Breeding takes place in the spring and summer, with spawning occurring in April and May in the Adriatic Sea, at depths between 60 and 70 m (200 and 230 ft). The larvae soon move to shallower depths and are pelagic, as are the juveniles at first. At a length of about 5 cm (2 in) the juveniles move to the coast and become demersal, often congregating in estuaries, and sometimes swimming a short distance upstream.