Clover and Jotter made plant lists and collected specimens throughout the trip, although the rigors of the journey—especially lack of space and difficulty keeping specimens dry—meant that they ended up with fewer specimens than they had hoped.[11] They described the canyon as having five plant zones, from the moist sand along the river's edge up to higher zones with shrubs and trees.[12] Most of what they found were typical riparian species, with a major exception being tamarisk, a non-native species that they saw in a few locations.[13] They found very little snakeweed, which has since become common throughout the canyon. Their survey remains the only comprehensive one done on the river's riparian species in the era before Glen Canyon Dam was built.[12] After the trip, Clover and Jotter published the botanical findings of the trip in a 1944 issue of American Midland Naturalist.