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However, the TPA responsiveness of both CMV elements proved to involve synergistic interactions between the core SRF binding site ( CCATATATGG ) and the adjacent inverted ETS binding motifs ( TTCC ), which correlated directly with formation of a bound tripartite complex containing both the cellular SRF and ELK-1 proteins .
 protein: ETS; protein: SRF; protein: tripartite complex; DNA: CMV elements; DNA: ETS binding motifs; protein: ELK-1 proteins; protein: SRF; DNA: SRF binding site
Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
 Entities: ETS, SRF, tripartite complex, CMV elements, ETS binding motifs, ELK-1 proteins, SRF, SRF binding site 
Answer:
 protein: ETS; protein: SRF; protein: tripartite complex; DNA: CMV elements; DNA: ETS binding motifs; protein: ELK-1 proteins; protein: SRF; DNA: SRF binding site