14695
Site-directed mutagenesis further demonstrated that the enhancer function of one of the two NF-E2/AP1 motifs of HS-40 is mediated through its binding to NF-E2 but not AP1 transcription factor .
 ( DNA: enhancer, None) , ( protein: AP1 transcription factor, None) , ( DNA: HS-40, None) , ( protein: NF-E2, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: enhancer, None) , ( protein: AP1 transcription factor, None) , ( DNA: HS-40, None) , ( protein: NF-E2, None) 