Abreast of the current knowledge of anatomy, physiology, and bacteriology, which together make up modern pathology, he was able to utilise to the best advantage the stores of specimens collected by John Hunter. His dissections enabled him to correlate many facts for the first time, and his results were set forth in his lectures. In 1885, he lectured on the structure and life history of the hydrozoa; in 1886 and 1887 on the organs of hearing; in 1889 and again in 1896 on the integumental system; in 1890 on phosphorescent organs and colour; in 1891 on secondary sexual characters; in 1895 on the endoskeleton; in 1897 on joints, and on the protection and nourishment of the young; in 1899 on the alternation of generations.