13216
However, this region directed extremely low reporter mRNA levels relative to endogenous IL-4 mRNA , suggesting that full gene activity requires additional enhancer elements .
 ( RNA: IL-4 mRNA, None) , ( DNA: enhancer elements, None) , ( protein: IL-4, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( RNA: IL-4 mRNA, None) , ( DNA: enhancer elements, None) , ( protein: IL-4, None) 