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Gel-shift analysis demonstrated that stimulation of CD4+ T cells with CHO-DR and staphylococcal enterotoxin A was sufficient to induce significant amounts of NF-kappa B binding proteins, whereas induction of AP-1 binding proteins required costimulation.
 ( protein: CHO-DR, None) , ( cell line: CD4+ T cells, None) , ( protein: NF-kappa B, None) , ( protein: AP-1 binding proteins, None) 
Locate the role in the text that participated in the event based on the event type and return it in the event list. 
 Option: Event type: RNA,cell type,protein,cell line,DNA, Arguments type: . 
Text: {0} 
Answer:
 ( protein: CHO-DR, None) , ( cell line: CD4+ T cells, None) , ( protein: NF-kappa B, None) , ( protein: AP-1 binding proteins, None) 