So protein complex two, also known as Succinate reductase, contains the Succinate dehydrogenase enzyme that is used by the citric acid cycle to transform Succinate into fumerate in the process forming Fadh two. Now, one important distinction between protein complex two and the other protein complexes is protein complex two is not a proton pump. It does not actually move the protons from the matrix side to the intermembrane side only protein complex two. And as we'll see in just a moment, protein complex three and four are actually proton pumps and are used to generate electrochemical gradients for protons. So let's move on to complex three. So complex three is shown here, and this complex is also known as cytochrome C oxidoreductase or Q cytochrome C oxidoreductase and sometimes known as cytochrome Reductase.