# Exploring the evolution of a trade-off between vigilance and foraging in group-living organisms 

Randal S. Olson $^{1,5,*}$, Patrick B. Haley$^{2,5,*}$, Fred C. Dyer$^{3,5}$, and Christoph Adami4,5 

$^{1}$Computer Science & Engineering Dept., Michigan State University, East Lansing, MI 48824 

$^{2}$Computer Science Dept., The University of Texas at Austin, Austin, TX 78712 

$^{3}$Zoology Dept., Michigan State University, East Lansing, MI 48824 

$^{4}$Microbiology & Molecular Genetics Dept., Michigan State University, East Lansing, MI 48824 

$^{5}$BEACON Center for the Study of Evolution in Action, East Lansing, MI 48824 

$^{*}$These authors contributed equally to this work 

E-mail: olsonran@msu.edu, patrick.haley@utexas.edu, fcdyer@msu.edu, adami@msu.edu 

Keywords: group foraging , many eyes hypothesis , anti-predator vigilance , genetic relatedness , reproductive strategy , tragedy of the commons 

Article submitted to the Journal of the Royal Society Interface, August 2014. 

