14437
When ligated to the murine c-fos promoter , however, the proIL-1 beta enhancer was inducible in phorbol myristate acetate -stimulated HeLa cells , suggesting the existence of a proIL-1 beta promoter-proximal requirement for tissue specificity .
 ( DNA: proIL-1 beta enhancer, None) , ( DNA: c-fos, None) , ( protein: proIL-1 beta, None) , ( DNA: murine c-fos promoter, None) , ( protein: proIL-1 beta, None) , ( cell line: phorbol myristate acetate -stimulated HeLa cells, None) , ( cell type: HeLa cells, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: proIL-1 beta enhancer, None) , ( DNA: c-fos, None) , ( protein: proIL-1 beta, None) , ( DNA: murine c-fos promoter, None) , ( protein: proIL-1 beta, None) , ( cell line: phorbol myristate acetate -stimulated HeLa cells, None) , ( cell type: HeLa cells, None) 