3590
By competition analysis with transcription factor consensus sequence oligonucleotides and by immunosupershift , transcription factor SP-1 or a closely related protein was shown to bind to this regulatory element .
 ( DNA: transcription factor consensus sequence oligonucleotides, None) , ( protein: transcription factor SP-1, None) , ( DNA: regulatory element, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: transcription factor consensus sequence oligonucleotides, None) , ( protein: transcription factor SP-1, None) , ( DNA: regulatory element, None) 