In terrestrial vertebrates, the earliest forms were large amphibious piscivores 400 million years ago. While amphibians continued to feed on fish and later insects, reptiles began exploring two new food types, other tetrapods (carnivory), and later, plants (herbivory). Carnivory was a natural transition from insectivory for medium and large tetrapods, requiring minimal adaptation (in contrast, a complex set of adaptations was necessary for feeding on highly fibrous plant materials).[1]