5086
Previous studies have shown that retinoic acid ( RA ), similar to tumor necrosis factor-alpha ( TNF-alpha ), can act as a bifunctional regulator of the growth of bone marrow progenitors , in that it can stimulate granulocyte-macrophage colony-stimulating factor (GM-CSF)- or interleukin-3 (IL-3)- induced GM colony formation , but potently inhibit G-CSF -induced growth .
 ( cell type: bone marrow progenitors, None) , ( protein: tumor necrosis factor-alpha, None) , ( protein: TNF-alpha, None) , ( protein: G-CSF, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( cell type: bone marrow progenitors, None) , ( protein: tumor necrosis factor-alpha, None) , ( protein: TNF-alpha, None) , ( protein: G-CSF, None) 