ACT_SITE 132; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 135; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 471; /note="Nucleophile"; /evidence="ECO:0000250"; ACT_SITE 474; /note="Nucleophile"; /evidence="ECO:0000250"