Le plus gros dinosaure prédateur découvert en Europe était une
Torvosaurus gurneyi, probablement le plus gros dinosaure prédateur à avoir été retrouvé en Europe, est un carnivore particulièrement puissant qui se servait de ...
Thomas R. Holtz Jr.
En effet, pour reprendre l’expression de <anchor>Thomas R. Holtz Jr.</anchor>, paléontologue spécialiste des vertébrés à l’Université du Maryland, T. gurneyi était une « grosse brute » qui utilisait la force pure plutôt que la vitesse ou la surprise pour attaquer ses victimes.
Thomas R. Holtz, Jr.
Thomas R. Holtz, Jr.

[![](geobanner.gif)](http://www.geol.umd.edu)[![](https://geol.umd.edu/~tholtz/webglobe.gif)](http://www.umd.edu)[![](http://www.geol.umd.edu/sgc/sharedimages/sunspot.gif)](http://www.scholars.umd.edu)[![](http://static.squarespace.com/static/516d98fde4b0f84b63148a3b/t/52729fe1e4b0a406b9069f03/1383243745844/orientalwhiteeye200.jpg?format=300w)](http://www.bisi.umd.edu/bees)

![Image](https://www.geol.umd.edu/~tholtz/G104/banners/MeandMeg.jpg)

## Thomas R. Holtz Jr.

### Vertebrate Paleontologist   Principal Lecturer

_Megalosaurus bucklandii_, the first non-bird dinosaur recognized by Science. July 2016.

_"There is nothing like geology; the pleasure of the first day's partridge shooting or first day's hunting cannot be compared to finding a fine group of fossil bones, which tell their story of former times with almost a living tongue."_ Charles Darwin, letter to his sister Catherine, 1834

* * *

![](MeandClaw.jpg)

Pedal (foot) claw of the latest Cretaceous oviraptorosaur [_Anzu_](http://static1.squarespace.com/static/51bf1cd3e4b0a897bf54112b/51bf3928e4b09edc5f83d0af/51bf5cf0e4b0cad7682bf91f/1395369704433/Anzu.jpg?format=1000w).
July 2013, near Ekalaka, MT

* * *

[Department of Geology](http://www.geol.umd.edu)

8000 Regents Drive

[University of Maryland](http://www.umd.edu)

College Park, MD 20742

Faculty Director, [Science & Global Change Program](http://www.geol.umd.edu/sgc) [College Park Scholars](http://www.scholars.umd.edu)

Faculty Director, former [Earth, Life & Time Program](http://www.geol.umd.edu/~jmerck/eltsite) [College Park Scholars](http://www.scholars.umd.edu)

2019 [Provost's Excellence Award for Professional Track Faculty in Teaching](https://faculty.umd.edu/awards/list_ptk.html)

Adjunct Faculty, [Behavior, Ecology, Evolution, Systematics](http://www.bees.umd.edu/) Program

Research Associate, [Department of Paleobiology](http://paleobiology.si.edu/), [National Museum of Natural History](http://www.mnh.si.edu/), [Smithsonian Institution](http://www.si.edu/)

Member, Scientific Council, [Maryland Science Center](http://www.mdsci.org/)

Email: [tholtz@umd.edu](mailto:tholtz@umd.edu)

Phone: (301) 405-4084 or (301) 405-6965

Fax (Geology): (301) 314-9661

Geology Office: 1225B [Chemistry Building](http://maps.umd.edu/map/index.html?Nav=Hide&MapView=Detailed&NoWelcome=True&LocationType=Building&LocationName=091)

Scholars Office: 1216 [Centreville Hall](http://reslife.umd.edu/halls/cambridge/centreville/)

Mailing Address:

Department of Geology

8000 Regents Drive

University of Maryland

College Park, MD 20742-4211

USA

![Image](https://www.geol.umd.edu/~tholtz/G104/banners/HoltzPratting.jpg)

Training _Jurassic World_ raptors, Chris Pratt-style. June 2015

* * *

[Current CV](Holtz-UMD CV.pdf)

**Research Interests**

- Phylogeny and functional morphology of the theropod dinosaurs, especially Tyrannosauridae

- Ecomorphology of terrestrial predators

- Cursoriality and terrestrial locomotion

- Mesozoic biogeography, especially quantitative approaches

- Functional character complexes and homoplasy

- Phylogenetic taxonomy of the theropod dinosaurs


**Education**

Ph.D., Yale University, [Department of Geology and Geophysics](http://www.yale.edu/geology), 1992

Dissertation: _[An unusual \_
_structure of the metatarsus of Theropoda (Archosauria, Dinosauria, Saurischia) of the \_
_Cretaceous](https://www.researchgate.net/publication/33700376_An_unusual_structure_of_the_metatarsus_of_Theropoda_Archosauria_Dinosauria_Saurischia_of_the_Cretaceous)_

B.A., Johns Hopkins University, [Department of Earth and Planetary Sciences](http://www.jhu.edu/~eps/), 1987

![Image](https://www.geol.umd.edu/~tholtz/G104/banners/MeAndKitteh.jpg)

_Felis catus_, with _Homo sapiens_ for scale. July 2017.

* * *

**Courses Taught at UMCP**

Department of Geology- [GEOL 102 Historical Geology](http://www.geol.umd.edu/~tholtz/G102/)
- [GEOL 104 Dinosaurs: A Natural History](http://www.geol.umd.edu/~tholtz/G104/)
- GEOL 120 Environmental Geology

- [GEOL 204 Dinosaurs, Early Humans, Ancestors & Evolution: The Fossil Record of Vanished Worlds of the Prehistoric Past](http://www.geol.umd.edu/~tholtz/G204/)
- [GEOL 331 Principles of Paleontology](http://www.geol.umd.edu/~tholtz/G331/)
- [GEOL 431 Vertebrate Paleobiology](https://www.geol.umd.edu/~jmerck/geol431/)
Field Programs

- GEOL 288A Field Studies I: [Natural History & Astronomy of Arizona](http://www.astro.umd.edu/~peel/GEOL288A/)
- GEOL 388G Field Studies II: [Natural History of the Galápagos Islands](https://www.geol.umd.edu/sgc/gal11/)
College Park Scholars Colloquia

- [CPSG 100, 101, 200, 230/240/250](http://www.geol.umd.edu/sgc/menus/syllabi.html), Science & Global Change Colloquia and Practicum

- CPSP 118G Earth, Life & Time Freshman Colloquium, [Fall](http://www.geol.umd.edu/~jmerck/eltsite/colloquium.html#sem1)
and  [Spring](http://www.geol.umd.edu/~jmerck/eltsite/colloquium.html#sem2)
- [CPSP 218G Earth, Life & Time Sophomore Colloquium](http://www.geol.umd.edu/~jmerck/eltsite/colloquium.html#sem3)
Honors Seminars

- [HONR 259C "Fearfully Great Lizards": Topics in Dinosaur Research](http://www.geol.umd.edu/~tholtz/H259C/)
Graduate Courses

- GEOL 789Q Recent Advances in Geology: Paleobiology of the Archosauria

- [GEOL 789W Recent Advances in Geology: Phanerozoic History, Macroevolution, and Earth Systems Science](http://www.geol.umd.edu/~tholtz/G789W/)

**Societies**

- Former President (2006-2009), Paleontological Society of Washington

- [Society of Vertebrate Paleontology](http://www.vertpaleo.org/)
- [The Paleontological Society](http://www.paleosoc.org)
- [The Palaeontological Association](http://www.palass.org/)
- [International Society of Vertebrate Morphologists](http://www.isvm.info/)
- [National Center for Science Education](http://www.natcenscied.org)

[ORCID: 0000-0002-2906-4900](https://orcid.org/0000-0002-2906-4900)

**Recent Publications**

Caspar, K.R., C. Gutiérrez-Ibáñez, O.C. Bertrand, T. Carr, J.A.D. Colbourne, A. Erb, H. George, T.R. Holtz, Jr., D. Naish, D.R. Wylie & G.R. Hulburt. 2024. [How smart was _T. rex_? Testing claims of exceptional cognition in dinosaurs and the application of neuron count estimates in palaeontological research.](https://anatomypubs.onlinelibrary.wiley.com/doi/10.1002/ar.25459)_The Anatomical Record_ in press. doi: 10.1002/ar.25459

Surring, L.A., M.E. Burns, E. Snively, D.E. Barta, T.R. Holtz, Jr., A.P. Russell, L.M. Witmer & P.J. Currie. 2022.  [Consilient evidence affirms expansive stabilizing ligaments in the tyrannosaurid foot.](https://journals.library.ualberta.ca/vamp/index.php/VAMP/article/view/29387) _Vertebrate Anatomy Morphology Palaeontology_ **10:** 49-64. doi: 10.18435/vamp29387

Carr, T.D., J.G. Napoli, S.L. Brusatte, T.R. Holtz, Jr., D.W.E. Hone, T.E. Williamson & L.E. Zanno. 2022. [Insufficient evidence for multiple species of _Tyrannosaurus_ in the latest Cretaceous of North America: a comment on "The Tyrant Lizard King, Queen and Emperor: multiple lines of morphological and stratigraphic evidence support subtle evolution and probable speciation within the North American genus _Tyrannosaurus_](https://link.springer.com/article/10.1007/s11692-022-09573-1?s=09) _Evolutionary Biology_ **(2022)**. doi: 10.1007/s11692-022-09573-1

Hone, D.W.E. & T.R. Holtz, Jr. 2022. [Comment on: Aquatic adaptation in the skull of carnivorous dinosaurs (Theropoda: Spinosauridae) and the evolution of aquatic habits in spinosaurids. 93: 275-284.](https://www.sciencedirect.com/science/article/pii/S0195667119300485) _Cretaceous Research_ **134**: 104152\. doi: 10.1016/j.cretres.2019.05.010

Holtz, T.R., Jr. 2021. [Theropod guild structure and the tyrannosaurid niche assimilation hypothesis: implications for predatory dinosaur macroecology and ontogeny in later Late Cretaceous Asiamerica.](https://cdnsciencepub.com/doi/full/10.1139/cjes-2020-0174) _Canadian Journal of Earth Sciences_ **58**: 778-795. doi: 10.1139/cjes-2020-0174 \[a non-formatted Open Access version of the final paper and its supplementary data is available [here](https://tspace.library.utoronto.ca/handle/1807/106039)\]

Hone, D.W.E., and T.R. Holtz, Jr. 2021. [Evaluating the ecology of _Spinosaurus_: shoreline generalist of aquatic pursuit specialist?](https://palaeo-electronica.org/content/2021/3219-the-ecology-of-spinosaurus) _Palaeontologia Electronica_ **24**(1): a03. doi: 10.26879/1110

Dececchi, T.A., A.M. Mloszewska, T.R. Holtz, Jr., M.B. Habib, and H.C.E. Larsson. 2020. [The fast and the frugal: divergent locomotor strategies drive limb lengthening in theropod dinosaurs.](https://journals.plos.org/plosone/article?id=10.1371/journal.pone.0223698) _PLoS ONE_ **15**(5): e0223698. doi: 10.1371/journal.pone.0223698

Naish, D., A. Cau, T.R. Holtz, Jr., M. Fabbri, and J.A. Gauthier. 2020. Theropoda O.C. Marsh 1881 \[D. Naish, A. Cau, T.R. Holtz, Jr., M. Fabbri, and J.A. Gauthier\], converted clade name. Pp. 1235-1246. In K. de Queiroz, P.D. Cantino, and J.A. Gauthier (eds.), _Phylonyms: A Companion to the PhyloCode._ CRC Press.

Snively, E., H. O'Brien, D.M. Henderson, H. Mallison, L. Surring, M.E. Burns, T.R. Holtz, Jr., A.P. Russell, L.M. Witmer, P.J. Currie, S.A. Hartman & J.R. Cotton. 2019. [Lower rotational inertia and larger leg muscles indicate more rapid turns in tyrannosaurids than in other large theropods.](https://peerj.com/articles/6432/) _PeerJ_ **7**: e6432. doi: 10.7717/peerj.6432

Holtz, T.R., Jr. 2018. [Evolution: New branches on the alvarezsaur tree](https://www.cell.com/current-biology/fulltext/S0960-9822(18)30913-8#.W5aY050XKSA.facebook). _Current Biology_ **28**: R941-R943.

Holtz, T.R., Jr. 2017. [Nomenclature: Share names for dinosaur divisions.](https://www.nature.com/articles/545030d) _Nature_ **545**: 30.

Hone, D.W.E. & T.R. Holtz, Jr. 2017. [A century of spinosaurs - a review and revision of the Spinosauridae with comments on their ecology.](https://onlinelibrary.wiley.com/doi/full/10.1111/1755-6724.13328) _Acta Geologica Sinica_ **91**: 1120-1132. doi: 10.1111/1755-6724.13328

Brusatte, S.L., T.D. Carr, T.E. Williamson, T.R. Holtz, Jr., D.W.E. Hone & S. A. Williams. 2016. [Dentary groove morphology does not distinguish ' _Nanotyrannus_' as a valid taxon of tyrannosauroid dinosaur. Comment on: \
"Distribution of the dentary groove of theropod dinosaurs: Implications for theropod phylogeny and the validity of the genus \
_Nanotyrannus_ Bakker et al., 1988".](http://www.sciencedirect.com/science/article/pii/S0195667116300192) _Cretaceous Research_ **65**: 232-237. doi: 10.1016/j.cretres.2016.02.007

Plotnick, R.E., J.M. Theodor & T.R. Holtz, Jr. 2015. [Jurassic Pork: What could a Jewish Time Traveler eat?](http://link.springer.com/article/10.1186/s12052-015-0047-2) _Evolution: Education and Outreach_ **8**:17\. doi: 10.1186/s12052-015-0047-2

Holtz, T.R., Jr. 2014. [Palaeontology: Mystery of the horrible hands solved.](http://www.nature.com/nature/journal/v515/n7526/abs/nature13930.html) _Nature_ **515**: 203-205. doi:10.1038/nature13930

Farlow, J.O., T.R. Holtz, Jr., T.H. Worthy & R.E. Chapman. 2013. Feet of the fierce (and not so fierce): pedal proportions in large theropods, other non-avian dinosaurs, and large ground birds. Pp. 88-132, in J.M. Parrish, R.E. Molar, P.J. Currie, and E.B. Koppelhus (eds.) _Tyrannosaurid Paleobiology._ Indiana University Press.

Brett-Surman, M.K., T.R. Holtz, Jr. & J.O. Farlow (eds.) 2012. _The Complete Dinosaur._ 2nd Edition. Indiana University Press. 1112 pp.

Holtz, T.R., Jr. 2012. Dinosaur hunters of the Southern Continents. Pp. 107-118, in M.K. Brett-Surman, T.R. Holtz, Jr. & J.O. Farlow (eds.) 2012. _The Complete Dinosaur._ 2nd Edition. Indiana University Press.

Holtz, T.R., Jr. & M.K. Brett-Surman. 2012. The osteology of dinosaurs. Pp. 135-149, in M.K. Brett-Surman, T.R. Holtz, Jr. & J.O. Farlow (eds.) 2012. _The Complete Dinosaur._ 2nd Edition. Indiana University Press.

Holtz, T.R., Jr. & M.K. Brett-Surman. 2012. The taxonomy and systematics of the dinosaurs. Pp. 209-223, in M.K. Brett-Surman, T.R. Holtz, Jr. & J.O. Farlow (eds.) 2012. _The Complete Dinosaur._ 2nd Edition. Indiana University Press.

Holtz, T.R., Jr. 2012. Theropods. Pp. 346-378, in M.K. Brett-Surman, T.R. Holtz, Jr. & J.O. Farlow (eds.) 2012. _The Complete Dinosaur._ 2nd Edition. Indiana University Press.

Holtz, T.R., Jr. 2008. [A critical re-appraisal of the obligate scavenging hypothesis for _Tyrannosaurus rex_ and other tyrant dinosaurs.](https://www.google.com/url?sa=t&rct=j&q=&esrc=s&source=web&cd=&cad=rja&uact=8&ved=2ahUKEwifw93ztrzuAhU2ElkFHbuMCmMQFjAAegQIAhAC&url=https%3A%2F%2Fwww.researchgate.net%2Fpublication%2F292603329_A_critical_reappraisal_of_the_obligate_scavenging_hypothesis_for_Tyrannosaurus_rex_and_other_tyrant_dinosaurs&usg=AOvVaw2sVh-Vs5f-fcEXk12N3Rvg) Pp. 370-396, in P. Larson and K. Carpenter (eds.), Tyrannosaurus rex _: The Tyrant King._ Indiana University Press.

France, C.A.M., P.M. Zelanko, A.J. Kaufman & T.R. Holtz. 2007. [Carbon and nitrogen isotopic analysis of Pleistocene mammals from the Saltville Quarry (Virginia, USA): Implications for trophic relationships](http://dx.doi.org/10.1016/j.palaeo.2007.02.002). _Palaeogeography, Palaeoclimatology, Palaeoecology_ **249**: 271-282.

Holtz, T.R., Jr. 2006. [Palaeontology: A Jurassic tyrant is crowned](http://www.nature.com/nature/journal/v439/n7077/pdf/439665a.pdf). _Nature_ **439**: 665-666.

Holtz, T.R., Jr. & H. Osmólska. 2004. Saurischia. Pp. 21-24, in D.B. Weishampel, P. Dodson and H. Osmólska (eds.), _The Dinosauria. Second Edition._ University of California Press.

Holtz, T.R., Jr., R.E. Molnar & P.J. Currie. 2004. Basal Tetanurae. Pp. 71-110, in D.B. Weishampel, P. Dodson and H. Osmólska (eds.), _The Dinosauria. Second Edition._ University of California Press.

Holtz, T.R., Jr. 2004. Tyrannosauroidea. Pp. 111-136, in D.B. Weishampel, P. Dodson and H. Osmólska (eds.), _The Dinosauria. Second Edition._ University of California Press.

Holtz, T.R., Jr., R.E. Chapman & M.C. Lamanna. 2004. Mesozoic biogeography of Dinosauria, Pp. 627-642 in D.B. Weishampel, P. Dodson and H. Osmólska (eds.), _The Dinosauria. Second Edition._ University of California Press.

Holtz, T.R., Jr. 2003. Dinosaur predation: evidence and ecomorphology. Pp. 325-340, in P.H. Kelley, M. Kowalewski and T.A Hansen (eds.), _Predator-Prey Interactions in the Fossil Record_, Topics in Geobiology Vol. 20. Kluwer Press.

Farlow, J.O. & T.R. Holtz, Jr. 2002. [The fossil record of predation in dinosaurs](http://www.academia.edu/293256/The_Fossil_Record_of_Predation_in_Dinosaurs). Pp. 251-266, in M. Kowalewski & P.H. Kelley (eds.), _The Fossil Record of Predation_. The Paleontological Society Papers 8.

Holtz, T.R., Jr. 2002. [Chasing _Tyrannosaurus_ and _Deinonychus_ around the Tree of Life: classifying dinosaurs](tyrdeintree.pdf). Pp. 31-38, in J.G. Scotchmoor, D.A. Springer, B.H. Breithaupt & A.F. Fiorillo (eds.), _Dinosaurs: The Science Behind the Stories_, Society of Vertebrate Paleontology, Paleontological Society & American Geological Institute.

Holtz, T.R., Jr. 2001. Arctometatarsalia revisited: the problem of homplasy in reconstructing theropod phylogeny. Pp. 99-122, in J. Gauthier and L.F. Gall (eds. ), _New Perspectives on the Origin and Early Evolution of Birds: Proceedings of the International Symposium in Honor of John H. Ostrom._ Yale Univ. Press.

Holtz, T.R., Jr. 2001. The phylogeny and taxonomy of the Tyrannosauridae. Pp. 64-83, in D.H. Tanke and K. Carpenter (eds.), _Mesozoic Vertebrate Life: New Research Inspired by the Paleontology of Philip J. Currie_. Indiana Univ. Press.

Farlow, J.O., S. M. Gatesy, T.R. Holtz, Jr., J.R. Hutchinson & J.M. Robinson. 2000. [Theropod locomotion.](http://www.bioone.org/bioone/?request=get-document&issn=0003-1569&volume=040&issue=04&page=0640) _American Zoologist_ **40**:640-663.

Holtz, T.R., Jr. 2000. Classification and evolution of dinosaur groups. Pp. 140-168, in G.S. Paul (ed.), _The Scientific American Book of Dinosaurs_. St. Martin's Press.

Holtz, T.R., Jr. 2000. [Theropod paleobiology: more than just bird origins.](gaiaintro.pdf) _Gaia_ **15**: 1-3.

Holtz, T.R., Jr. 2000. [A new phylogeny of the carnivorous dinosaurs.](gaiaphyl.pdf) _Gaia_ **15**: 5-61.

Holtz, T.R., Jr., D.L. Brinkman & C.L. Chandler. 2000. [Denticle morphometrics and a possibly omnivorous feeding habit for the theropod dinosaur _Troodon_.](gaiatroo.pdf) _Gaia_ **15**: 159-166.

Padian, K., J.R. Hutchinson & T.R. Holtz, Jr. 1999. Phylogenetic definitions and nomenclature of the major taxonomic categories of the carnivorous Dinosauria (Theropoda). _Journal of Vertebrate Paleontology_ **19**: 69-80.

Holtz, T.R., Jr. 1998. [Spinosaurs as crocodile mimics](http://www.sciencemag.org/cgi/content/full/282/5392/1276). _Science_ **282**: 1276-1277.

Holtz, T.R., Jr. 1997. Dinosaur hunters of the Southern Continents. Pp. 43-51, in J.O. Farlow & M.K. Brett-Surman (eds.), _The Complete Dinosaur_. Indiana Univ. Press.

Holtz, T.R., Jr. & M.K. Brett-Surman. 1997. The osteology of the dinosaurs. Pp. 78-91, in J.O. Farlow & M.K. Brett-Surman (eds.), _The Complete Dinosaur_. Indiana Univ. Press.

Holtz, T.R., Jr. & M.K. Brett-Surman. 1997. The taxonomy and systematics of the dinosaurs. Pp. 92-106, in J.O. Farlow & M.K. Brett-Surman (eds.), _The Complete Dinosaur_. Indiana Univ. Press.

Holtz, T.R., Jr. 1996. [Phylogenetic taxonomy of the Coelurosauria (Dinosauria: Theropoda).](http://www.jstor.org/stable/1306452) _Journal of Paleontology_ **70**: 536-538.

Holtz, T.R., Jr. 1995. [The arctometatarsalian pes, an unusual structure of the metatarsus of Cretaceous Theropoda (Dinosauria: Saurischia).](http://www.jstor.org/stable/4523590) _Journal of Vertebrate Paleontology_ **14**: 480-519.

Holtz, T.R., Jr. 1994. [The phylogenetic position of the Tyrannosauridae: implications for theropod systematics.](http://www.jstor.org/stable/1306180) _Journal of Paleontology_ **68**: 1100-1117.

**Web-based Publications**

[Tyrannosauroidea](http://tolweb.org/tree?group=Tyrannosauroidea&contgroup=Coelurosauria&dynnodeid=12799), [Tyrannosauridae](http://tolweb.org/tree?group=Tyrannosauridae&contgroup=Tyrannosauroidea&dynnodeid=12975),
and [Tyrannosaurid Systematics](http://tolweb.org/tree/eukaryotes/animals/chordata/dinosauria/tyrannosauroidea/tyransystem.html) pages
for the [Tree of Life](http://tolweb.org/tree/phylogeny.html) project

**Phylogenetic Databases**

- **Theropoda (with emphasis on non-avian Tetanurae)** (from _Dinosauria. 2nd Edition_ chapters)

  - [Character descriptions](http://www.ucpress.edu/books/pages/2601001/Dino4.1BasalTetaDesc.doc) (MS Word format)

  - [Data Matrix](http://www.ucpress.edu/books/pages/2601001/Dino4.2BasalTetaMatrix.doc) (MS Word format)

  - [All data matricies](http://www.ucpress.edu/books/pages/2601001/2601001.supplement.html) from _Dinosauria. 2nd Edition._
- **Theropoda** (from _Gaia_ and _New Perspectives on the Origin and Early Evolution of Birds_ papers)

  - [Nexus File](gaia.mc)
  - [Updated](Gaia2000) version of above, with some corrections for _Ceratosaurus_
  - [Character description](gaiamatrix.rtf)
- **Tyrannosauridae** (from the _Mesozoic Vertebrate Life_ chapter and [Tyrannosauridae](http://tolweb.org/tree?group=Tyrannosauridae&contgroup=Tyrannosauroidea&dynnodeid=12975)
  page of the [Tree of Life](http://tolweb.org/tree/phylogeny.html) project)

  - [Nexus File](tyrants.mc)
  - [Character description](tyrmatrix.rtf)

![](http://www.geol.umd.edu/~tholtz/dinoappendix/Holtzcover.jpg)

Holtz, T.R., Jr. (illustrations by L. Rey). 2007. _Dinosaurs: The Most Complete Up-To-Date Encyclopedia_
_for Dinosaur Lovers of All Ages_. Random House, 428 pp. [Supplementary information](http://www.geol.umd.edu/~tholtz/dinoappendix/)
(including chapter updates) available at [http://www.geol.umd.edu/~tholtz/dinoappendix/](http://www.geol.umd.edu/~tholtz/dinoappendix/).

**Some Links**

[Wikipedia](http://en.wikipedia.org/wiki/Thomas_R._Holtz_Jr.) entry, and [EvoWiki](http://evolutionwiki.org/wiki/Thomas_Holtz), too!

["Bizarre Dinosaurs: What's Wrong With This Picture? An Audio Critique"](http://ngm.nationalgeographic.com/2007/12/bizarre-dinosaurs/holtz-dinosaur-photography), audio commentary by me on National Geographic's review of some of their historic dinosaur art

An [interview with me](https://www.robotbutt.com/2015/06/12/an-interview-with-thomas-r-holtz-dinosaur-rock-star/) on [Robot Butt](https://www.robotbutt.com)
**For Fun**

Tom Holtz's science fictional appearances:
- Michael Swanwick's short short story "Proving Dr. Tom's Hypothesis" in _Field Guide to_
  _Mesozoic Megafauna_
- Michael Swanwick's novel _Bones of the Earth_
[Parry and Carney, Friends for Life](http://www.parryandcarney.com/), a dinosaur-based webcomic by former UMd student (and veteran of my first Honors Seminar), Mike Keesey

Charley Parker's [Dinosaur Cartoons](http://www.dinosaurcartoons.com/index.html)

###### Last modified: 26 April 2024

![Image](https://www.geol.umd.edu/~tholtz/G104/banners/HoltzChelon.jpg)

_Chelonoidis_-selfie. June 2011.