Spawns only during daytime (Ref. 27547). At spawning, males lie
close to the bottom in the current of the spawning area while
females stay along the banks and in still water (Ref. 27547). A
female moves from the bank of the stream usually escorted by 2-4
males to the spawning area at the center of the stream. The
males crowd beside her; egg deposition occurs as the males try
to clasp the female with their pelvic fins or vibrate against
her with their anal fins. This spawning act lasts for 3-5
seconds and may occur as often as 6-40 times per hour. After the
eggs are deposited, the sexes separate and return to their
previous stream positions (Ref.1998). Fish that moved out of a
lake to spawn generally return to the lake a few days after
spawning. However, river-resident fish may stay on or near the
spawning area for much of the summer (Ref. 27547). Many spawn in
two or even three consecutive years but others may skip one or
two years between spawning (Ref. 10928). - Scott, W.B. and E.J.
Crossman 1973 Freshwater