Read the following paragraph and extract the answer for the question: Who is the jungle king's mentor indebted to?

Five years after socialite Ursula Stanhope left civilization to marry George of the Jungle, George finds himself hard-pressed to fulfill the roles of jungle king, father, and husband. George's stress level increases when the "Mean Lion" challenges him for leadership of the jungle, and when Ursula's mother Beatrice teams up with Ursula's ex-fiancé, Lyle, in a plot to forcibly take away all that George holds most dear. 
To do this, Beatrice invites Ursula, George, and George Junior to visit Las Vegas, which they accept. Throughout the visit, Beatrice and some of Ursula's fellow socialites try constantly to convince Ursula that George is unworthy of her affection. George, observing the threats but not his wife's resistance, begins to think himself unworthy of Ursula.
During the same time, George's mentor Ape has become a gambler and is in debt to several creditors, including Lyle. Upon discovering that Ape does not possess the exploitation rights of Ape Mountain, but George does, Lyle instead makes Ape work off his debts from the gambling for the next 17 years. He then engages Ape as a staged song performer and steals the deed to Ape Mountain from George's wardrobe. He thereafter sends two agents, Sally and Kowalski, to Ape Mountain, where they begin to demolish the jungle. The animals, terrified, turn to the Lion for guardianship.
Having failed to convince Ursula to divorce George, Beatrice hires a master of hypnosis to hypnotize Ursula into having no memory of having known George. The hypnotist brainwashes Ursula, giving her the idea that she had married Lyle. George, upon hearing from Beatrice that Ursula has left him, leaves his luck-charm with Ursula as she sleeps, then departs. He rescues Ape and leaves Las Vegas. Their departure triggers much commotion when the police force and the Animal Control Agency join forces to recapture them.
user
Lyle
assistant