So even though at the time when mendel made the discovery, we knew what gametes were and we knew that gametes must fertilize to form the Zygote. We knew nothing about how Meiosis takes place and so we knew nothing about how the gametes are actually formed. So although this was actually correct, it wasn't exactly correct. Because we knew nothing about meiosis. So now let's actually try to combine the concept of meiosis with the principle of segregation. So basically, nowadays we know that segregation is a direct result of the separation of the homologous chromosomes.