The story follows characters from Germany, Britain, the United States and the Soviet Union, who become linked by events leading to World War II, and continues through the war and its immediate aftermath. The major characters are often children of the characters who were seen in Fall of Giants. The novel covers a wide range of world events during the period, including the rise of Nazism, the ascent of Franco in Spain, the short-lived growth of British fascism, Action T4, the Battle of Moscow, the Blitz, the Normandy landings, the attack on Pearl Harbor, the development of the American atom bomb and the Soviet one, the Battle of Berlin and many more.