There are several causes:
1) Physical vibrations during the printing process, which can be caused by a poorly calibrated printer, unstable printing environment, or a machine in need of maintenance.
2) The corners of the print lifting off the bed due to uneven cooling, which can be caused by a lack of bed adhesion, an overheated bed, or an incorrectly calibrated extruder temperature.
3) Insufficient adhesion of the first layer to the print bed, which can be caused by an unclean bed, poor bed levelling, or incorrect bed temperature.
There might be and some other reasons. Let me know if I can help with something else.
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