10535
When macrophages were pretreated with low doses of IFN-gamma and then stimulated with IFN-alpha , clearly enhanced formation of specific transcription factor complexes was detected.
 ( cell type: macrophages, None) , ( protein: IFN-gamma, None) , ( protein: transcription factor, None) , ( protein: IFN-alpha, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( cell type: macrophages, None) , ( protein: IFN-gamma, None) , ( protein: transcription factor, None) , ( protein: IFN-alpha, None) 