According to zoologist Jakob Bro-Jørgensen, the colour of the male's coat can reflect the levels of androgen, a male hormone, which is highest during rutting.[22] Comparing the subspecies, T. d. derbianus is characterised by 15 body stripes, smaller size, and a rufous colour, while T. d. gigas is larger, a sandy colour, and has 12 body stripes.[22]