In the early dynastic period, he was depicted in animal form, as a black canine.[44] Anubis's distinctive black color did not represent the animal, rather it had several symbolic meanings.[45] It represented "the discolouration of the corpse after its treatment with natron and the smearing of the wrappings with a resinous substance during mummification."[45] Being the color of the fertile silt of the River Nile, to Egyptians, black also symbolized fertility and the possibility of rebirth in the afterlife.[46] In the Middle Kingdom, Anubis was often portrayed as a man with the head of a jackal.[47] An extremely rare depiction of him in fully human form was found in the tomb of Ramesses II in Abydos.[45][10]