By identifying the ancient type of DNA attributed to the Phoenicians, geneticist Pierre Zalloua was also able to chart their spread out of the eastern Mediterranean. These markers were found in unusually high proportions in non-Lebanese samples from other parts of the "Mediterranean coast where the Phoenicians are known to have established colonies, such as Carthage in today's Tunisia."[1] The markers were also found among samples of Maltese and Spaniards, where the Phoenicians were also known to have established colonies.