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Leslie Barnett ( born 12 October 1920 in London United Kingdom as Margaret Leslie Collard died 10 February 2002 ) was a  British  biologist who worked with  Francis Crick  Sydney Brenner and Richard J. Watts-Tobin to genetically demonstrate the triplet nature of the code of protein translation through the Crick Brenner Barnett Watts-Tobin et al. experiment of 1961 which discovered frameshift mutations ; this insight provided early elucidation of the nature of the genetic code .
 origin: British, Francis Crick
Given options, please tell me the relationships of all the listed entity pairs.Output format is "relation1: word1, word2; relation2: word3, word4". 
Option: cities of residence, top members employees, members, alternate names, origin, state or provinces of residence, title of person, spouse, city of headquarters, NA, founded by, state or province of headquarters, employee of, countries of residence, country of birth, founded, subsidiaries, country of headquarters 
Text: {0} 
 Entity Pairs:  British, Francis Crick 
Answer:
 origin: British, Francis Crick