From August 1940, a flotilla of 27 Italian submarines operated from the BETASOM base in Bordeaux to attack Allied shipping in the Atlantic, initially under the command of Rear Admiral Angelo Parona, then of Rear Admiral Romolo Polacchini and finally of Ship-of-the-Line Captain Enzo Grossi. The Italian submarines had been designed to operate in a different way than U-boats, and they had a number of flaws that needed to be corrected (for example huge conning towers, slow speed when surfaced, lack of modern torpedo fire control), which meant that they were ill-suited for convoy attacks, and performed better when hunting down isolated merchantmen on distant seas, taking advantage of their superior range and living standards. While initial operation met with little success (only 65,343 GRT sunk between August and December 1940), the situation improved gradually over time, and up to August 1943 the 32 Italian submarines that operated there sank 109 ships of 593,864 tons,[31][32][page needed] for 17 subs lost in return, giving them a subs-lost-to-tonnage sunk ratio similar to Germany's in the same period, and higher overall.[6] The Italians were also successful with their use of "human torpedo" chariots, disabling several British ships in Gibraltar.