Fujihara Award, Fujihara Foundation of Science (2005) Japan Academy Prize, Japan Academy (2006) Asahi Prize, Asahi Shimbun (2009) Kyoto Prize in Basic Sciences (2012)[3] Gairdner Foundation International Award (2015) International Prize for Biology (2015) Keio Medical Science Prize (2015) Rosenstiel Award (2015) Wiley Prize in Biomedical Sciences (2016) Nobel Prize in Physiology or Medicine (2016)[4]