ACT_SITE 84; /evidence="ECO:0000250|UniProtKB:Q96FW1"; ACT_SITE 87; /note="Nucleophile"; /evidence="ECO:0000250|UniProtKB:Q96DC9"; ACT_SITE 244; /evidence="ECO:0000250|UniProtKB:Q96FW1"; ACT_SITE 268; /evidence="ECO:0000250|UniProtKB:Q96FW1"