The El Castillo cave contains the oldest known cave painting: a large red stippled disk in the Panel de las Manos was dated to more than 40,000 years old using uranium-thorium dating in a 2012 study.[1] This is consistent with the tradition of cave painting originating in the Proto-Aurignacian, with the first arrival of anatomically modern humans in Europe.[2] A 2013 study of finger length ratios in Upper Paleolithic hand stencils found in France and Spain determined that the majority were of female hands, overturning the previous widely held belief that this art form was primarily a male activity.[3]