12208
Mutational analysis of UAS1 sequences combined with in vitro bandshift assays revealed the presence of three binding sites for cellular factors in this region.
 ( DNA: binding sites, None) , ( DNA: UAS1 sequences, None) , ( DNA: UAS1, None) , ( protein: cellular factors, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: binding sites, None) , ( DNA: UAS1 sequences, None) , ( DNA: UAS1, None) , ( protein: cellular factors, None) 