9968
Subnuclear fractionation reveals that there are two ATF1 isoforms which appear to differ with respect to DNA binding activity and partition selectively between nuclear matrix and nonmatrix compartments , consistent with the role of the nuclear matrix in regulating gene expression .
 ( protein: ATF1 isoforms, None) , ( protein: ATF1, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: ATF1 isoforms, None) , ( protein: ATF1, None) 