The work of Frederic E. Clements (1916) was particularly influential. He proposed that a community of vegetation is a superorganism and that communities develop in a fixed pattern of successional stages from inception through to some single climax state or to a self-regulating state of equilibrium. By analogy, an individual is born, grows, matures, and dies, but the community which the individual inhabited continues to grow and exhibit properties which are greater than the sum of the properties of the parts.