text:
A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Inthanon, Chiang Mai Province, northern Thailand
Zootaxa 3905(4):573-584
DOI:10.11646/zootaxa.3905.4.9
Kirati Kunya
Montri Sumontha
Ranong Marine Fisheries Research and Development Station
Nonn Panitvong
Societas Medicinae Sinensis
Wuttipong Dongkumfu
We describe a new forest-dwelling Cyrtodactylus from Doi Inthanon, Chiang Mai Province, northern Thailand. Cyrtodactylus inthanon sp. nov. is characterized by a maximum known SVL of 87.3 mm; 18 to 20 longitudinal rows of dorsal tubercles; a continuous series of 34 to 37 enlarged femoro-precloacal scales, including four to six pitted (female) or porebearing (male) scales on each femur separated by a diastema from five pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and three to five irregular beige dorsal bands between limb insertions. The discovery of a new reptile endemic to Doi Inthanon reinforces the high importance of this mountain in terms of biodiversity conservation.
Live adult male holotype of Cyrtodactylus inthanon sp. nov. from Doi Inthanon, Chiang Mai Province, Thailand. Photo by M. Sumontha.
Preserved type series of Cyrtodactylus inthanon sp. nov. in dorsal view. Photo by O.S.G. Pauwels.
Preserved type series of Cyrtodactylus inthanon sp. nov. in ventral view. Photo by O.S.G. Pauwels.
Adult female Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang Mai Province. Photo by T. Sirisamphan.
Another adult female Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang Mai Province. Photo by W. Dongkumfu.
Figures - uploaded by Olivier S. G. Pauwels
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Content uploaded by Olivier S. G. Pauwels
All content in this area was uploaded by Olivier S. G. Pauwels on Nov 17, 2015
Accepted by A. Bauer: 9 Dec. 2014; published: 14 Jan. 2015
ISSN 1175-5326 (print edition)
(online edition)
Copyright © 2015 Magnolia Press
Zootaxa 3905 (4): 573
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http://dx.doi.org/10.11646/zootaxa.3905.4.9
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A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus)
from Doi Inthanon, Chiang Mai Province, northern Thailand
, MONTRI SUMONTHA
, NONN PANITVONG
, WUTTIPONG DONGKUMFU
THANA SIRISAMPHAN
& OLIVIER S. G. PAUWELS
Nakhon Ratchasima Zoo, 111 M. 1, Ratchasima-Pak Tongchai Rd., Chaimongkol, Mueang, Nakhon Ratchasima 30000, Thailand.
E-mail: kkunya@yahoo.com
Ranong Marine Fisheries Station, 157 Saphanpla Rd., Paknam, Mueang, Ranong 85000, Thailand.
E-mail: montri.sumontha@gmail.com
siamensis.org, 408/144 Phaholyothin Place Bldg 34FL, Phaholyothin Rd., Phayathai, Bangkok 10400, Thailand.
E-mail: npanitvong@gmail.com
Doi Inthanon National Park, 119 M. 7 Ban Luang Sub-district, Jom Thong District, Chiang Mai Province 50160, Thailand.
E-mail: w.peak60@hotmail.com
Département des Vertébrés Récents, Institut Royal des Sciences naturelles de Belgique, Rue Vautier 29, B-1000 Brussels, Belgium
Corresponding author. E-mail: osgpauwels@yahoo.fr
We describe a new forest-dwelling Cyrtodactylus from Doi Inthanon, Chiang Mai Province, northern Thailand. Cyrtodac-
tylus inthanon sp. nov. is characterized by a maximum known SVL of 87.3 mm; 18 to 20 longitudinal rows of dorsal tu-
bercles; a continuous series of 34 to 37 enlarged femoro-precloacal scales, including four to six pitted (female) or pore-
bearing (male) scales on each femur separated by a diastema from five pitted (females) or pore-bearing (male) precloacal
scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and three to five irregular beige dorsal
bands between limb insertions. The discovery of a new reptile endemic to Doi Inthanon reinforces the high importance of
this mountain in terms of biodiversity conservation.
Key words: Cyrtodactylus inthanon sp. nov., taxonomy, new species, Doi Inthanon National Park
With its summit culminating at 2565 m a.s.l., Doi Inthanon is the highest mountain of Thailand and shows a forest
species composition that is unique in the country (Khamyong et al. 2004). Its reptile fauna has never been
thoroughly investigated, and few records are available and are mostly undocumented (Nabhitabhata et al. 2004,
Nabhitabhata & Chan-ard 2005). No Cyrtodactylus has been so far reported. Pursuing our taxonomic work and
field surveys on this genus, that have already brought to light the existence of several micro-endemics in the
mountains of northern Thailand (Bauer et al. 2009, 2010, Ellis & Pauwels 2012, Kunya et al. 2014, Pauwels et al.
2014), we visited Doi Inthanon and encountered a population that is distinct from all known congeners in scalation
and color pattern. This population is consequently described hereafter as a new species, Cyrtodactylus inthanon sp.
Measurements and meristic counts follow Sumontha et al. (2012), Panitvong et al. (2014), Pauwels & Sumontha
(2014) and Pauwels et al. (2014). Paired meristic characters are given left/right. Numbers of supralabial and
infralabial scales are counted from the largest scale immediately posterior to the dorsal inflection of the posterior
portion of the upper jaw to the rostral and mental scales, respectively. The number of longitudinal rows of body
KUNYA ET AL.
Zootaxa 3905 (4) © 2015 Magnolia Press
tubercles was counted transversely across the center of the dorsum from one ventrolateral skin fold to the other.
The number of longitudinal rows of ventral scales was counted transversely across the center of the abdomen from
one ventrolateral skin fold to the other. The numbers of subdigital lamellae beneath the toes were counted from the
base of the first phalanx to the claw. The following measurements were taken with a digital caliper to the nearest
0.1 mm: AG: axilla to groin length, taken from the posterior margin of the forelimb at its insertion point on the
body to the anterior margin of the hind limb at its insertion point on the body; EarL: ear length, the greatest
horizontal distance of the ear opening; ForeaL: forearm length, taken on the dorsal surface from the posterior
margin of the elbow while flexed 90° to the inflection of the flexed wrist; HeadH: head height, the maximum depth
of head from the occiput to the throat; HeadL: head length, from the posterior margin of the retroarticular process
of the lower jaw to the tip of the snout; HeadW: head width, measured at the angle of the jaws; Internar: internarial
distance, measured between the nares across the rostrum; Interorb: interorbital distance, measured between the
anterior edges of the orbits; NosOrb: nostril to orbit distance, from the posterior margin of the external nares to the
anterior margin of the orbit; OrbD: orbit diameter, the greatest horizontal diameter of the orbit; OrbEar: orbit to ear
distance, from the anterior edge of the ear opening to the posterior edge of the orbit; SnOrb: snout to orbit distance,
from the tip of the snout to the anteriormost margin of the orbit; SVL: snout-vent length, taken from the tip of snout
to the vent; TailL: tail length, taken from the vent to the tip of the tail, original or regenerated; TailW: tail width,
taken at the base of the tail immediately posterior to the postcloacal swelling; TibiaL: tibia length, taken on the
ventral surface from the posterior surface of the knee while flexed 90° to the base of heel. Meristic characters
abbreviations: DorTub: dorsal tubercles; FemPi: femoral pits; FemPo: femoral pores; IL: infralabial scales;
InterorbSc: interorbital scales; PreclPi: precloacal pits; PreclPo: precloacal pores; SL: supralabial scales; Ven:
ventral scales. Museum and other acronyms: CUMZ-R: Chulalongkorn University Museum of Zoology, Reptile
Collection, Bangkok; IRSNB: Institut Royal des Sciences naturelles de Belgique, Brussels; MS: Montri
Sumontha's field number series; THNHM: Thailand Natural History Museum, National Science Museum,
Technopolis, Pathum Thani.
Cyrtodactylus inthanon sp. nov.
(Figs 1–7)
Holotype. THNHM 22550; adult male from Doi Inthanon (ca. 18°35'32" N, 098°29'12" E, alt. ca. 700 m asl),
Amphoe (= District) Jom Thong, Chiang Mai Province, northern Thailand. Collected by W. Dongkumfu on 7
Paratypes. THNHM 25605 (field no. MS 320) and CUMZ-R-0.2320 (field no. MS 503), adult females, same
locality, date and collector data as the holotype.
Diagnosis. Cyrtodactylus inthanon sp. nov. can be distinguished from all other congeneric species by its
maximum known SVL of 87.3 mm; 18 to 20 longitudinal rows of dorsal tubercles; a continuous series of 34 to 37
enlarged femoro-precloacal scales, including four to six pitted (female) or pore-bearing (male) scales on each
femur separated by a diastema from five pitted (females) or pore-bearing (male) precloacal scales; no precloacal
groove nor depression; transversely enlarged subcaudal scales; and three to five irregular beige dorsal bands
between limb insertions.
Description of holotype. Adult male. SVL 87.3 mm. TailL 102.0 mm (only first 12.4 mm original). Head
relatively long (HeadL/SVL 0.30), wide (HeadW/HeadL 0.63), not markedly depressed (HeadH/HeadL 0.41),
distinct from slender neck. Loreal region inflated, canthus rostralis not prominent. Snout elongate (SnOrb/HeadL
0.38), rounded, longer than orbit diameter (OrbD/SnOrb 0.70); scales on snout small, rounded to oval, granular to
weakly conical, mostly homogeneous, larger than those on crown, interorbital and occipital regions. Eye large
(OrbD/HeadL 0.27); pupil vertical with crenelated margins; supraciliaries short, those at posterior part of orbit
bearing small conical spines. Ear opening rounded, relatively small (EarL/HeadL 0.06); orbit to ear distance
subequal to orbit diameter (OrbEar/OrbD 1.01). Rostral much wider (3.3 mm) than deep (2.0 mm), rostral crease
nearly half of rostral height. Two enlarged supranasals separated from one another by an elongate internasal.
Rostral in contact with first supralabials, nostrils, supranasals and internasal. Nostrils oval, more or less laterally
directed, each surrounded by supranasal, rostral, first supralabial and two enlarged postnasals. Three or four rows
A NEW BENT-TOED GECKO FROM DOI INTHANON
of small scales separate orbit from supralabials. Mental triangular, wider (3.4 mm) than deep (2.9 mm). A single
pair of greatly enlarged postmentals in broad contact behind mental, each bordered anteromedially by mental,
anterolaterally by first infralabial, posterolaterally by an enlarged lateral chinshield, and posteriorly by two
granules. Supralabials to mid-orbital position 7/7, enlarged supralabials to angle of jaws 10/11. Infralabials 9/9.
Interorbital scale rows across narrowest point of frontal bone 24.
Body slender, relatively short (AG/SVL 0.43) with well-defined non-denticulate ventrolateral folds. Dorsal
scales weakly heterogeneous, domed to conical; regularly distributed tubercles (about five times size of adjacent
scales) extending from shoulder region onto tail base, smaller tubercles on postocular region, crown, occiput and
nape; most tubercles bearing a strong keel, less marked on lower flank tubercles; tubercles on posterior trunk and
sacral region most prominent; tubercles in 20 regular rows at midbody, typically separated from one another by two
or three dorsal granules. Ventral scales larger than dorsals, smooth, oval and subimbricate, largest on posterior
abdomen and in precloacal region. Midbody scale rows across belly between ventrolateral folds 34. Gular region
with homogeneous, smooth, juxtaposed granular scales. A continuous row of 35 enlarged femoro-precloacal scales,
as follows, from left to right: one unpitted poreless femoral scale + six pore-bearing femoral scales + a diastema of
nine unpitted poreless scales + five pore-bearing precloacal scales + a diastema of eight unpitted poreless scales +
six pore-bearing femoral scales. Postcloacal spurs each bearing three enlarged, conical scales.
Scales on palm and sole smooth, rounded to oval or hexagonal, slightly domed. Scalation on dorsal surface of
hind and forelimbs similar to body dorsum with enlarged tubercles interspersed among smaller scales. Fore and
hind limbs relatively long, slender (ForeaL/SVL 0.16, TibiaL/SVL 0.20). Digits long, slender, inflected at
interphalangeal joints, all bearing robust, slightly recurved claws. Basal subdigital lamellae broad, oval to
rectangular, without scansorial surfaces (7-8-8-7-7 right manus, 7-7-9-11-9 right pes); narrow lamellae distal to
digital inflection and not including ventral claw sheath: 9-9-12-13-10 (right manus), 11-11-14-12-12 (right pes); no
interdigital webbing. Relative lengths of digits: IV>III>V>II>I (manus), V>IV>III>II>I (pes). Mostly regenerated
tail, gently tapering to pointed tip, longer than SVL (TailL/SVL 1.17). Original and regenerated parts of tail with
enlarged median subcaudal scales.
FIGURE 1. Live adult male holotype of Cyrtodactylus inthanon sp. nov. from Doi Inthanon, Chiang Mai Province, Thailand.
Photo by M. Sumontha.
FIGURE 2. Preserved type series of Cyrtodactylus inthanon sp. nov. in dorsal view. Photo by O.S.G. Pauwels.
FIGURE 3. Preserved type series of Cyrtodactylus inthanon sp. nov. in ventral view. Photo by O.S.G. Pauwels.
FIGURE 4. Adult female Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang Mai
Province. Photo by T. Sirisamphan.
FIGURE 5. Another adult female Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang
Mai Province. Photo by W. Dongkumfu.
Coloration in life. Dorsal ground color of head, dorsum and tail brown. Three irregular light bands on dorsum
between fore- and hind limb insertions, bifurcated at their base on the flanks; their central part shows the same
color as the brown dorsal bands, and they bear prominent whitish tubercles, most contrasting on lower flanks
(Figure 1). Continuous nuchal loop connects orbits. Reticulate pattern on top of head. Ground color of upper
surfaces of fore- and hind limbs light brown, with irregular and discontinuous thin beige bands with scattered
whitish tubercles. Original part of tail showing two light rings which do not encircle the tail. Dorsal surface of
regenerated part of tail marbled with beige and light brown, darker ventrally. Iris dark brown. Throat, venter and
undersides of fore- and hind limbs uniformly beige.
Vari a t i o n. The paratypes resemble the holotype in most aspects of morphology and coloration (Figures 2–3).
Main morphometric and meristic characters of the type series are provided in Table 1. THNHM 25605 shows a
continuous series of 35 enlarged femoro-precloacal scales, as follows, from left to right: three unpitted poreless
femoral scales + four pitted femoral scales + a diastema of six unpitted poreless scales + five pitted precloacal
scales + a diastema of nine unpitted poreless scales + five pitted femoral scales + two unpitted poreless femoral
scales. CUMZ-R-0.2320 shows a continuous series of 37 enlarged femoro-precloacal scales, as follows, from left
to right: three unpitted poreless femoral scales + five pitted femoral scales + a diastema of eight unpitted poreless
scales + five pitted preanal scales + a diastema of eight unpitted poreless femoral scales + six pitted femoral scales
+ two unpitted poreless femoral scales. In THNHM 25605, supranasals partly separated in their inferior part by an
internasal. In CUMZ-R-0.2320, supranasals totally separated by an elongate internasal; rostral crease less than a
fourth of rostral height; postcloacal spurs each bearing four conical scales. Three to five irregular beige dorsal
bands between limb insertions (Figures 1–2, 4–7). Original tails show 11 or 12 light dorsal rings (Figures 4, 6–7).
Color pattern in juvenile (Figure 7) same as in subadult (Figure 6) and adults (Figures 1–2, 4–5).
TABLE 1. Meristic and morphometric (in mm) data for the type series of Cyrtodactylus inthanon sp. nov. Paired
meristic characters are given left/right.
Distribution and natural history. The species is known only from Doi Inthanon (Figure 8), from 700 to 1010
m a.s.l., where it is common. We encountered it while it was active at night on trees and large rocks along streams.
It moved slowly when disturbed by torch light and bit when handled. It was found at direct proximity to the reptiles
Acanthosaura lepidogaster (Cuvier) (Agamidae), Gekko gecko (Linnaeus), Hemidactylus frenatus Duméril &
Bibron and H. platyurus (Schneider), Hemiphyllodactylus chiangmaiensis Grismer, Wood & Cota (Gekkonidae),
Ahaetulla prasina (Boie) (Colubridae), Amphiesma khasiense (Boulenger) (Natricidae) and Trimeresurus
popeiorum Smith (Viperidae), and the amphibians Ansonia inthanon Matsui, Nabhitabhata & Panha (Bufonidae),
Leptolalax pelodytoides (Boulenger), Megophrys major Boulenger and M. minor Stejneger (Megophryidae),
Amolops marmoratus (Blyth), Hylarana nigrovittata (Blyth) and Odorrana livida (Blyth) (Ranidae). Captive
specimens ate meal worms and crickets and seemed to quickly dehydrate with decreasing hygrometry. The new
species' known range entirely falls within Doi Inthanon National Park.
Etymology. The specific epithet inthanon refers to the type locality. It is a noun in apposition, invariable. We
suggest the following common names: Took-kai Doi Inthanon (Thai), Doi Inthanon bent-toed gecko (English),
Cyrtodactyle du Doï Inthanon (French), Doi Inthanon Bogenfingergecko (German), Doiinthanonkromvingergekko
(Dutch).
Cyrtodactylus inthanon sp. nov. shows enlarged precloacal pore-bearing scales separated from femoral pore-
THNHM 22550 Holotype THNHM 25605 Paratype CUMZ-R-0.2320 Paratype
Sex Male Female Female
SVL 87.3 86.7 83.5
ForeaL 13.6 13.8 13.2
TibiaL 17.2 17.1 16.1
TailL 102.0 (last 89.6 regenerated) 51.7 (last 44.6 regenerated) 102.0 (last 42.0 regenerated)
TailW 6.2 7.2 6.1
AG 37.5 35.4 36.9
HeadL 26.4 25.0 24.2
HeadW 16.6 16.5 15.2
HeadH 10.7 10.6 9.5
OrbD 7.0 6.9 6.1
OrbEar 7.1 6.8 6.4
SnOrb 10.0 9.9 9.0
NosOrb 7.4 7.3 6.8
Interorb 5.6 4.8 4.6
EarL 1.6 1.4 1.3
Internar 2.4 2.4 2.1
DorTub201918
PreclPi/PreclPo 5 PreclPo 5 PreclPi 5 PreclPi
FemPi/FemPo 6/6 FemPo 5/4 FemPi 6/5 FemPi
Ven342929
SL 10/11 11/12 10/10
IL 9/9 10/10 9/9
InterorbSc 24 18 18
bearing scales by a diastema, and we thus hereafter provide a comparison with all other species known from
Cambodia, China, Laos, Myanmar, Peninsular Malaysia, Thailand and Vietnam sharing this condition (see
intraspecific comparisons and latest descriptions for this set of species in Bauer 2003, Nazarov et al. 2008, Ngo &
Bauer 2008, Rösler & Glaw 2008, Bauer et al. 2010, David et al. 2011, Sumontha et al. 2010, Grismer 2008, 2011,
Luu et al. 2011, Ziegler et al. 2013, Kunya et al. 2014, Pauwels et al. 2014a-b, Phung et al. 2014).
FIGURE 6. Subadult female Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang Mai
Province. Photo by W. Dongkumfu.
Among all these species, the ones showing similar dorsal patterns to Cyrtodactylus inthanon sp. nov. are the
Northern Thailand C. dumnuii and C. khelangensis Pauwels, Sumontha, Panitvong & Varaguttanonda, 2014, the
Northeastern Thailand C. kunyai Pauwels, Sumontha, Keeratikiat & Phanamphon, 2014, the Northern Vietnamese
C. huongsonensis Luu, Nguyen, Do & Ziegler, 2011 and the Southern Vietnamese C. huynhi Ngo & Bauer, 2008,
C. kingsadai Ziegler, Phung, Le & Nguyen, 2013 and C. ziegleri Nazarov, Orlov, Nguyen & Ho, 2008. From C.
dumnuii, C. inthanon sp. nov. differs by its lower number of ventral scale rows (29–34 vs. 40), its absence of
prominent white tubercles within the brown dorsal bands (vs. their conspicuous presence in C. dumnuii), its dark
brown iris (vs. green in C. dumnuii) and its lower number of light rings on original tail (11 or 12 vs. 13 or 14). It is
separated from C. huongsonensis by its higher number of dorsal tubercle rows (18–20 vs. 14–16), its lower number
of ventral scale rows (29–34 vs. 41–48) and 5 vs. 6 precloacal pores. It differs from C. huynhi by its enlarged
subcaudals (absent in C. huynhi), its lower number of precloacal pores (five vs. seven to nine) and its lower number
of ventral scale rows (29–34 vs. 43–46). C. inthanon sp. nov. can be distinguished from C. khelangensis by the
absence of precloacal pores in females (vs. presence in C. khelangensis), the presence of prominent white tubercles
on the nape and along the anterior and posterior border of the nuchal collar (absent in C. khelangensis), its poorly
contrasted reticulum on dorsal surface of head (vs. sharply contrasted in C. khelangensis) and its higher number of
light rings on the original tail (11 or 12 vs. 8). It differs from C. kingsadai by its lower number of precloacal pores
(5 vs. 7–9) and its lower number of ventral scale rows (29–34 vs. 39–46). C. inthanon sp. nov. differs from C.
kunyai by its higher number of precloacal pores in males (five vs. three) and its continuous nuchal loop (vs.
medially interrupted). It differs from C. ziegleri by its enlarged subcaudal scales (absent in C. ziegleri), by having
generally less dorsal tubercle rows (18–20 vs. 20–24) and ventral scale rows (29–34 vs. 33–39). Besides these
species, one should mention here another one which, although it lacks femoral pores, presents a similar dorsal
pattern, and also lives in Northern Thailand, namely C. doisuthep Kunya, Panmongkol, Pauwels, Sumontha,
Meewasana, Bunkhwamdi & Dangsri, 2014. C. inthanon sp. nov. can be separated from it, in addition to the
possession of femoral pores in males, by its lighter dorsal coloration, and its light dorsal bands which are wider
than in C. doisuthep and in lower number (three to five vs. six or seven). The regenerated tail portion in C. inthanon
sp. nov. is marbled light brown and beige, while it is velvet black in C. doisuthep (compare with figures in Kunya
et al. 2014).
FIGURE 7. Juvenile Cyrtodactylus inthanon sp. nov. (not collected) from type locality on Doi Inthanon, Chiang Mai Province.
Cyrtodactylus inthanon sp. nov. is distinct from all the remaining Burmese, Cambodian, Lao, Vietnamese and
Thai species showing precloacal and femoral pores separated by a diastema by the combination of the following
characters: enlarged subcaudals (absent in C. aequalis Bauer), five precloacal pores in males (11–12 in C.
annandalei Bauer, six in C. auribalteatus Sumontha, Panitvong & Deein, 2010, ten in C. bichnganae Ngo &
Grismer, seven to nine in C. brevipalmatus Smith, nine in C. caovansungi Orlov, Nguyen, Nazarov, Ananjeva &
Nguyen, 9–11 in C. consobrinus (Peters), 15 in C. russelli Bauer, nine in C. slowinskii Bauer, three or four in C.
takouensis Ngo & Bauer, none or one in C. thuongae Phung, van Schingen, Ziegler & Nguyen, eight or nine in C.
tigroides Bauer, Sumontha & Pauwels, 2003), 18 to 20 longitudinal tubercle rows (24 in C. aequalis, 22–24 in C.
auribalteatus, 22 in C. russelli, nine or ten in C. takouensis, 13 in C. tigroides), 29–34 ventral scale rows (24 in C.
aequalis, 43 in C. annandalei, 38-40 in C. auribalteatus, 37 in C. brevipalmatus, 44–48 in C. caovansungi, 58–65
in C. consobrinus, 35–41 in C. russelli, 39–40 in C. takouensis) and three to five irregular light bands between limb
insertions and reticulated dorsal head pattern (blotched dorsum pattern in C. aequalis; no reticulum on dorsal head
surface in C. annandalei; three straight bands between limb insertions in C. auribalteatus; blotched dorsum pattern
in C. brevipalmatus; blotched dorsum pattern and absence of dorsal head surface reticulum in C. russelli; blotched
dorsum pattern in C. slowinskii; three or four straight yellow bands between limb insertions in C. takouensis;
blotched dorsum pattern in C. thuongae, four straight bands between limb insertions in C. tigroides).
Cyrtodactylus inthanon sp. nov. does not seem to face any immediate specific conservation threat, but there
are general threats to the whole Doi Inthanon National Park, including use of forest resources by locals, disturbance
of wildlife by tourists and trails and facilities development (Hvenegaard & Dearden 1998). We did not record it in
the local or international pet trade. The discovery of a new reptile species endemic to Doi Inthanon reinforces the
high importance of this mountain in terms of biodiversity conservation. Since the species is relatively common, it
indicates that although Doi Inthanon is one of the Thai natural sites most visited by researchers and ecotourists, its
faunal inventory is obviously incomplete, and additional surveys would probably reinforce the value of the park.
FIGURE 8. Map of Thailand showing the position of Doi Inthanon, the type locality of Cyrtodactylus inthanon sp. nov.
We are grateful to Kriangsak Thanomphan, Chief of Doi Inthanon National Park, for supporting our study. We
thank Tanya Chan-ard (THNHM, Pathum Thani), Georges Lenglet (IRSNB, Brussels) and Kumthorn Thirakhupt
(CUMZ, Bangkok) for providing access to the respective herpetological collections under their care, to Thomas
Ziegler (Cologne Zoo) for commenting on the manuscript, and to Watchira Sodob for preparing the map.
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APPENDIX. Comparative material examined.
Cyrtodactylus auribalteatus: see Sumontha et al. (2010: type series); C. brevipalmatus: see Pauwels and Chan-ard (2006); C.
doisuthep: see Kunya et al. (2014: type series); C. dumnuii: see Bauer et al. (2010: type series); C. intermedius: IRSNB 17011,
Nakhon Ratchasima, Thailand; C. khelangensis: see Pauwels et al. (2014: type series); C. oldhami complex: see Pauwels &
Chan-ard (2006) and Pauwels et al. (2000); C. peguensis: see Pauwels et al. (2000); C. sanook: see Pauwels et al. (2013: type
series); C. sumonthai: see Bauer et al. (2002: type series); C. tigroides: see Bauer et al. (2003: type series).
... The abbreviations are as follows: / = data not available in literature; -=; characteristic not present; + = characteristic present but not uniquely determined; * = broken or regenerated tail. Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Pauwels et al. 2004Pauwels et al. , 2013Pauwels et al. , 2014Pauwels et al. , 2018Nguyen et al. 2010;Ngo & Pauwels 2010;Sumontha et al. 2010Sumontha et al. , 2012Sumontha et al. , 2014Sumontha et al. , 2015Shi & Zhao 2010;Chan-ard & Makchai 2011;David et al. 2011;Schneider et al. 2011;Grismer et al. 2012Grismer et al. , 2018Johnson et al. 2012;Kunya et al. 2014Kunya et al. , 2015Luu et al. 2014Luu et al. , 2015Luu et al. , 2016Luu et al. , 2017Nazarov et al. 2014;Panitvong et al. 2014 King 1962;Das 1993;Günther & Rösler 2002;Rösler et al. 2007;Rösler & Glaw 2008;Oliver et al. 2009;Welton et al. 2009Welton et al. , 2010Iskander et al. 2011;Kathriner et al. 2014;Riyanto et al. 2014Riyanto et al. , 2015Riyanto et al. , 1016Riyanto et al. , 2018Harvey et al. 2015;Hartmann et al. 2016;, Riyanto et al. 2018from Uetz et al. 2019). The abbreviations are as follows: / = data not available in literature; -=; characteristic not present; + = characteristic present but not uniquely determined; * = broken or regenerated tail. ...
A new species of Cyrtodactylus (Squamata: Gekkonidae) from southern Vietnam
Sabrina Ostrowski
Dang Trong Do
Minh Le
We describe a new species of Cyrtodactylus on the basis of four specimens collected from Phu My, Binh Dinh Province, southern Vietnam. Cyrtodactylus phumyensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: size small (SVL up to 66.8 mm); two internasals; dorsal tubercle rows 18 or 19 at midbody; ventral scale rows 33-41; ventrolateral folds slightly developed; each thigh with 5-7 enlarged femoral scales; femoral pores absent in males and female; a series of 5-7 precloacal pores plus a pitted, enlarged precloacal scale in males; 6 pitted, enlarged precloacal scales in female; paravertebral tubercles 20-23; lamellae under toe IV 18-21; small subcaudal scales, not transversely enlarged; two irregular dark longitudinal stripes on shoulder. In phylogenetic analyses, the new species is recovered as a member of the Cyrtodactylus irregularis species group, and strongly supported as a sister taxon of C. cucdongensis from Khanh Hoa Province.
... Comparisons. We compared the new species with its congeners from Vietnam and neighboring countries in mainland Indochina, including Laos, Cambodia, Myanmar, China (Yunnan), Thailand, and peninsular Malaysia based on examination of specimens (see Appendix) and data obtained from the literature (Smith 1917(Smith , 1921a(Smith ,b, 1935Taylor 1963;Ulber & Grossmann 1991;Ulber 1993;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Ziegler et al. 2002Ziegler et al. , 2010Ziegler et al. , 2013Pauwels et al. 2004Pauwels et al. , 2013Pauwels et al. , 2014aPauwels et al. ,b, 2016Nguyen et al. 2006Hoang et al. 2007;Orlov et al. 2007;Grismer & Ahmad 2008;Ngo 2008Ngo , 2011Ngo & Bauer 2008;Ngo & Chan 2010, 2011Ngo & Pauwels 2010;Sumontha et al. 2010Sumontha et al. , 2012Sumontha et al. , 2014Chan-ard & Makchai 2011;David et al. 2011;Schneider et al. 2011;Luu et al. 2011Luu et al. , 2014Luu et al. , 2015Luu et al. , 2016aGrismer et al. 2012Grismer et al. , 2016Grismer et al. , 2017Kunya et al. 2014Kunya et al. , 2015Nazarov et al. 2014;Panitvong et al. 2014;Le et al. 2016;Connette et al. 2017;Pham et al. 2017;andNguyen et al. 2015a, 2017). The new species can be distinguished from other known species of Cyrtodactylus by morphological characters (see Table 2). ...
... It is therefore likely that the sequence of C. bichnganae was erroneously assigned and we recommend that the sample from the type specimen be re-sequenced in future study to clarify this issue. Smith 1917Smith , 1921aSmith ,b, 1935Taylor 1963;Ulber & Grossmann 1991;Ulber 1993;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Ziegler et al. 2002Ziegler et al. , 2010Ziegler et al. , 2013Pauwels et al. 2004Pauwels et al. , 2013Pauwels et al. , 2014aPauwels et al. ,b, 2016Nguyen et al. 2006Nguyen et al. , 2014Hoang et al. 2007;Orlov et al. 2007;Grismer & Ahmad 2008;Ngo 2008Ngo , 2011Ngo & Bauer 2008;Ngo & Chan 2010, 2011Ngo & Pauwels 2010;Sumontha et al. 2010Sumontha et al. , 2012Sumontha et al. , 2014Chan-ard & Makchai 2011;David et al. 2011;Schneider et al. 2011;Luu et al. 2011Luu et al. , 2014Luu et al. , 2015Luu et al. , 2016aGrismer et al. 2012Grismer et al. , 2016Grismer et al. , 2017Kunya et al. 2014Kunya et al. , 2015Nazarov et al. 2014;Panitvong et al. 2014;Le et al. 2016;Connette et al. 2017;Pham et al. 2017; ...
A new species of Cyrtodactylus (Squamata: Gekkonidae) from northwestern Vietnam
Anh Van Pham
Hanh Ngo
Truong Nguyen
We describe a new species of Cyrtodactylus on the basis of seven specimens collected from karst forests of Son La and Dien Bien provinces, Vietnam. Cyrtodactylus taybacensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: medium size (SVL up to 97.5 mm); dorsal tubercles in 13-16 irregular rows; ventral scale rows 30-38; ventrolateral folds present without interspersed tubercles; each thigh with 11-13 enlarged femoral scales; femoral pores absent in males and females; precloacal pores 11-13 in males, five or 15 pitted scales in females, in a continuous row; postcloacal tubercles two or three; lamellae under toe IV 16-20; subcaudal scales transversely enlarged; dorsal head with dark brown marking, oval, triangle and arched shape; five brown dorsal bands between limb insertions. In phylogenetic analyses, the new species is recovered as a member of the Cyrtodactylus wayakonei species group, and strongly supported as the sister taxon of C. cf. bichnganae from Son La City in all analyses.
... Doi Inthanon National Park in Chiang Mai Province spanned three districts (Chom Thong, Mae Wang, and Mae Chaem) with altitudes ranging from 400 to 2562 m, and temperatures ranging from 1 o C to 23 o C (Anonimus 2001). Located in a mountainous area and covered with rich plant diversities (Khamyong et al. 2004), many new species of bryophytes (Akiyama et al. 2010), ferns (Iwatsuki et al. 1998), reptiles (Kunya et al. 2015) and insects (Takaoka & Choochote 2005) were uncovered. Fungal diversity in Doi Inthanon National Park has been studied, specifically aquatic fungi (Chuaseeharonnachai et al. 2013) and Xylariaceae (Laessøe et al. 2013), however, none reported on discomycetes. ...
Morpho-phylogenetic insights reveal Bisporella montana as Calycina montana comb. nov. (Pezizellaceae, Helotiales)
Anis S Lestari
Thilini Chethana Kandawatte
Calycina-like specimens were collected in Doi Inthanon National Park, Thailand. The combined phylogeny of ITS and LSU based on maximum likelihood and bayesian inference analyses confirmed the taxonomic placement of our specimen in Pezizellaceae. Based on morphology and multi-gene phylogeny, the new discomycete specimen was identified as Calycina montana comb. nov., and was documented for the first time in Thailand. Calycina montana is characterized by the amber color, stipitate apothecia, gelatinized ectal excipulum, cylindrical asci and ellipsoidal ascospores. The morphology was observed together with the type strain, Bisporella montana. Our phylogenetic analyses and observation of the morphology of the type strain also confirmed the new combination of Bisporella shangrilana as Calycina shangrilana comb. nov., which resolves the confusion of Bisporella placement in Pezizellaceae.
... However, a few Cyrtodactylus spp. have been found in the northern part of Thailand including Cyrtodact ylus doisuthep, Cyrtodact ylus dumnuii, C yrtodact ylus inthanon, C yrtodact ylus er ythrops, Cyrtodact ylus maelanoi, and Cyrtodact ylus khelangensis, which were found in only three provinces (Chiang Mai, Mae Hong Son, and Lampang) (Bauer et al., 2009;Bauer et al., 2010;Grismer et al., 2020a;Kunya et al., 2014;Kunya et al., 2015;Pauwels et al., 2014b). This finding is inconsistent with those for many lineages of Cyrtodactylus in the surrounding countries, e.g., Myanmar and Laos, which have continuous massif to those in northern Thailand with many limestone ecosystems that are suitable habitats. ...
A New Cyrtodactylus Species (Reptila: Gekkonidae) from Nan Province, Northern Thailand
Siriwadee Chomdej
Waranee Pradit
Parinya Pawangkhanant
Chatmongkon Suwannapoom
Here, a new species of bent-toed gecko, Cyrtodactylus phukhaensis sp. nov., is described from Doi Phu Kha, Nan province, Thailand based on molecular and morphological evidence. A phylogeny based on NADH dehydrogenase subunit 2 (ND2) and its flanking tRNAs places the new species in the chauquangensis group as a sister taxon to Cyrtodact ylus wayakonei. The new species can be differentiated from other members of the chauquangensis group by having a unique combination of 7 or 8 supralabials, 23–28 longitudinal rows of dorsal tubercles, 8–10 infralabials, 9 femoral pores, 7 precloacal pores, 40–47 ventral scales, and a lack of bands crossing the temporal area. In addition, results from a chromosome study of C. phukhaensis sp. nov. showed that the new species has a diploid chromosome number of 40 with a fundamental number of 46. Te formula of the karyotype was as follows: 2n (40) = 2m + 4sm + 34t. Our findings suggest that further studies of Cyrtodactylus biodiversity in northern Tailand are needed.
... Doi Inthanon, formerly known as Doi Angka, is the highest mountain of Thailand, with its summit at 2565 m a.s.l. The small area represents a classic example of a montane hotspot for biodiversity and endemicity (e.g., Khamyong et al. 2003;Teejuntuk et al. 2003;Sungkajanttranon et al. 2018), although little is documented about the herpetofauna (Kunya et al. 2015;Wu et al. 2019). ...
Integrative taxonomy reveals a new species of Leptobrachella (Anura: Megophryidae) from the mountains of northern Thailand
Jinmin Chen
Yunhe Wu
Jing Che
The genus Leptobrachella is one of the most speciose and taxonomically troubling groups of Asian anurans. Herein, we describe a new species of Leptobrachella from Doi Inthanon, Chiang Mai Province, Thailand based on the integration of morphological, acoustic and molecular data. The new species, Leptobrachella murphyi sp. nov., is morphologically distinguished from its congeners on the basis of body size, dorsal skin texture and coloration, ventral coloration and pattern, degree of webbing and fringing on the fingers and toes, iris coloration in life, pattern of markings on flanks and pattern of body macroglands. It also differs from its phylogenetically close congeners by an uncorrected p-distance of >9.6% for a fragment of 16S rRNA. The advertisement call of the new species consists of 4.5–4.7 kHz (at 15 °C) and without a distinct introductory note. Leptobrachella murphyi sp. nov. likely occurs across the Thanon Thong Chai Range and analyses provide evidence of unknown biodiversity and species composition on Doi (mountain) Inthanon. In addition, the congeneric species L. minima was also confirmed in Doi Inthanon. The coexistence pattern of Leptobrachella in Doi Inthanon deserves further study. As Thailand's highest mountain and biodiversity reservoir, the need for further biological exploration is urgent given ongoing habitat loss and degradation.
... Additionally, mountain ranges in the northern and western areas show a similar pattern to those in some parts of southern Thailand where many species of bent-toed geckos have previously been found 14,15,21,22,[24][25][26] . Although, a few studies of Cyrtodactylus species in northern Thailand have been performed, nevertheless molecular phylogenetic analysis was not included in these works [27][28][29][30][31][32] . Several species of Cyrtodactylus have recently been described as cryptic species both in Thailand (C. ...
Phylogenetic analyses of distantly related clades of bent-toed geckos (genus Cyrtodactylus) reveal an unprecedented amount of cryptic diversity in northern and western Thailand
Shiping Gong
Cyrtodactylus species are the most diverse of the geckos and are widely distributed in Southeast Asia, including Thailand. However, their patterns of distribution, especially in northern and western parts of Thailand, remain unknown because few Cyrtodactylus species in these regions have been described. Thus, a data set of mitochondrial NADH dehydrogenase 2 (ND2) gene and flanking tRNAs from Cyrtodactylus found in northern and western Thailand, including contiguous areas, was assembled to elucidate phylogenetic relationships and identify the distribution patterns of these geckos. The results showed four well-supported clades, a northwestern clade (A), a northern clade (B), a western clade (C), and a special clade characterized by specific morphological features (D). Clades A-C were grouped with strong support by the geography of their localities from northern Thailand (Mae Hong Son and Chiang Mai Provinces) along the Tenasserim mountain ranges to Phang-Nga Province, Thailand. Clade D is a distinct clade of Cyrtodactylus species characterized by a tuberculate and prehensile tail and distributed widely in mainland Southeast Asia. Overall, the results suggest a pattern of geographic separation and distribution of Cyrtodactylus in northern and western Thailand. Additionally, this study provides evidence of a hidden biodiversity of Cyrtodactylus in these regions.
... Femoral and precloacal pore counts were excluded from the PCA due to their absence in females. PV, ET4, UT4, and TT4 were also excluded because these diagnostic characters were not evaluated for Cyrtodctylus inthanon by Kunya et al. (2015). Because the data were skewed by values ranging from 4-36, these data were log-transformed prior to analysis in order to normalize their distribution so as to ensure characters with very large and very low values could not over-leverage the results owing to intervariable nonlinearity. ...
A new species of Cyrtodactylus Gray (Squamata; Gekkonidae) from the Thai Highlands with a discussion on the evolution of habitat preference
Larry Lee Grismer
Attapol Rujirawan
Korkhwan Termprayoon
Anchalee Aowphol
A new gekkonid lizard, Cyrtodactylus maelanoi sp. nov., from Mae Hong Son Province of the Thai Highlands is described using an integrative taxonomic analysis based on morphology, color pattern, and the mitochondrial gene NADH dehydrogenase subunit 2 (ND2) and its flanking tRNAs. Phylogenetic analyses place the new species within clade 1 of the C. sinyineensis group and as the sister species to C. inthanon with an uncorrected pairwise sequence divergence of 3.9%. Collection data gathered in the field indicate that C. maelanoi sp. nov. is a habitat generalist. Reconstruction of the ancestral habitat preference for the C. sinyineensis group by way of stochasitc character mapping (SCM) indicates that karstic environments were the ancestral condition out of which the general habitat preference of the ancestor of C. maelanoi sp. nov. and C. inthanon and that of C. amphipetreaus and C. doisuthep evolved three times independently. Additionally, SCM demonstrated that the evolution of a granitic habitat preference from a karst-adapted ancestor happened in C. aequalis. The discovery of a new upland species in the Thai Highlands brings into focus the understudied nature of the mountain systems of western Thailand and the need for their continued exploration and conservation.
... Prior to this study, there were 36 species of Cyrtodactylus Gray known from Thailand which is comparable to that of neighboring Myanmar (45 species), Laos (23 species), Vietnam (43 species), and Peninsular Malaysia (33 species) but far exceeds that of China (five species) and Cambodia (eight species) (Uetz et al. 2020). However, in stark contrast to the neighboring countries, the phylogenetic relationships of 44% (16) of the Thai species are unknown because most of the recent species descriptions did not include molecular data (Bauer et al. 2009(Bauer et al. , 2010Chan-ard & Makchai, 2011;Kunya et al. 2014Kunya et al. , 2015Pauwels et al. 2014aPauwels et al. ,b, 2016, thus precluding any downstream comparative analyses (see Grismer et al. 2020a). To this end, we add C. inthanon and C. doisuthep and lower that value to 38%. ...
A new species Cyrtodactytlus Gray (Squamata: Gekkonidae) from western Thailand and the phylogenetic placement of C. inthanon and C. doisuthep
Nikolay Poyarkov
A new species of Cyrtodactylus from Tak Province, Thailand, Cyrtodactylus amphipetraeus sp. nov., is described using an integrative taxonomic analysis based on morphology, color pattern, and the mitochondrial gene NADH dehydrogenase subunit 2 (ND2). The phylogenetic analyses place the new species within the C. sinyineensis group which was previously thought to be endemic to the Salween Basin in southern Myanmar. The phylogeny also places C. inthanon in the C. sinyneensis group which is expanded herein to also include the group's sister species C. doisuthep. Along with C. amphipetraeus sp. nov., these are the first three species of the C. sinyineensis group to be found outside of Myanmar east of the Tenasserim Mountains. The Tenasserim Mountain region is discussed as an area of cladogeneic turnover.
... The herpetological diversity of montane forest is high. Herpetological surveys in the montane forests in northern Thailand during the last decade have led to numerous discoveries of species of amphibians and reptiles, including the first record of snakes, Plagiopholis blakewayi Boulenger (Tillack et al., 2006), Sinonatrix yunnanensis Rao and Yang (Pauwels et al., 2009), Ptyas nigromarginata (Vogel and Hauser, 2013), Parafimbrios laos Teynié et al. (Teynié and Hauser, 2017), Protobothrops (Vasaruchapong et al., 2017), Dendrelaphis nigroserratus Vogel et al., (2012), Liopeltis frenatus (Günther) and Ptyas multicincta (Roux) (Figueroa et al., 2016;Hauser, 2018); rare species were rediscovered, such as Opisthotropis spenceri Smith (Chuaynkern et al., 2014) and Paratapinophis praemaxillaris Angel (Murphy et al., 2008); new species and new country records were also recorded of lizards, Diploderma yunnanense (Anderson) (Manthey and Denzer, 2012;Wang et.al., 2018), Hemiphyllodactylus chiangmaiensis Grismer et al. (2014), Cyrtodactylus doisuthep Kunya et al. (2014), C. inthanon Kunya et al. (2015) and Pseudocalotes kakhienensis (Anderson) from Doi Pui, Doi Inthanon and Omkoi District, Chiangmai Province (Montri Sumontha and Kirati Kunya observed data); new species of amphibians include Limnonectes taylori Matsui et al. (2010), Leptolalax zhangyapingi Jiang et al. (2013), Tylototriton anguliceps Le et al. (2015), T. uyenoi Nishikawa et al. (2013), Gracixalus seesom Matsui et al. (2015), Minervarya chiangmaiensis (Suwannapoom et al.) (Suwannapoom et al., 2016;Niyomwan et al., 2019). We suggest that the further research in montane forests in northern Thailand and adjacent Myanmar and Laos requires additional attention. ...
Protobothrops kelomohy sp. nov. (Squamata: Viperidae), the Second Known Species of Lance-Headed Pit Viper from Thailand
Taksa Vasaruchapong
Niruth Chomngam
Lawan Chanhome
Protobothrops kelomohy sp. nov. is described from dry evergreen forest in Chiang Mai Province, northern Thailand, based on morphology and a molecular phylogeny. It differs from congeners by the following combination of characters: 1) relatively large body size (total length up to 1,310 mm); 2) dorsal scale rows 23-23-17, all keeled; 3) ventral scales 231-234; 4) subcaudal scales 80-84, paired; 5) supralabials 8-9; 6) infralabials 12-13; 7) typical lance-pattern on upper head surface; 8) three bold vertical facial stripes; and 9) dorsal and tail brown, each scale with dark reddish-brown transverse blotches, edged in black, somewhat fused together as a median zigzag line on dorsum, a row of large ventrolateral blotches on each side, each darker marking edged by a narrow yellow line. The new discovery and other recent findings suggest that further herpetological research is required in the montane forest of northern Thailand and adjacent Myanmar and Laos. Data on the natural history of the new species and an updated key to the species of Protobothrops are provided. This is the second species of Lance-headed Pit Viper recorded from Thailand after P. mucrosquamatus.
... Comparisons. We compared the new species with its congeners from Vietnam and neighboring countries in mainland Indochina, including Laos, Cambodia, Myanmar and Thailand based on examination of specimens (see Appendix) and data obtained from the literature (Smith 1917(Smith , 1921a(Smith ,b, 1935Taylor 1963;Ulber 1993;Bauer 2002Bauer , 2003Bauer et al. 2002Bauer et al. , 2003Bauer et al. , 2009Bauer et al. , 2010Ziegler et al. 2002Ziegler et al. , 2010Ziegler et al. , 2013Pauwels et al. 2004Pauwels et al. , 2013Pauwels et al. , 2014aPauwels et al. ,b, 2016Nguyen et al. 2006Nguyen et al. , 2014Hoang et al. 2007;Orlov et al. 2007;Ngo 2008;Ngo & Bauer 2008;Ngo & Pauwels 2010;Ngo & Chan 2010Sumontha et al. 2010Sumontha et al. , 2012Sumontha et al. , 2014Chan-ard & Makchai, 2011;David et al. 2011;Schneider et al. 2011;Luu et al. 2011Luu et al. , 2014Luu et al. , 2015Luu et al. , 2016aGrismer et al. 2012;Kunya et al. 2014Kunya et al. , 2015Nazarov et al. 2014;Panitvong et al. 2014;Le et al. 2016;and Connette et al. 2017). ...
A new species of Cyrtodactylus (Squamata: Gekkonidae) and the first record of C. otai from Son La Province, Vietnam
We describe a new species of Cyrtodactylus on the basis of four specimens collected from the limestone karst forest of Phu Yen District, Son La Province, Vietnam. Cyrtodactylus sonlaensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: maximum SVL of 83.2 mm; dorsal tubercles in 13–15 irregular rows; ventral scales in 34–42 rows; ventrolateral folds prominent without interspersed tubercles; enlarged femoral scales 15–17 on each thigh; femoral pores 14–15 on each thigh in males, absent in females; precloacal pores 8, in a continuous row in males, absent in females; postcloacal tubercles 2 or 3; lamellae under toe IV 18–21; dorsal head with dark brown markings, in oval and arched shapes; nuchal loop discontinuous; dorsum with five brown bands between limb insertions, third and fourth bands discontinuous; subcaudal scales distinctly enlarged. In phylogenetic analyses, the new species is nested in a clade consisting of C. huongsonensis and C. soni from northern Vietnam and C. cf. pulchellus from Malaysia based on maximum likelihood and Bayesian analyses. In addition, we record Cyrtodactylus otai Nguyen, Le, Pham, Ngo, Hoang, Pham & Ziegler for the first time from Son La Province based on specimens collected from Van Ho District.
A new species of the genus Cyrtodactylus Gray, 1827 from central Laos (Squamata: Gekkonidae)
Patrick David
Nicole Schneider
A new species of the gekkonid genus Cyrtodactylus Gray, 1827, Cyrtodactylus teyniei spec. nov., is described based on a single adult female from Borikhamxay Province, central Laos. It is distinguished from any other Cyrtodactylus species by the combination of the following characters: (1) SVL 89.9 mm, (2) longitudinal rows of dorsal tubercles 19, (3) ventral scales in 38 rows between ventrolateral skin folds, (4) ventrolateral skin folds without tubercles, (5) enlarged femoral and precloacal scales 54 (precloacal scales 8, plus 23 femoral scales on each side), (6) precloacal depression lacking, (7) precloacal pores 14, (8) subdigital lamellae under the firth toe 19-20, (9) enlarged median subcaudals, and (10) nuchal band absent, dorsum with blotched pattern. An extended comparison with other species of the genus Cyrtodactylus, in particular species recently described from Laos and Vietnam, is provided. Cyrtodactylus teyniei spec. nov. is the seventh species of Cyrtodactylus known to occur in Laos.
Cyrtodactylus Auribalteatus (Squamata: Gekkonidae), A New Cave-Dwelling Gecko From Phitsanulok Province, Thailand
GRIDSADA DEEIN
A new cave-dwelling species of the gekkonid lizard genus Cyrtodactylus Gray, C. auribalteatus sp. nov. is described on the basis of animals collected from a cave in Thung Salaeng Luang National Park, Phitsanulok Province, Thailand, bringing the number of species in this genus in Thailand to 18. The new species shares over all appearances with other cave-dwelling Cyrtodactylus spp. in having large eyes, a long snout, reduced tubercles, absence of caudal tubercles, and slender limbs for climbing. The new species is characterized by its moderate size (snout-vent length to at least 98.07 mm), weakly conical tubercles in 22-24 rows at midbody, high number of ventral scale across midbody (38-40), absence of precloacal groove, present of a single series of 6 precloacal pores and 4-5 femoral pores on each thigh separated by a diastema, broad subcaudal plates, and its dorsal pattern of three dark bands between limb insertions. It is the seventh species of cave-dwelling Cyrtodactylus, including the sandstone cave species, C. jarujini, recorded from Thailand.
Cyrtodactylus saiyok sp. nov., a new dry evergreen forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Kanchanaburi Province, western Thailand
Jitthep Tunprasert
Olivier S. G. Pauwels
We describe Cyrtodactylus saiyok sp. nov. from a dry evergreen forest on a limestone hill in Khao Krajae, Sai Yok District, Kanchanaburi Province, western Thailand. It is characterized by a maximal known SVL of 61 mm; 18-19 longitudinal rows of dorsal tubercles; 23 or 24 ventral scale rows between ventrolateral skin folds; a continuous series of enlarged femoro-precloacal scales, including 5 pore-bearing precloacal scales (males); no precloacal groove or depression; transversely enlarged subcaudal scales; a complete black nuchal loop; a W-shaped band above shoulders and 3-5 irregular, medially interrupted or not, black dorsal bands between limb insertions. It is the sixth reptile species that is possibly endemic to Sai Yok District.
A new Cyrtodactylus (Squamata: Gekkonidae) from Huong Son limestone forest, Hanoi, northern Vietnam
Vinh Quang Luu
Huy Quang Do
We describe a new species of the genus Cyrtodactylus based on two adult specimens from Huong Son limestone forest, Hanoi, Vietnam. Cyrtodactylus huongsonensis sp. nov. is distinguished from the remaining Indochinese bent-toed geckos by a combination of the following characters: medium-sized, with a maximum SVL of 89.8 mm; dorsal pattern consisting of dark nuchal loop, neck band and five in part irregular transverse body bands between limbs; two enlarged lateral chin-shields in contact with first postmental pair; dorsal tubercles present on occiput, body, forearms, hind limbs and tail base; 14–16 irregularly running dorsal tubercle rows; ventrals in 41–48 longitudinal rows at midbody; lateral skin folds present, without interspersed tubercles; six precloacal pores plus in total 17 femoral pores in males, which are separated by 8–12 poreless scales; enlarged femoral scales and precloacal scales present; three postcloacal spurs in males; subcaudal scales transversely enlarged. This is the 24 th species of Cyrtodactylus known from Vietnam.
Cyrtodactylus erythrops (Squamata: Gekkonidae), a new cave-dwelling gecko from Mae Hong Son Province, Thailand
A. M. Bauer
Tunyakorn Kunya
A new cave-dwelling species of Cyrtodactylus Gray is described from Mae Hong Son Province in northern Thailand. The new species, C. erythrops sp. nov., is characterized by its moderate size (snout-vent length to at least 78 mm), relatively large, closely-spaced, flattened tubercles in 18–20 irregular rows at midbody, low number of ventral scales across midbody (28), absence of precloacal groove, presence of precloacal and femoral pores separated by a diastema, broad subcaudal plates, and dorsal pattern of dark spots and blotches. It is the fifth species of cave-dwelling Cyrtodactylus recorded from Thailand and its discovery adds to the mounting evidence that this genus exhibits unprecedented levels of localized endemism throughout tropical Southeast Asia.
A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Malaysia including a literature survey of mensural and meristic data in the genus
Herbert Rösler
Frank Glaw
We describe a new gekkonid lizard from Batang Padang, Perak province, Malaysia, based on a single specimen collected almost 100 years ago. Cyrtodactylus stresemanni sp. nov. apparently differs from all other species in the genus by large tubercles on the ventral side of the tail, suggesting an isolated position within the genus. A literature survey of meristic and mensural data of all described Cyrtodactylus species revealed further diagnostic characters to distinguish C. stresemanni from all other species.
Descriptions of two new species of Cyrtodactylus Gray 1827 (Squamata: Gekkonidae) endemic to southern Vietnam
Ngo Van Tri
Two new species of the gekkonid lizard genus Cyrtodactylus Gray, 1827 are described on the basis of voucher specimens collected in isolated mountains separated from the Truong Son Mountain Range, southern Vietnam. Both species were collected from shallow caves, as have been many of the recently described Cyrtodactylus from across Southeast Asia. Both species, C. takouensis sp. nov. from Ta Kou Nature Reserve, Binh Thuan Province and C. huynhi sp. nov. from Chua Chan Mountain, Dong Nai Province, are characterized by a small number of enlarged femoral scales separated from the precloacal scales by a large diastema. They may be distinguished from one another and from all other congeners by differences in rows of tubercles and ventral scales, subcaudal scalation and dorsal color pattern. These new taxa bring the number of currently recognized Cyrtodactylus species in Vietnam to twelve.
We describe a new gekkonid lizard from Batang Padang, Perak province, Malaysia, based on a single specimen collected almost ICK) years ago. Cyrtodactylus stresemanni sp. nov. apparently differs from all other species in the genus by large tubercles on the ventral side of the tail, suggesting an isolated position within the genus. A literature survey of meristic and mensural data of all described Cyrtodactylus species revealed further diagnostic characters to distinguish C. stresemanni from all other species.
Descriptions of seven new Cyrtodactylus (Squamata: Gekkonidae) with a key to the species of Myanmar (Burma)
Lizards of Peninsular Malaysia, Singapore and their Adjacent Archipelagos
Discover more about: Thailand
Malayemys Systematics & Distribution
Timothy R. Brophy
Suthep Wiboonatthapol
These studies were conducted to get a better idea of the number and kinds of species within the Malayemys complex of Southeast Asia.
Taxonomic diversity, phylogeny and biogeography of Southeast Asian herpetofauna
Anna B Vassilieva
Thiti Ruangsuwan
Project is focused on application of molecular, morphological and bioacoustic approaches to studies of taxonomic diversity, phylogenetic relationships and historical biogeography of amphibians and reptiles in Southeast Asia, in particularly, the Indochinese Peninsula (Vietnam, Laos, Cambodia and Vietnam) and adjacent territories. ... [more]
marine fisheries
Thammanoon Temchai
Veerayuth Lauhachinda
A new forest-dwelling Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Doi Suthep, Chiang...
June 2014 · Zootaxa
Aumporn Panmongkol
Siriwat Dangsri
We describe a new forest-dwelling Cyrtodactylus (Gekkonidae) from Doi Suthep, Chiang Mai Province, northern Thailand. Cyrtodactylus doisuthep sp. nov. is characterized by a maximal known SVL of 90.5 mm; 19 or 20 longitudinal rows of dorsal tubercles; a continuous series of 34 or 35 enlarged femoro-precloacal scales, including six or seven pitted scales on each femur (male and females) separated ... [Show full abstract] by a diastema from six pitted (females) or pore-bearing (male) precloacal scales; no precloacal groove or depression; transversely enlarged subcaudal scales; and six or seven irregular thin beige dorsal bands between limb insertions.
First record of the Cyrtodactylus brevipalmatus group (Squamata: Gekkonidae) from Vietnam with descr...
May 2021 · Zootaxa
Dzung Trung Le
Saly Sitthivong
Tung Thanh Tran
A new species of the genus Cyrtodactylus is described from Dien Bien Province, northwestern Vietnam based on morphological and molecular data. Cyrtodactylus ngati sp. nov. can be distinguished from remaining congeners by the following combination of characters: maximum SVL 69.3 mm; dorsal pattern consisting of six dark irregular transverse bands between limb insertions; inter-supranasals one; ... [Show full abstract] dorsal tubercles present on occiput, body, hind limbs and on first half of tail; 17-22 irregular dorsal tubercle rows at midbody; lateral folds clearly defined, with interspersed tubercles; 32-38 ventral scales between ventrolateral folds; 13 precloacal pores separated by a diastema of 5/5 poreless scales from a series of 7/7 femoral pores in enlarged femoral scales; precloacal and femoral pores absent in females; 1-3 postcloacal tubercles on each side; transversely enlarged median subcaudal scales absent. In the molecular analyses, the new species is shown to be the sister taxon to C. interdigitalis from Thailand. This is the 47 th species of the genus Cyrtodactylus and the first member of the C. brevipalmatus species group recorded from Vietnam.
Cyrtodactylus khelangensis, a new cave-dwelling Bent-toed Gecko (Squamata: Gekkonidae) from Lampang...
January 2014 · Zootaxa
Varawut Varaguttanonda
We describe Cyrtodactylus khelangensis sp. nov. from a limestone cave in Pratu Pha, Mae Mo District, Lampang Province, northern Thailand. It is characterized by a maximal known SVL of 95.3 mm; 16-20 longitudinal rows of dorsal tubercles; a continuous series of 37-40 enlarged femoroprecloacal scales, including six-seven pitted or pore-bearing scales (males) or one or two pitted scales (females) on ... [Show full abstract] each femur separated by a diastema from 2-6 pore-bearing precloacal scales (males and females); no precloacal groove nor depression; transversely enlarged subcaudal scales; and four irregular brown dorsal bands between nuchal loop and hind limb insertions.
Cyrtodactylus kunyai (Squamata: Gekkonidae), a new cave-dwelling Bent-toed Gecko from Loei Province,...
Kaweesak Keeratikiat
Eakarit Phanamphon
We describe a new cave-dwelling Bent-toed Gecko, Cyrtodactylus kunyai (Squamata: Gekkonidae) from Suan Hin Pha Ngam, Nong Hin District, Loei Province, northeastern Thailand. It is characterized by a maximal known snout-vent length of 87.9 mm, a banded dorsal pattern with a medially interrupted nuchal loop and four of five brown bands between nuchal loop and hind limb insertions and three ... [Show full abstract] bicolored band interspaces between limb insertions, a dark orangeish iris, a continuous series of enlarged femoro-precloacal scales with 5-6 femoral pores on each side separated by a diastema from 3 precloacal pores in males (no pores in females), 19 irregularly arranged dorsal longitudinal tubercle rows at midbody, 34 ventral scale rows between ventrolateral skin folds, transversely enlarged subcaudal plates, and no precloacal groove.
A new Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Phetchaburi Province, Thailand
March 2016 · Zootaxa
A new Bent-toed Gecko, Cyrtodactylus phetchaburiensis sp. nov. is described from the Tha Yang District of Phetchaburi Province, western Thailand. It is a medium-sized Cyrtodactylus (SVL to at least 63.2 mm), with small, mostly keeled tubercles in 20 regular longitudinal rows on dorsum; 33 scales across mid-venter between lowest rows of flank tubercles; enlarged row of femoral scales present; five ... [Show full abstract] precloacal pores in male, femoral pores and precloacal groove absent; 5–6 broad basal lamellae and 11 narrow distal lamellae beneath digit IV of pes; and a single median row of transversely enlarged subcaudal scales present. It has a dorsal colour pattern of large, dark, diffusely-edged markings on a fawn background and a pair of dark scapular patches. The species is a member of the Central Indochinese (Thai-Myanmar) clade of Cyrtodactylus and is most closely related to C. oldhami (Theobald), from which it differs in colour pattern.
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text:
Watch a Movie, Feed the Hungry
If you are fortunate enough to catch a screening of Ready, Set, Bag! in a movie theater, you will be doing more than just supporting a delightful independent film about state champions as they prepare for the National Grocers Best Bagger Competition.
via Repair The World Read More
By Dvora Meyers / October 25, 2010
This article originally appeared on Repair The World.
If you are fortunate enough to catch a screening of Ready, Set, Bag! in a movie theater, you will be doing more than just supporting a delightful independent film about state champions as they prepare for the National Grocers Best Bagger Competition. You will also be feeding the hungry. A dollar from each ticket sale is donated to a local food bank. This is the film distribution strategy of Oren Jacob, the film's executive producer and his wife, co-director and producer, Justine Jacob.
The initial idea was not their own. It originated at film festivals in Austin, TX and Prescott, AZ where local organizers suggested that the pair do a fundraiser with the local food bank. "And it was just a really fun event," Oren recalls of those first two screenings. "The audience was super stoked. They saw a fun, entertaining film and at the same time helped support the food banks and fed folks in their communities. When we decided to take the film out ourselves, we thought we should just do the same." During the course of a single 100 person screening, they generate enough in donations to purchase 400-500 meals.
The most obvious beneficiaries are the needy people being fed-the film screenings have funded over 8,000 meals (and counting) thus far. But the hungry and food banks have not been the only ones to benefit from the fundraiser-film pairing. The partnership has greatly helped the movie. When asked if donating a portion of ticket sales has negatively affected the film's bottom line, he emphatically insists that it's quite the reverse. "No," he says, "it actually enables us to have a bottom line. It's a pretty central part of our operating procedures in the business. On a secondary level, to put together the kind of publicity and marketing team that can get our film out as effectively as the food banks' publicists can would be very hard. It [the pairing] has increased our operational complexity and decreased our running costs pretty substantially." He hopes that other filmmakers consider this sort of charitable partnership as not just something good to do for PR but as intrinsically helpful to their business.
Oren and his wife have been surprised at how gratifying it was to combine their professional pursuits and charitable impulses. "In my normal life, I go to work and make a salary. There's not this whole other thing going on by helping people in need. But now every time we screen the film, or sell tickets to it or do anything that helps us get Ready, Set, Bag! out there, we are feeding people in need at the same time. And that adds a very positive, meaningful undercurrent to the business and it's very motivating. Distributing an independent film is a very hard thing to do-millions of hours, countless sleepless nights-and it's not like we're going to make a 100 billion dollars on the movie. We're not Avatar or Toy Story 3."
Also, this has given them yet another opportunity to impress the importance of helping others on their three young children. Tikkun Olam is one of the rules in the Jacob house (along with other equally important gems such as "Safety first" and "Manage your own gear"). "We talk about tikkun olam with them a lot, about bringing positive change to the community that we live in," Oren says.
If you'd like to bring Ready, Set, Bag! to you area, contact Oren Jacob at info at readysetbag dot com or reach out to him on Facebook or Twitter.
Tagged with: Groceries , hunger , justine jacob , oren jacob , ready set bag
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Unique Feather and leather earrings made to order. Tribal, gypsy, bohemian, or hippie, these striking primal adornments are the perfect compliment to any bold and daring ensemble! Perfect for festival season!
This listing is for one made to order pair of the above pictured earrings. Please allow 1-2 weeks for construction of your handmade feather earrings. If you have a specific deadline, please let me know and I'll do my best to accommodate you! Please note, due to variations in natural materials, your feathers will vary slightly from the photo, but will be just as beautiful! If you would like to see your earrings before they ship, please request a preview photo!
Each of my feather creations is lovingly hand crafted with meticulous attention to detail and aesthetic. I use only the highest quality feathers and source my materials in small batches so no two items are exactly alike. Great care is taken to construct pieces which will endure travel and adventure, and everything from the connecting pieces to the wire and earrings have been tested for quality and durability. Some feathers, however, are naturally fragile, so handling your jewelry gently is recommended.


Bail vs Bond: What's the Difference?
What Happens After a DV Charge in North Carolina?
At Raleigh Bail Bonds, our team is composed of licensed bondsmen who understand that the service we provide makes a difference. Throughout the entire time you'll be working with us, you can count on experiencing professional, discreet, prompt and friendly service.


Growing up, I would imagine that behind the last row of houses, which I could see from my balcony, there is a landscape. Usually, I would be on the balcony at dusk, looking at the space between two buildings…because I wasn't able to see anything but the sky and the top of one tree, I would imagine that there is a meadow leading to the forest and onwards to hills, instead of a big factory that is really there. I noticed that the balcony was turned towards the west and that fact become symbolic for me – since the nineties, everyone in my surroundings considered that the "West" meant spiritual freedom and a normal relaxed life.
I discovered the book "De rerum natura" (On the nature of things) written by Roman philosopher Lucretius (1st century AD), who tried to summarize some of the Greek and Roman viewpoints. It motivated me to start investigating the parallels that can be drawn between contemporary society and ancient theories about nature. Using atomism, ethics and history of observing heavens I am developing works that reflect ancient attitude and contemporary potential of personal autonomy.
Works are analogies of modern person's choices and prospects in contemporary society along with their aftermath. Tangible, repeating / time lapse drawings of celestial phenomena, quantum level quivers, changing landscapes and manmade objects talk about an individual in contemporary society, (the illusion of) free will and fate.


These are the FULL SHARE distributions. Half Shares are a little more than 1/2 of this. As you can see, shares start out smaller when little is available for harvest and evolve over the growing season. Splitting a half share is not recommended.


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One aspect of US securities laws that frequently trips up the unwary is the fact that US law regulates both the offer and the sale of securities. This is unlike most other securities regulation regimes, and it seems counterintuitive to most folks. After all, if you offer securities in a way you shouldn't have and no-one buys anything, no harm, no foul, right? Sorry, that's not the way it works.
To make it even worse, "offer" has a special meaning in Securities World. You might think an offer is something like "Would you like to buy my Preferred Shares at $100 each?" Most normal people would, but sorry, wrong again.
So let's back way up to basics. The Securities Act of 1933 says that you cannot offer or sell securities to the public unless (a) the offering is registered with the SEC, or (b) there is an available exemption from registration. The crowdfunding exemption under Section 4(a)(6) of the Securities Act won't be available until the SEC issues its regulations, so right now you cannot sell or offer any securities.
A chapter in a book describing your R&D process.
A statement that a company's officer makes on a TV interview.
A tweet that you are looking forward to doing crowdfunding when it's legal.
A discussion of your capital-raising alternatives.
If you do any of these things you may end up making an unregistered offer of securities in violation of the Securities Act. This could lead to problems when you actually come to do an offer of crowdfunding securities, especially as we get closer to the point at which it's legal.
So before you say anything about future capital-raising plans, talk to a lawyer!
*The SEC reads it for the articles.


Welcome to Shelldale Family Gateway! We are a non-profit, community benefit, charitable organization that aims to enrich the quality of life for the Willow Road area of Guelph. Our multicultural approach strives to meet the needs of our community, offering programs and supports in over 29 languages.
The Shelldale Family Gateway community hub is located in the Shelldale Centre, home to several community organizations, and provides an integrated service model to the community.
We are a non-profit, community benefit, charitable organization that aims to enrich the quality of life for the Willow Road area of Guelph. Our multicultural approach strives to meet the needs of our community, offering programs and supports in over 29 languages.
Shelldale offers approximately 30 programs over the course of a year. New programs are constantly being added, so please check in frequently to make sure you are up to date.
Volunteering is a great way to contribute to your community, meet people, and gain new skills and experiences. Many of our programs depend on the support of volunteers to be successful. We have a wide variety of opportunities throughout the week in all of our program areas.
to build a better world through building a better community.


I love this block. Pieces of type like this are usable works of art. The letterform itself is so heavy but it isn't brutish. The arrow shapes that appear in the counter spaces are fascinating to me. The serifs aren't as thick as many styles of clarendon, which makes the letter look a bit more modern.
You can see the pantograph marks in the counter spaces. The face is one of the most beautiful I've seen. The color of the wood is rich and clear, and you can see the grain crossing the face vertically. This block was cut from a particularly large tree, as it is 4 inches tall and you can still barely see the curve to the grain. The splotches left on the block are some result of ink being left on them, and not all of it came off.


We stayed in Alain and Paula's apartment for almost a month. We had several questions before booking the apartment / arriving, and they went out of their way to help us. The apartment itself is nice and cozy, very clean, and on a quiet residential street. The best part is to see the mountains from the window when you wake up in the morning. You open your eyes and it's the first thing you see through the big windows. There is a small organic grocery literally around the corner, and to get to the big grocery store it's a 10 minute walk. It's also 10 minutes to walk down to the lake, and about 20-25 to Vieille annecy, which is the beautiful old town where you can just get lost and wander for hours. I would also recommend checking out the bar / restaurant Saint Charles. We never ate there but they have good coffee / drinks and very nice people! If you are coming to Annecy, this is a really great place to stay for a good price, and it is close to many things where you can easily walk! Thanks Alain and Paula!
Nous sommes restés 6 mois dans cet appartement pendant la durée d'octobre 2013 jusqu'à mars 2014 et nous devons dire que nous aimions beaucoup notre séjour à Annecy. Cet appartement est très lumineux, bien équipé avec le wifi haut débit. Le quartier est très calme et tranquille et bien aussi situé près de grand magasins Galeries La Fayette, la poste, supermarché Casino, et pharmacie. Il est également à distance de marche du lac d'Annecy environ 10-15mins. Le propriétaire est facilement joignable en cas de besoin. Nous bien sûr reviendrons à l'avenir!!
Je suis venu pour la première fois à Annecy à l'âge de 10 ans et franchement je ne regrette pas du tout d'y être revenu et nous nous allons y revenir l'année prochaine avec un grand plaisir.
Une semaine qui restera gravée dans nos mémoires, avec beaucoup de beaux souvenirs.
The place was perfect for what we wanted for our week in Annecy. The location is about a 15 minute walk from the center and about 10 mins from the lake in a quiet residential apartment block. The apartment is small but perfect for two people with a small, but adequate kitchen, living space and semi private bed. Alain and Paula had just completed a renovation when we rented the place and it looks great.
We would recommend anyone staying there to walk up to Annecy Vieux and eat up there as it was a completely different experience to the central tourist area. The meals we had up there were the best we had in Annecy, but you will need to make a booking, or be prepared to wait.
I would recommend this place for someone who is looking for a place to base themselves in Annecy away from the center.
We spent a happy week in this cozy apartment! It is located in a quiet residential neighborhood about 20 minutes' walk from the historic center, in a unit that is two flights up. The decor is as charming as it looks in the pictures. The apartment has two comfortable, full-size sofa-beds. Annecy itself is a magnificent town and if you come in the Spring, as we did, then you may see the town's tulip displays, which are spectacular. Also -- Annecy has lots of public playgrounds for children and fun things to do on the lake, such as paddle-boating. Thank you Alain and Paula!
Beautiful and sweet apartment situated close to the centre of annecy. myself and 3 other girls stayed here for a week and it was a perfect apartment for us. Every thing you need was provided, Internet, bed sheets and a wonderful shower. Best of all the owners are really nice and easy to talk with. I would suggest this apartment for every one.
Cet appartement est vraiment dans une copropriété très calme dans un quartier très agréable.
Le grand plus le linge de maison est fourni!
Je suis reste 2 mois dans l'appartement et tout etait agreable.Le fait d'avoir l'internet est d'une grande utitlite.
I have already rented the apartment few times in a winter time and I am going to rent it again because it is very well equipped - it has just everything I need for comfortable stay, very well situated - I mean short distance from the historical center, the lake, the mountains and shopping centers and the owner is a very friendly and kind person. The flat is always bright and the view from the windows is fantastic.
L'appartement est très agreable avec tout le confort nécéssaire à un bon séjour.
La proximité du lac et des commerces est appréciable.
Découvrez le site de notre autre location meublée.


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The average distance from London to Bergerac is 740 kilometers (equals to 460 miles or 399 nautical miles). Your flight will departure from London, United Kingdom and arrive on Bergerac, France.
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Short Business Description: Waters International is a company dealing in oil field drilling equipment. We have a large collection of industrial drilling rigs mud pumps, surplus oilfield equipment, triplex mud pumps, mud tanks, rotary tables and many other drilling products. We also provide mud pump parts and accessories.
Since its establishment in 1970, Waters International has come a long way and established itself as a reliable and a leading provider of oil and gas drilling equipment for sale and rent. Right from the beginning, the leadership in our company has collected in-depth experience of the oil and gas business and developed a clear understanding of its nature too. We go all-out to provide complete customer satisfaction quickly and effectively.


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The young driver will receive a warm welcome in registration before being introduced to their instructor who will make sure they are sitting comfortably and have good visibility before an in-car briefing explaining the controls of the car.
The young driver will then take the wheel of their first dream supercar for a mind blowing 10 minutes of sheer exhilaration driving a real supercar at our York or Teesside venues, or 6 laps driving in Stafford.
For car number 3 the young driver will be feeling more confident and will, by now, be more relaxed and will thoroughly enjoy driving 10 minutes in supercar number 3. (6 laps in Stafford). The smile should be from ear to ear and very prominent on the photographs that will have been taken by our friendly resident photographer.
After collecting a certificate the driver can view the photographs which are available to purchase.
Drives in York or Middlesbrough will be 10 minutes per car and in Stafford the youngster will drive each car for 6 laps of the track.
Cars to choose from include Ferrari, Lamborghini, Audi R8, Porsche, Subaru, Ford Mustang GT, Nissan GT-R (York and Stafford only) Lotus Elise (not in Stafford) and Aston Martin.
Drivers should be 4ft 10 ins tall or above and aged between 11-17 years old without a driving license.


Follow the questions below and send us your appointment request. Once Kresta receives your information, she'll get right on it and get things rolling.
If you believe this is an emergency, we recommend you call us at (540) 483-9382 where you'll be able to immediately talk with a Wisler Plumbing employee (24/7).
How Can Wisler Plumbing Help You?


text:
The Sentinel Watch
Home » Student Profiles » Musical Nurse Earns BSN, MSN and Forges Plans for DNP Educational Leadership
Musical Nurse Earns BSN, MSN and Forges Plans for DNP Educational Leadership
When Georgiana Ibach first became a nurse, it was to support her first love: music. The University of Delaware graduate (piano) was working as a piano teacher and organist, but eventually craved a more regular schedule. "I went to Beebe School of Nursing, graduated, and went into the operating room at Beebe Medical after working the medical-surgical floor," says the Delaware native. Later, Georgiana joined Bayhealth's Milford Campus in surgical services—all while raising her triplets (two girls and one boy).
In 2009, Georgiana returned to Beebe Medical, joining the cardiovascular/thoracic team in the operating room. "I decided it was time to open some doors, so I went back to school," she says. By then, Georgiana was a single parent and needed to support herself and her four children. Her search for the right online program brought her to American Sentinel. "I really liked the flexibility and the way it fit my life." She enrolled that same year, taking a two-year break from her musical endeavors—she is an accompanist/organist in her community for churches, musicals, and at various high schools—so that she could focus on school.
A move to utilization review
One year after earning the BSN, Georgiana got a job in utilization review, reviewing Medicare and commercial insurances and working with various agencies on approvals, denials and appeals. "When it came up, I thought, 'I'm going to give this a try,'" she says. "I had the clinical background and now the education to do it. It felt so great to get hired for the position."
A few years went by and Georgiana decided to go back to school for her master's degree. When it came to choosing the right program, she didn't hesitate to call American Sentinel once again to explore her MSN specialization options. "I returned to American Sentinel because of the great teachers and positive experience," she says. She started the MSN, nursing education specialization in hopes that she will one day become an educator. "In my job, I do a little education of new trainees and that sort of thing and I really enjoy it. I could definitely see myself in the classroom one day."
Another degree on her resume
In 2016, Georgiana graduated with the MSN. Not one to rest on her laurels, she is already plotting her return to school—this time for a DNP. It is a longtime goal that stems back to her childhood. "I'm the only one of my seven siblings to have gone to college," she says. Her endeavors have certainly rubbed off on her children. Her daughter is in culinary school at Johnson and Wales, her other daughter is studying theater and drama at Towson University and her son is eyeing law or business school while studying at Albright College. Georgiana also has a seventh-grader, adopted from Russia, who studies piano.
An American Sentinel ambassador
Georgiana is currently an ambassador for American Sentinel University, speaking to prospective students about her experience. "I love encouraging other people to go to school," she says. Her advice: if she can do it, so can anyone else. Despite having one child still at home, a full-time job and a full calendar of upcoming shows in which she is playing piano—she played in 22 shows last summer and substituted many Sundays as a church organist. She is currently rehearsing Cinderella at the local theater, which debuts in November. At American Sentinel, Georgiana managed to graduate with a 3.8 GPA.
"American Sentinel was a great experience and I'm proud I did it," she says. "I have wanted to earn these degrees since I had my children, but life tends to take over. Once I did this for myself, it really inspired me to keep going. I'm happy I proved to myself and to my kids that you can get an education at any age."
Inspired by Georgiana's story? An MSN program can be your passport to a specialty nursing field, like nursing education, informatics, nursing management and organizational leadership, or infection control. Specialized knowledge forms the foundation of these nursing fields. When you acquire new knowledge, you can apply it to nursing practice in ways that enhance patient care and improve outcomes.
Have you dreamed of earning your BSN, MSN or DNP? With American Sentinel, you can make that dream a reality.
Read the other student success stories for more inspiration.
Posted in Student Profiles
Topics: bsn bsn student stories msn msn student stories
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American Sentinel University is accredited by the Distance Education Accrediting Commission, DEAC, 1101 17th Street NW, Suite 808, Washington, DC 20036, (202) 234-5100, www.deac.org, which is listed by the U.S. Department of Education as a nationally recognized accrediting agency. The Distance Education Accrediting Commission is recognized by the Council for Higher Education Accreditation (CHEA). American Sentinel University's next on-site evaluation by DEAC will be in Fall, 2019. DEAC Consumer Information Disclosure Form.
American Sentinel's RN to BSN, RN to BSN/MSN and Master of Science Nursing programs are accredited by the CCNE, 655 K Street, NW, Suite 750, Washington, DC 20001, (202) 887-6791. Recognized by the U.S. Secretary of Education as a specialized professional accreditation agency, the CCNE is an autonomous branch of the American Association of Colleges of Nursing, the only accrediting body devoted exclusively to evaluating baccalaureate and graduate nursing degree programs. American Sentinel University's next on-site evaluation by CCNE will be in Spring 2024.
ACEN Accreditation
American Sentinel University's Doctor of Nursing Practice program is accredited by the Accreditation Commission for Education in Nursing (ACEN), formerly NLNAC, 3343 Peachtree Road, NE, Suite 850, Atlanta, GA 30326, (404) 975-5000, fax (404) 975-5020, a specialized accrediting body recognized by the Council for Higher Education Accreditation (CHEA). American Sentinel University's next on-site evaluation by ACEN will be in Fall 2026.
Candidate for regional accreditation by the Higher Learning Commission (HLC)
At its meeting on June 30, 2016, the Higher Learning Commission (HLC) Board of Trustees voted to grant American Sentinel University the status of candidate for accreditation.
Institutions seeking accreditation by HLC are required to complete a four-year period of candidacy before being evaluated for full accreditation. To earn and maintain candidacy, American Sentinel must fully meet the Commission's Eligibility Requirements and Assumed Practices and demonstrate the capacity to meet all the Criteria for Accreditation within the candidacy period. American Sentinel University's hosted an on-site evaluation for initial accreditation by HLC on September 9 and 10 of 2019. The decision-making process involves multiple steps and occurs over several months. No decision will be made before the board meets in February of 2020 and it may take until July of 2020 before the University is notified of the decision.
Higher Learning Commission • 230 South LaSalle Street, Suite 7-500 • Chicago, IL 60604 • info@hlcommission.org • 800.621.7440
© 2020 American Sentinel University, all rights reserved.
American Sentinel University 10065 E. Harvard Avenue, Ste. 450, Denver, CO 80231 USA Phone: 1.800.729.2427 • Email: info@AmericanSentinel.edu
Program offerings include
• Registered Nurse to B.S. Nursing (RN to BSN) • M.S. Nursing (MSN) (7 Specializations) • Nurse Practitioner (2 Specializations) • Doctor of Nursing Practice (DNP)