The forest floor serves as a bridge between the above ground living vegetation and the soil, and it is a crucial component in nutrient transfer through the biogeochemical cycle. Much of the energy and carbon fixed by forests is periodically added to the forest floor through litterfall, and a substantial portion of the nutrient requirements of forest ecosystems is supplied by decomposition of organic matter in the forest floor and soil surface. The sustained productivity of forests is closely linked with the decomposition of shed plant parts, particularly the nutrient-rich foliage.