ACT_SITE 225; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 304; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 388; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 392; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 421; /evidence="ECO:0000250|UniProtKB:Q38860"; ACT_SITE 425; /evidence="ECO:0000250|UniProtKB:Q38860"