ACT_SITE 23; /note="Nucleophile"; /evidence="ECO:0000255|PROSITE-ProRule:PRU00331"; ACT_SITE 132; /note="Proton acceptor"; /evidence="ECO:0000255|PROSITE-ProRule:PRU00331"; ACT_SITE 147; /evidence="ECO:0000255|PROSITE-ProRule:PRU00331"