Mollisols occur in savannahs and mountain valleys (such as Central Asia, or the North American Great Plains). These environments have historically been strongly influenced by fire and abundant pedoturbation from organisms such as ants and earthworms. It was estimated that in 2003, only 14 to 26 percent of grassland ecosystems still remained in a relatively natural state (that is, they were not used for agriculture due to the fertility of the A horizon).