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The cell cycle-dependent transcription factor , E2F-1 , regulates the cyclin -like species of the DNA repair enzyme uracil-DNA glycosylase ( UDG ) gene in human osteosarcoma ( Saos-2 ) cells .
 ( DNA: uracil-DNA glycosylase ( UDG ) gene, None) , ( cell line: Saos-2, None) , ( protein: DNA repair enzyme, None) , ( protein: E2F-1, None) , ( protein: uracil-DNA glycosylase, None) , ( protein: UDG, None) , ( cell line: human osteosarcoma ( Saos-2 ) cells, None) , ( protein: cyclin, None) , ( protein: cell cycle-dependent transcription factor, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: uracil-DNA glycosylase ( UDG ) gene, None) , ( cell line: Saos-2, None) , ( protein: DNA repair enzyme, None) , ( protein: E2F-1, None) , ( protein: uracil-DNA glycosylase, None) , ( protein: UDG, None) , ( cell line: human osteosarcoma ( Saos-2 ) cells, None) , ( protein: cyclin, None) , ( protein: cell cycle-dependent transcription factor, None) 