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However, the TPA responsiveness of both CMV elements proved to involve synergistic interactions between the core SRF binding site ( CCATATATGG ) and the adjacent inverted ETS binding motifs ( TTCC ), which correlated directly with formation of a bound tripartite complex containing both the cellular SRF and ELK-1 proteins .
 ( protein: ETS, None) , ( protein: SRF, None) , ( protein: tripartite complex, None) , ( DNA: CMV elements, None) , ( DNA: ETS binding motifs, None) , ( protein: ELK-1 proteins, None) , ( protein: SRF, None) , ( DNA: SRF binding site, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: ETS, None) , ( protein: SRF, None) , ( protein: tripartite complex, None) , ( DNA: CMV elements, None) , ( DNA: ETS binding motifs, None) , ( protein: ELK-1 proteins, None) , ( protein: SRF, None) , ( DNA: SRF binding site, None) 