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We demonstrate, through the deletion of the human UDG promoter sequences , that expression of E2F-1 activates the UDG promoter through several E2F sites .
 ( DNA: E2F sites, None) , ( protein: UDG, None) , ( DNA: UDG promoter, None) , ( protein: E2F-1, None) , ( protein: UDG, None) , ( DNA: human UDG promoter sequences, None) , ( protein: E2F, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: E2F sites, None) , ( protein: UDG, None) , ( DNA: UDG promoter, None) , ( protein: E2F-1, None) , ( protein: UDG, None) , ( DNA: human UDG promoter sequences, None) , ( protein: E2F, None) 