5656
The proteins encoded downstream of rpoN are known to negatively regulate sigma 54 activity .
 GENE: rpoN; GENE: sigma 54
Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: GENE 
Text: {0} 
 Entities: rpoN, sigma 54 
Answer:
 GENE: rpoN; GENE: sigma 54