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Protein-DNA binding studies suggested that Sp1 and two other CCGCCC binding proteins from erythroid and non-erythroid cells could bind to the Sp1 binding motif .
 ( DNA: Sp1 binding motif, None) , ( protein: CCGCCC binding proteins, None) , ( protein: Sp1, None) , ( protein: Sp1, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: Sp1 binding motif, None) , ( protein: CCGCCC binding proteins, None) , ( protein: Sp1, None) , ( protein: Sp1, None) 