Sensor networks are expected to evolve into long-lived, open, ubiquitous, multi-purpose networked systems. We propose a new concept called AutoNomouS netWorked sEnsoR system (ANSWER) whose mission is to provide in situ users with real-time, secure information that enhances their situational and location awareness. To the best of our knowledge, solutions that accomplish this goal do not yet exist. ANSWER finds immediate applications to both overt and covert operations ranging from tactical battlefield surveillance to crisis management and homeland security. The architectural model of ANSWER is composed of a large number of sensors and of a set of (mobile) aggregation-and-forwarding nodes that organize and manage the sensors in their vicinity. In this paper we present the main features that enable ANSWER to effectively and efficiently provide secure, QoS-aware information services to in situ mobile users; namely, secure dynamic task-based networking and in-network storage to support application-level tasks and queries (each specified with desired QoS and security attributes), while hiding network-level details; and a model-based methodology exploiting QoS and security trade-offs for smart AFN mobility subject to application and network requirements and constraints. This new concept is in sharp departure from the prevalent view in NSS design that networking is independent of the task(s) at hand and information processing and storage are, primarily, the responsibility of remote entities.