13. Tribe ATHROISMEAE 山黄菊族 shan huang ju zu Authors: Yousheng
Chen & Arne A. Anderberg Annual or perennial herbs, shrubs, or
small trees, sometimes aromatic. Leaves simple, alternate or
fasciculate, petiolate or sessile, abaxial surface with or
without sessile-glandular trichomes. Capitula terminal, in
corymbiform cymes, or solitary, sometimes congested, glomerule-
like or in rounded secondary capitula, pedunculate, sometimes
sessile, radiate, disciform, or discoid. Phyllaries 1-4-seriate,
sometimes much reduced or absent, herbaceous, subequal or
gradate. Receptacles convex to conical, sometimes subglobose to
globose, paleate or epaleate. Ray florets when present female;
corollas yellow or white. Anthers usually ecalcarate, shortly
caudate or not caudate, rarely tails well developed and
branched; endothecium with polarized thickenings. Achenes brown
or black, obcompressed to subterete, sometimes with thickened
striations or ribs, sometimes with a proliferation of spongy
cells apically, glabrous or with ciliate