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Dry pollen grains often show preformed clefts that allow extreme volume changes accompanying desiccation and hydration. Some of them additionally serve as portals for pollen tube exit during germination and are called apertures. They are highly variable in size, number and complexity. Pollen grains from monocotyle plant species, as for example those from tropical epiphytes, usually possess only a single meridional cleft, which spans along the entire length.