12480
In situ immunolabelling revealed that upregulation of protein expression by IFN-alpha is a consequence of a marked increase in both the number and the intensity of the staining of so-called PML nuclear bodies .
 ( protein: PML nuclear bodies, None) , ( DNA: PML, None) , ( protein: IFN-alpha, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: PML nuclear bodies, None) , ( DNA: PML, None) , ( protein: IFN-alpha, None) 