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We investigated the role of this G + C-rich region in augmenting alpha-globin promoter activity in the presence of the far-upstream alpha-globin enhancer , HS-40 .
 ( DNA: HS-40, None) , ( DNA: G + C-rich region, None) , ( DNA: alpha-globin promoter, None) , ( DNA: far-upstream alpha-globin enhancer, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: HS-40, None) , ( DNA: G + C-rich region, None) , ( DNA: alpha-globin promoter, None) , ( DNA: far-upstream alpha-globin enhancer, None) 