Fearing of what recently occurred, Axum shifted its capital near Agew] In the middle of the sixteenth century Adal Sultanate armies led by Harar leader Ahmed Gragn invaded Habesha lands in what is known as the ""Conquest of Habasha"". Following Gragn invasions the southern part of the Empire was lost to Ethiopia and scattered Habesha like the Gurage people were cut off from the rest of Abyssinia. In the late sixteenth century the nomadic Oromo people penetrated the Habesha plains occupying large territories during the Oromo migrations. Abyssinian warlords often competed with each other for dominance of the realm. The Amharas seemed to gain the upper hand with the accession of Yekuno Amlak of Ancient Bete Amhara in 1270, after defeating the Agaw lords of Lasta (in those days a non-Semitic-speaking region of Abyssinia)  The Gondarian dynasty, which since the 16th century had become the centre of Royal pomp and ceremony of Abyssinia, finally lost its influence as a result of the emergence of powerful regional lords, following the murder of Iyasu I, also known as Iyasu the Great. The decline in the prestige of the dynasty led to the semi-anarchic era of Zemene Mesafint ("Era of the Princes"), in which rival warlords fought for power and the Yejju Oromo "enderase"s ("regents") had effective control. The emperors were considered to be figureheads. Until a young man named Kassa Haile Giorgis also known as Emperor Tewodros brought end to "Zemene Mesafint" by defeating all his rivals and took the throne in 1855. The Tigrayans made only a brief return to the throne in the person of Yohannes IV in 1872, whose death in 1889 resulted in the power base shifting back to the dominant Amharic-speaking elite. His successor Menelik II an Emperor of Amhara origin seized power. League of Nations in 1935 reported that after the invasion of Menelik's forces into non Abyssinian lands of Somalis, Harari, Oromo, Sidama, Shanqella etc, the inhabitants were enslaved and heavily taxed by the gebbar system leading to depopulation. Some scholars consider the Amhara to have been Ethiopia's ruling elite for centuries, represented by the Solomonic line of Emperors ending in Haile Selassie I. Marcos Lemma and other scholars dispute the accuracy of such a statement, arguing that other ethnic groups have always been active in the country's politics.