7452
It is perhaps possible that all of Stat4 actions on Th1 development may be exert directly by Stat4 at the IFN-gamma gene , however we suggest that, more likely, Stat4 may act to induce Th1 development through the induction of other non-cytokine genes , whose stable expression maintains the transcriptional state of a Th1 cell .
 ( protein: IFN-gamma, None) , ( protein: Stat4, None) , ( DNA: IFN-gamma gene, None) , ( protein: Stat4, None) , ( cell type: Th1 cell, None) , ( DNA: non-cytokine genes, None) , ( protein: Stat4, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: IFN-gamma, None) , ( protein: Stat4, None) , ( DNA: IFN-gamma gene, None) , ( protein: Stat4, None) , ( cell type: Th1 cell, None) , ( DNA: non-cytokine genes, None) , ( protein: Stat4, None) 