Another find at Tenlik in eastern Zhetysu contained the grave of a high-ranking warrior, whose clothing had been decorated with around 100 golden bosses. Some scholars such as Peter B. Golden have proposed that the Wusun may have been identical with the people described by Herodotus (IV. 16–25) and in Ptolemy's "Geography" as Issedones (also "Issedoni", "Issedoi" or "Essedoni"). Their exact location of their country in Central Asia is unknown. The Issedones are "placed by some in Western Siberia and by others in Chinese Turkestan," according to E. D. Phillips. French historian Iaroslav Lebedynsky suggests that the Wusun may have been the Asii of "Geographica". A genetic study published in "Nature" in May 2018 examined the remains of four Wusun buried between ca. 300 BC and 100 BC. The sample of Y-DNA extracted belonged to haplogroup R1. The samples of mtDNA extracted belonged to C4a1, HV6, J1c5a and U5b2c. The authors of the study found that the Wusun and Kangju had less East Asian admixture than the Xiongnu and Sakas. Both the Wusun and Kangju were suggested to be descended from Western Steppe Herders (WSHs) of the Late Bronze Age who admixed with Siberian hunter-gatherers and peoples related to the Bactria–Margiana Archaeological Complex.