In a seminal study, Stephens et al.[30] demonstrated this inter-brain link through an fMRI analysis of speakers and listeners. Using the speaker's spatial and temporal neural responses to model the listener's responses during natural verbal communication, they found that brain activity synchronized in dyads in both a delayed and anticipatory manner, but this synchrony failed to occur when subjects did not communicate (e.g., speaking in a language the listener does not understand). Greater synchrony across brains, especially in the predictive anticipatory responses, indicated better scores on comprehension measures.