Passage: The scapula (shoulder blade) was fused with the coracoid, forming a single bone without any visible demarcation between the two original elements. The C. nasicornis holotype was found with an articulated left fore limb including an incomplete manus (hand). Although disarticulated during preparation, a cast had been made of the fossil beforehand to document the original relative positions of the bones. Carpal bones were not known from any specimen, leading some authors to suggest that they were lost in the genus. In a 2016 paper, Matthew Carrano and Jonah Choiniere suggested that one or more cartilaginous (not bony) carpals were probably present, as indicated by a gap present between the forearm bones and the metacarpals, as well as by the surface texture within this gap seen in the cast. In contrast to most more-derived theropods, which showed only three digits on each manus (digits I–III), Ceratosaurus retained four digits, with digit IV reduced in size. The first and fourth metacarpals were short, while the second was slightly longer than the third. The metacarpus and especially the first phalanges were proportionally very short, unlike in most other basal theropods. Only the first phalanges of digits II, III, and IV are preserved in the holotype; the total number of phalanges and unguals (claw bones) is unknown. The anatomy of metacarpal I indicates that phalanges had originally been present on this digit, as well. The pes (foot) consisted of three weight-bearing digits, numbered II–IV. Digit I, which in theropods is usually reduced to a dewclaw that does not touch the ground, is not preserved in the holotype. Marsh, in his original 1884 description, assumed that this digit was lost in Ceratosaurus, but Charles Gilmore, in his 1920 monograph, noted an attachment area on the second metatarsal demonstrating the presence of this digit.Uniquely among theropods, Ceratosaurus possessed small, elongated, and irregularly formed osteoderms (skin bones) along the midline of its body. Such osteoderms have been found...

Question: Ceratosaurus had two metacarpals that were considered short, which were they?
first and fourth