ACT_SITE 89; /evidence="ECO:0000250|UniProtKB:Q96DC9"; ACT_SITE 92; /note="Nucleophile"; /evidence="ECO:0000250|UniProtKB:Q96DC9"; ACT_SITE 259; /evidence="ECO:0000250|UniProtKB:Q96FW1"; ACT_SITE 288; /evidence="ECO:0000250|UniProtKB:Q96DC9"