939
By contrast, three lymphoblastoid cell lines (immortalized in vitro with Epstein-Barr virus ) were weakly positive and MNDA was up-regulated by interferon-alpha ( IFN-alpha ) treatment .
 ( protein: MNDA, None) , ( cell line: lymphoblastoid cell lines, None) , ( protein: IFN-alpha, None) , ( protein: interferon-alpha, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: MNDA, None) , ( cell line: lymphoblastoid cell lines, None) , ( protein: IFN-alpha, None) , ( protein: interferon-alpha, None) 