The wolf-like canids are a group of large carnivores that are genetically closely related because they all have 78 chromosomes. The group includes genus Canis, Cuon, and Lycaon. The members are the dog (C. lupus familiaris), gray wolf (C. lupus), coyote (C. latrans), golden jackal (C. aureus), Ethiopian wolf (C. simensis), black-backed jackal (C. mesomelas), side-striped jackal (C. adustus), dhole (Cuon alpinus), and African wild dog (Lycaon pictus).[6] The latest recognized member is the African golden wolf (C. anthus), which was once thought to be an African branch of the golden jackal.[4] As they possess 78 chromosomes, all members of the genus Canis are karyologically indistinguishable from each other, and from the dhole and the African hunting dog.[7][8] The two African jackals are shown to be the most basal members of this clade, indicating the clade's origin from Africa.[3] Canis arnensis arrived in Mediterranean Europe 1.9 million years ago and is probably the ancestor of modern jackals.[9]