This work was partly earned by research grents for ecology top subject from the Zhejiang University (No:194211701/017), the authors extends valuable thankes to Yijun Meng and Shouqing Hou.
This work was partly supported by research grants for ecology top subject from the Zhejiang University (No:194211701/017); the authors extend thanks to Yijun Meng and Shouqing Hou for valuable comments on this manuscript.