2552
A 500-bp fragment encompassing this hypersensitive site ( nt 4481 to 4982 ) exhibited transcription-enhancing activity (two- to threefold) when it was cloned in its natural position with respect to the HIV-1 promoter after transient transfection in U937 and CEM cells .
 DNA: 500-bp fragment; cell line: U937; cell line: CEM cells; DNA: hypersensitive site; DNA: nt 4481 to 4982; DNA: HIV-1 promoter
Given options, please tell me the categories of all the listed entity words.Output format is "type1: word1; type2: word2". 
Option: cell line, protein, RNA, DNA, cell type 
Text: {0} 
 Entities: 500-bp fragment, U937, CEM cells, hypersensitive site, nt 4481 to 4982, HIV-1 promoter 
Answer:
 DNA: 500-bp fragment; cell line: U937; cell line: CEM cells; DNA: hypersensitive site; DNA: nt 4481 to 4982; DNA: HIV-1 promoter