Few fossil deposits are known from South Asia, the suspected region of honey bee origin, and fewer still have been thoroughly studied. No "Apis" species existed in the New World during human times before the introduction of "A. mellifera" by Europeans. Only one fossil species is documented from the New World, "Apis nearctica", known from a single 14 million-year-old specimen from Nevada. The close relatives of modern honey bees – "e.g." bumblebees and stingless bees – are also social to some degree, and social behavior seems a plesiomorphic trait that predates the origin of the genus. Among the extant members of "Apis", the more basal species make single, exposed combs, while the more recently evolved species nest in cavities and have multiple combs, which has greatly facilitated their domestication. Most species have historically been cultured or at least exploited for honey and beeswax by humans indigenous to their native ranges. Only two species have been truly domesticated: "Apis mellifera" and "Apis cerana indica". "A. mellifera" has been cultivated at least since the time of the building of the Egyptian pyramids, and only that species has been moved extensively beyond its native range. Honey bees are the only extant members of the tribe Apini. Today's honey bees constitute three clades: "Micrapis" (dwarf honey bees), "Megapis" (giant honey bee), and "Apis" (domestic honey bees and close relatives). "Apis florea" and "Apis andreniformis" are small honey bees of southern and southeastern Asia. They make very small, exposed nests in trees and shrubs. Their stings are often incapable of penetrating human skin, so the hive and swarms can be handled with minimal protection. They occur largely sympatrically, though they are very distinct evolutionarily and are probably the result of allopatric speciation, their distribution later converging. Given that "A. florea" is more widely distributed and "A. andreniformis" is considerably more aggressive, honey is, if at all, usually harvested from the former only. They are the most ancient extant lineage of honey bees, maybe diverging in the Bartonian (some 40 million years ago or slightly later) from the other lineages, but do not seem to have diverged from each other a long time before the Neogene. "Apis florea" have smaller wing spans than its sister species.