selfish behavior, dominance interactions, or cooperative behavior. Hamilton (1971) proposed that in a non-related "herd" of animals, spatial organization is likely a result of the selfish interests of individuals trying to acquire food or avoid predation. On the other hand, spatial arrangements have also been observed among highly related members of eusocial groups, suggesting that the arrangement of individuals may provide some advantage for the group. Individuals in a social insect colony can be spatially organized, or arranged non-randomly inside the nest. These miniature territories, or spatial fidelity zones have been described in honey bees ("Apis mellifera"), ants ("Odontomachus brunneus"; "Temnothorax albipennis"; "Pheidole dentata"), and paper wasps ("Polistes dominulus", "Ropalidia revolutionalis"). While residing in these zones, workers perform the task appropriate to the area they reside. For example, individuals that remain in the center of an ant nest are more likely to feed larvae, whereas individuals found at the periphery of the nest are more likely to forage. Wilson proposed that by remaining in small, non-random areas inside the nest, the distance an individual moves between tasks may be minimized, and overall colony efficiency would increase. There are a variety of ways in which individuals can divide space inside a nest. According to the “foraging-for-work” hypothesis, adult workers begin performing tasks in the area of the nest where they emerged, and gradually move towards the periphery of the nest as demands to perform particular tasks change. This hypothesis is based on two observations: "(1) that there is spatial structure in the layout of tasks in social insect colonies and (2) that workers first become adults in or around the center of the nest". Individuals can remain in an area for an extended period of time, as long as tasks need to be performed there. Over time, an individual's zone may shift as tasks are accomplished and workers search for other areas where tasks need to be performed. Honey bees, for example, begin their adult life caring for brood located in the area near where they emerged (i.e. nurse bees).