956
However, analysis of sucrose gradient fractions in the gel retardation assay provided evidence that the LMP promoter -binding proteins form a complex of higher M(r) in EBNA-2 -positive cell extracts .
 ( DNA: LMP promoter, None) , ( cell type: EBNA-2 -positive cell, None) , ( protein: LMP promoter -binding proteins, None) , ( protein: EBNA-2, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: LMP promoter, None) , ( cell type: EBNA-2 -positive cell, None) , ( protein: LMP promoter -binding proteins, None) , ( protein: EBNA-2, None) 