5391
Our results suggest that the bulk of protein-DNA interactions that underlies the developmental control of globin genes takes place in the gamma- and beta- globin gene promoters , and that GT motifs of the beta-globin locus LCR may play a role in the developmental regulation of human beta-globin gene expression , perhaps by increasing the probability of interaction of the LCR holocomplex with the fetal or the adult globin gene .
 ( DNA: LCR, None) , ( DNA: beta-globin locus, None) , ( protein: LCR holocomplex, None) , ( DNA: LCR, None) , ( DNA: GT motifs, None) , ( DNA: beta-globin gene, None) , ( DNA: globin genes, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( DNA: LCR, None) , ( DNA: beta-globin locus, None) , ( protein: LCR holocomplex, None) , ( DNA: LCR, None) , ( DNA: GT motifs, None) , ( DNA: beta-globin gene, None) , ( DNA: globin genes, None) 