He also found marks inside the flaps' front edges that he interpreted as internal channels connecting the gills to the interior of the body, much as Whittington interpreted the mark along the proboscis as an internal channel.[5] Zhang and Briggs (2007) however, interpreted all flaps have posterior spacing where the gill blades attached.[5] Budd and Daley (2011) reject the reconstruction by Zhang & Briggs, showing the flaps have complete posteroir edges as previous reconstructions.