Later, overnight September 27–28, the hurricane moved directly over Dry Tortugas National Park in the Florida Keys with sustained winds of 120 mph (195 km/h),[166] as it was undergoing an eye replacement cycle. Once that cycle was complete, Ian rapidly strengthened, reaching Category 4 intensity on the morning of September 28, according to data from a Reserve Hurricane Hunter aircraft,[167] with its maximum sustained winds rising to a high-end Category 4 speed of near 155 mph (250 km/h) by 10:35 UTC.[168] Later, at 19:05 UTC, Ian made landfall at Cayo Costa Island, with winds of 150 mph (240 km/h).[169] And then, an hour and a half later, on the Florida mainland, just south of Punta Gorda, with maximum sustained winds of 145 mph (230 km/h).[170] It then proceeded across Central Florida before moving off shore into the Atlantic Ocean at 15:00 UTC near Cape Canaveral as strong tropical storm.[171] At the time, the storm was exhibiting some extra tropical characteristics, such as a comma-pattern and some frontal features in its outer circulation.