5480
Neutralizing mAbs to IL-10 and TNF-alpha indicated that both cytokines were essential for the induction and were required to generate a synergistic increase in virus expression .
 ( protein: mAbs, None) , ( protein: IL-10, None) , ( protein: TNF-alpha, None) , ( protein: cytokines, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: mAbs, None) , ( protein: IL-10, None) , ( protein: TNF-alpha, None) , ( protein: cytokines, None) 