Although stringent water conservation measures have been implemented, the threat of severe shortfalls in the Colorado River basin continues to increase each year. After a much lower-than-average snowpack in the 2018 water year, Bureau of Reclamation officials projected the odds of an official shortage declaration in 2020 at 52 percent, in 2021 at 64 percent, and in 2022 at 68 percent. The Colorado River and its tributaries often nourish extensive corridors of riparian growth as they traverse the arid desert regions of the watershed. Although riparian zones represent a relatively small proportion of the basin and have been affected by engineering projects and river diversion in many places, they have the greatest biodiversity of any habitat in the basin. The most prominent riparian zones along the river occur along the lower Colorado below Davis Dam, especially in the Colorado River Delta, where riparian areas support 358 species of birds despite the reduction in freshwater flow and invasive plants such as tamarisk (salt cedar). Reduction of the delta's size has also threatened animals such as jaguars and the vaquita porpoise, which is endemic to the gulf. Human development of the Colorado River has also helped to create new riparian zones by smoothing the river's seasonal flow, notably through the Grand Canyon. More than 1,600 species of plants grow in the Colorado River watershed, ranging from the creosote bush, saguaro cactus, and Joshua trees of the Sonoran and Mojave Deserts to the forests of the Rocky Mountains and other uplands, composed mainly of ponderosa pine, subalpine fir, Douglas-fir and Engelmann spruce. Before logging in the 19th century, forests were abundant in high elevations as far south as the Mexico–U.S. border, and runoff from these areas nourished abundant grassland communities in river valleys. Some arid regions of the watershed, such as the upper Green River valley in Wyoming, Canyonlands National Park in Utah and the San Pedro River valley in Arizona and Sonora, supported extensive reaches of grassland roamed by large mammals such as buffalo and antelope as late as the 1860s. Near Tucson, Arizona, "where now there is only powder-dry desert, the grass once reached as high as the head of a man on horse back". Rivers and streams in the Colorado basin were once home to 49 species of native fish, of which 42 were endemic. Engineering projects and river regulation have led to the extinction of four species and severe declines in the populations of 40 species.