It occupied Delphi from 290 BC and gained territory steadily until, by the end of the 3rd century BC, it controlled the whole of central Greece outside Attica and Boeotia. At its peak, the league's territory included Locris, Malis, Dolopes, part of Thessaly, Phocis, and Acarnania. In the latter part of its power, certain Greek city-states joined the Aetolian League such as the Arcadian cities of Mantineia, Tegea, Figaleia and Kydonia on Crete.[2]