1311
Since the human GM-CSF promoter has a base difference within the GC-box , we speculate that this may explain why the human promoter is weak and that an upstream enhancer is required for the induction of the human GM-CSF gene .
 ( DNA: GC-box, None) , ( DNA: upstream enhancer, None) , ( DNA: human GM-CSF gene, None) , ( DNA: human GM-CSF promoter, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: GC-box, None) , ( DNA: upstream enhancer, None) , ( DNA: human GM-CSF gene, None) , ( DNA: human GM-CSF promoter, None) 