1999: Seymour S. Kety 2000: Sydney Brenner 2002: James E. Darnell 2004: Matthew Meselson 2006: Joseph G. Gall 2008: Stanley Falkow 2010: David Weatherall 2012: Donald D. Brown and Tom Maniatis [2] 2014: Mary-Claire King 2016: Bruce M. Alberts [3] 2018: Joan Argetsinger Steitz