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The major putative downstream site for E2F , located in the first exon , serves as a target for E2F-1/DP1 complex binding in vitro.
 ( DNA: major putative downstream site, None) , ( protein: E2F, None) , ( protein: E2F-1/DP1 complex, None) , ( DNA: first exon, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: major putative downstream site, None) , ( protein: E2F, None) , ( protein: E2F-1/DP1 complex, None) , ( DNA: first exon, None) 