	The holotype, MNHN 1961-28, was found in a layer of the Tiouraren Formation dating from the Bathonian-Oxfordian. De Lapparent had presumed that the strata dated from the Early Cretaceous. It consists of a cervical vertebra, seven pieces of the dorsals, three pieces of the sacrum, five tail vertebrae, a humerus, the lower end of a pubic bone, the lower end of a thighbone, a piece of a shinbone, a piece of a fibula, a metatarsal, four additional pieces of the metatarsus and a phalanx of a toe. The paratypes were an ulna, a metatarsal and a second partial skeleton consisting of vertebrae and limb elements. In 2004, Sereno e.a. referred a third skeleton, specimen MNN TIG6 consisting of a series of cervical and dorsal vertebrae together with some ribs.