What was the growth rate of Google Other revenues in the first quarter of 2021 
or the past four years, and we are working toward operating on carbon-free energy around the clock by 2030, a far more ambitious goal that we hope will be transformative for the industry. Five of our data centers in Europe and North America are already operating near or at 90% carbon-free energy around the clock. We are also working toward our commitment to help one billion people make more sustainable choices with our services by 2022. One example of how we are doing this is the new AI-powered feature in Maps that will show the route with the lowest carbon footprint when ETAs are the same. You'll also be able to compare the carbon impact between routes.
Finally, as we look ahead to the rest of 2021, our four big themes continue to guide us: first, building and providing the most helpful products and services; second, continuing to earn the trust of our users by investing in high-quality information and keeping users' data safe and private; third, strong execution as a company, particularly as we start to reopen our offices; and fourth, building sustainable value in our own business and for our partners.
As always, thank you to our Googlers around the world for a great start to the year. To everyone, I look forward to seeing you at IO. Over to Philipp.
Philipp Schindler -- Chief Business Officer
Thanks, Sundar, and good afternoon, everyone. It's great to be joining you again today.
We're pleased with the strong growth in Google Services revenues in the first quarter. Year-on-year performance reflects elevated consumer online activity, broad-based strength in advertiser spend, and lapping of the initial impact of the pandemic on advertising revenues that began in March last year.
In the first quarter in Search, we saw sustained strength across most categories led by retail. We also saw strong performance in tech and CPG. In YouTube, we had phenomenal growth driven by direct response followed by continued strength in brand. We've seen great momentum in TrueView for Action ads, with a number of advertisers using the format doubling over the past year. In Network, exceptional growth was driven by AdMob and Ad Manager with particular strength in app campaigns. Google Other revenues were driven by growth in Google Play and YouTube's non-advertising revenues, followed by hardware.
I would now like to take a few minutes to dive deeper into the trends we're seeing in our business. As Sundar touched on earlier, the pandemic is evolving in different ways across the world. Some countries are in advanced stages of reopening, others are facing reacceleration of cases. And there is everything in between. It's never been more important to help businesses navigate the pandemic as circumstances change.
On travel, we're starting to see renewed interest from users as they turn to Google to plan their next trip even before they're ready to book. Every travel partners looking to understand where demand is going and we are helping them to find these opportunities through insights and automation. For American Airlines, for example, that meant using our insight tools to anticipate demand on untapped routes. These newly prioritized routes had significantly higher booking rate in Search this quarter compared to last quarter. Also just last month, we made it free for hotels and travel companies to list their bookings links, similar to what we did with shopping last year. For consumers, it means more choice. For hotels and travel companies, it means free exposure on Google. For advertisers, it means paid campaigns can be augmented with free listings. We are already seeing positive results across the board.
Let me switch gears now and talk about retail, where we had a very strong quarter. As you know, we've taken important steps over the past year to accelerate an open retail ecosystem. We made product listings free, removed commission fees and opened our shopping platform to Shopify and PayPal. We're also helping retailers with some key opportunities, such as innovating an omnichannel as the line between digital and physical retail continues to blur
or the past four years, and we are working toward operating on carbon-free energy around the clock by 2030, a far more ambitious goal that we hope will be transformative for the industry. Five of our data centers in Europe and North America are already operating near or at 90% carbon-free energy around the clock. We are also working toward our commitment to help one billion people make more sustainable choices with our services by 2022. One example of how we are doing this is the new AI-powered feature in Maps that will show the route with the lowest carbon footprint when ETAs are the same. You'll also be able to compare the carbon impact between routes.
Finally, as we look ahead to the rest of 2021, our four big themes continue to guide us: first, building and providing the most helpful products and services; second, continuing to earn the trust of our users by investing in high-quality information and keeping users' data safe and private; third, strong execution as a company, particularly as we start to reopen our offices; and fourth, building sustainable value in our own business and for our partners.
As always, thank you to our Googlers around the world for a great start to the year. To everyone, I look forward to seeing you at IO. Over to Philipp.
Philipp Schindler -- Chief Business Officer
Thanks, Sundar, and good afternoon, everyone. It's great to be joining you again today.
We're pleased with the strong growth in Google Services revenues in the first quarter. Year-on-year performance reflects elevated consumer online activity, broad-based strength in advertiser spend, and lapping of the initial impact of the pandemic on advertising revenues that began in March last year.
In the first quarter in Search, we saw sustained strength across most categories led by retail. We also saw strong performance in tech and CPG. In YouTube, we had phenomenal growth driven by direct response followed by continued strength in brand. We've seen great momentum in TrueView for Action ads, with a number of advertisers using the format doubling over the past year. In Network, exceptional growth was driven by AdMob and Ad Manager with particular strength in app campaigns. Google Other revenues were driven by growth in Google Play and YouTube's non-advertising revenues, followed by hardware.
 I would now like to take a few minutes to dive deeper into the trends we're seeing in our business. As Sundar touched on earlier, the pandemic is evolving in different ways across the world. Some countries are in advanced stages of reopening, others are facing reacceleration of cases. And there is everything in between. It's never been more important to help businesses navigate the pandemic as circumstances change.
On travel, we're starting to see renewed interest from users as they turn to Google to plan their next trip even before they're ready to book. Every travel partners looking to understand where demand is going and we are helping them to find these opportunities through insights and automation. For American Airlines, for example, that meant using our insight tools to anticipate demand on untapped routes. These newly prioritized routes had significantly higher booking rate in Search this quarter compared to last quarter. Also just last month, we made it free for hotels and travel companies to list their bookings links, similar to what we did with shopping last year. For consumers, it means more choice. For hotels and travel companies, it means free exposure on Google. For advertisers, it means paid campaigns can be augmented with free listings. We are already seeing positive results across the board.
Let me switch gears now and talk about retail, where we had a very strong quarter. As you know, we've taken important steps over the past year to accelerate an open retail ecosystem. We made product listings free, removed commission fees and opened our shopping platform to Shopify and PayPal. We're also helping retailers with some key opportunities, such as innovating an omnichannel as the line between digital and physical retail continues to blur