In the 2014 study, of the three successfully generated SNP profiles of Neolithic Starčevo culture samples from Vinkovci, two belonged to Y-DNA haplogroup G2a-P15 and one to I2a1-P37.2, which could indicate G2a as potential representatives of the spread of farming from the Near East to Europe, while I2a as Mesolithic substratum in Europe.[1] In the 2018 study, 10 out of 17 samples from Croatia had a successful Y-DNA sequencing; two Croatia Cardial Neolithic (6005-5786 BCE) samples from Zemunica Cave belonged to C1a2 and E1b1b1a1b1, Early-Neolithic Starčevo culture (5837-5659 BCE) from Beli Manastir-Popova zemlja to C, Early-Neolithic Croatia Impressa culture (5641-5560 BCE) from Kargadur to G2a2a1, two Middle-Neolithic Sopot culture (5207-4546 BCE) samples from Osijek to G2a2a1 and J2a1, Late-Neolithic Sopot (4790-4558 BCE) from Beli Manastir-Popova zemlja to I, two Vučedol culture (2884-2582 BCE) samples from Beli Manastir-Popova zemlja and Vučedol Tell to R1b1a1a2a2 and G2a2a1a2a, and the Early-Middle Bronze Age (1631-1521 BCE) sample from Veliki Vanik belonged to J2b2a.[2]