3914
Thus, IL-2 -induced IL-2R alpha promoter activity requires a complex upstream element , which appears to contain binding sites for both positive and negative regulatory factors .
 DNA: IL-2R alpha promoter; DNA: binding sites; protein: IL-2; DNA: complex upstream element
Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
 Entities: IL-2R alpha promoter, binding sites, IL-2, complex upstream element 
Answer:
 DNA: IL-2R alpha promoter; DNA: binding sites; protein: IL-2; DNA: complex upstream element