Lake Tahoe is a tributary watershed drainage element within the Truckee River Basin, and its sole outlet is the Truckee River, which continues on to discharge to Pyramid Lake. Because of the sensitivity of Truckee River water quality (involving two protected species, the cui-ui[90] sucker fish and the Lahontan cutthroat trout), this drainage basin has been studied extensively. The primary investigations were stimulated by the U.S. Environmental Protection Agency, which funded the development of the DSSAM model to analyze water quality below Lake Tahoe.