ACT_SITE 391; /note="Nucleophile; for glutamine hydrolysis"; /evidence="ECO:0000255|HAMAP-Rule:MF_01227"; ACT_SITE 517; /evidence="ECO:0000255|HAMAP-Rule:MF_01227, ECO:0000255|PROSITE-ProRule:PRU00605"; ACT_SITE 519; /evidence="ECO:0000255|HAMAP-Rule:MF_01227, ECO:0000255|PROSITE-ProRule:PRU00605"