9969
Site-directed mutational studies demonstrate that Site I and Site IV together support ATF1- and CREB- induced trans-activation of the H4 promoter .
 ( DNA: Site I, None) , ( DNA: Site IV, None) , ( DNA: H4 promoter, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: Site I, None) , ( DNA: Site IV, None) , ( DNA: H4 promoter, None) 