ACT_SITE 128; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 131; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 467; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 470; /note="Nucleophile"; /evidence="ECO:0000250"