ACT_SITE 247; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 326; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 415; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 419; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 452; /evidence="ECO:0000250|UniProtKB:Q38860"