2740
In addition , a wild - type strain containing a temperature - sensitive threonyl - tRNA synthetase mutation showed increased thr operon expression at the non - permissive temperature , whereas none of the mutants showed any change .
 GENE: temperature - sensitive threonyl - tRNA synthetase mutation; GENE: thr operon
Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
 Entities: temperature - sensitive threonyl - tRNA synthetase mutation, thr operon 
Answer:
 GENE: temperature - sensitive threonyl - tRNA synthetase mutation; GENE: thr operon