2835
Treatment of Jurkat cells with mAb J393 induced tyrosine phosphorylation of specific protein substrates that underwent hyperphosphorylation upon antigen receptor costimulation .
 ( cell line: Jurkat cells, None) , ( protein: protein substrates, None) , ( protein: mAb J393, None) , ( protein: antigen receptor, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( cell line: Jurkat cells, None) , ( protein: protein substrates, None) , ( protein: mAb J393, None) , ( protein: antigen receptor, None) 