Fulani culture continued to emerge in the area of the upper Niger and Senegal Rivers. The Fulani were cattle-keeping farmers who shared their lands with other nearby groups, like the Soninke, who contributed to the rise of ancient Ghana, with eastward and westward expansion being led by nomadic groups of cattle breeders or the Fulɓe ladde. While the initial expansionist groups were small, they soon increased in size due to the availability of grazing lands in the Sahel and the lands that bordered it to the immediate south.