title: Pterostichinae
text: # Pterostichinae
  
  Pterostichinae Bonelli, 1810 è una sottofamiglia di coleotteri carabidi diffusa in tutto il mondo.
  
  ## Descrizione
  
  Gli Pterostichinae sono coleotteri di medie o piccole dimensioni, forma ovale e colore scuro, spesso metallico. Mancano quasi completamente specie dotate di decorazioni a colori contrastanti. I caratteri che permettono di distinguerli dagli altri carabidi sono: le tibie anteriori con una incavatura all'apice, le elitre a curvatura regolare, le antenne pubescenti dal 4° articolo e gli occhi sormontati da due setole. Quest'ultima caratteristica permette di riconoscerli facilmente dagli Harpalinae, con cui possono venire confusi.
  
  ## Sistematica
  
  La sottofamiglia è suddivisa in otto tribù che comprendono i seguenti generi:
  
  * Tribù Abacetini Chaudoir, 1872
     * Abacetus Dejean, 1828
     * Abacidus LeConte, 1873
     * Aristopus Laferte-Senectere, 1853
     * Celioinkosa Straneo, 1951
     * Chlaeminus Motschulsky, 1865
     * Colpodichius Straneo, 1952
     * Cosmodiscus Sloane, 1907
     * Cyrtomoscelis Chaudoir, 1874
     * Distrigidius Jeannel, 1948
     * Ecnomolaus Bates, 1892
     * Haptoderidius Straneo, 1975
     * Haptoderodes Straneo, 1986
     * Holconotus Schmidt-Goebel, 1846
     * Inkosa Peringuey, 1926
     * Mateuellus Deuve, 1990
     * Metabacetus Bates, 1892
     * Metaxellus Straneo, 1960
     * Metaxys Chaudoir, 1857
     * Novillidius Straneo, 1941
     * Oodinkosa Straneo, 1939
     * Ophonichius Straneo, 1942
     * Pediomorphus Chaudoir, 1878
     * Pioprosopus Tschitscherine, 1899
     * Pollicobius Vinson, 1939
     * Prostalomus Basilewsky, 1950
     * Pseudabacetus Burgeon, 1935
     * Pterostillichus Straneo, 1949
     * Rhagadillius Straneo, 1951
     * Tiferonia Darlington, 1962
     * Trachelocyphoides Alluaud, 1934
     * Trachelocyphus Tschitscherine, 1900
  * Tribù Chaetodactylini Tschitscherine, 1903
     * Chaetodactyla Tschitscherine, 1897
  * Tribù Cnemalobini Germain, 1911
     * Cnemalobus Guerin-Meneville, 1838
  * Tribù Cratocerini Lacordaire, 1854
     * Sottotribù Catapieseina Bates, 1882
        * Catapiesis Solier, 1835
        * Homalomorpha Brulle, 1835
     * Sottotribù Cratocerina sensu stricto
        * Brachidius Chaudoir, 1852
        * Cratocerus Dejean, 1829
        * Oxyglychus Straneo, 1938
     * Sottotribù Drimostomatina Chaudoir, 1872
        * Abacaelostus Straneo, 1952
        * Andrewesinulus Straneo, 1938
        * Apsidocnemus Alluaud, 1936
        * Barylaus Liebherr, 1985
        * Caecocaelus Straneo, 1949
        * Caelostomus W.S. MacLeay, 1825
        * Camptogenys Tschitscherine, 1899
        * Capabatus Csiki, 1930
        * Crenulostrigus Straneo, 1942
        * Cyrtolaus Bates, 1882
        * Dactyleurys Tschitscherine, 1899
        * Dactylinius Straneo, 1941
        * Diachipteryx Alluaud, 1926
        * Diceromerus Chaudoir, 1872
        * Dromistomus Jeannel, 1948
        * Drymonaxus Straneo, 1941
        * Eurystomis Chaudoir, 1878
        * Feostoma Straneo, 1941
        * Hemitelestus Alluaud, 1895
        * Hoplizomenus Chaudoir, 1872
        * Leleuporites Straneo, 1960
        * Madapelmus Dajoz, 1985
        * Monodryxus Straneo, 1942
        * Pachycaecus Straneo, 1971
        * Pachyroxochus Straneo, 1942
        * Platyxythrius Straneo, 1942
        * Stegazopteryx Will, 2004
        * Stomonaxellus Tschitscherine, 1901
        * Strigomerodes Straneo, 1939
        * Strigomerus Chaudoir, 1872
        * Trichillinus Straneo, 1938
  * Tribù Microcheilini Jeannel, 1948
     * Microcheila Brulle, 1834
  * Tribù Morionini Brulle, 1835
     * Buderes Murray, 1857
     * Hyperectenus Alluaud, 1935
     * Hyperion Castelnau, 1834
     * Megamorio Chaudoir, 1880
     * Morion Latreille, 1810
     * Morionidius Chaudoir, 1880
     * Moriosomus Motschulsky, 1864
     * Platynodes Westwood, 1846
     * Stereostoma Murray, 1857
  * Tribù Pterostichini Bonelli, 1810
     * Sottotribù Euchroina Chaudoir, 1874
        * Abacillius Straneo, 1949
        * Abacillodes Straneo, 1988
        * Abaris Dejean, 1831
        * Apsaustodon Tschitscherine, 1901
        * Argutoridius Chaudoir, 1876
        * Blennidus Motschulsky, 1865
        * Blenniventer Straneo, 1986
        * Bothynoproctus Tschitscherine, 1900
        * Cephalostichus Straneo, 1977
        * Cynthidia Chaudoir, 1873
        * Euchroa Brulle, 1834
        * Eumara Tschitscherine, 1901
        * Gastrogmus Sloane, 1915
        * Haplobothynus Tschitscherine, 1901
        * Hybothecus Chaudoir, 1874
        * Litarthrum Sloane, 1915
        * Lobobrachus Sharp, 1885
        * Marsyas Putzeys, 1845
        * Meropalpus Tschitscherine, 1900
        * Microcephalus Dejean, 1828
        * Mireius Straneo, 1984
        * Nelidus Chaudoir, 1878
        * Neotalus Will, 2002
        * Oribazus Chaudoir, 1874
        * Pachythecus Chaudoir, 1874
        * Phaenaulax Tschitscherine, 1898
        * Prosopogmus Chaudoir, 1865
        * Pseudabarys Chaudoir, 1873
        * Setalimorphus Sloane, 1895
        * Setalis Castelnau, 1867
        * Simodontus Chaudoir, 1843
        * Trirammatus Chaudoir, 1838
     * Sottotribù Loxandrina Erwin & Sims, 1984
        * Adrimus Bates, 1871
        * Cerabilia Castelnau, 1867
        * Feronista Moore, 1965
        * Haploferonia Darlington, 1962
        * Homalonesiota Maindron, 1908
        * Loxandrus LeConte, 1852
        * Metoncidus Bates, 1871
        * Nebrioferonia Straneo, 1939
        * Oxycrepis Reiche, 1843
        * Stolonis Motschulsky, 1866
        * Zeodera Castelnau, 1867
     * Sottotribù Metiina Straneo, 1951
        * Abropus Waterhouse, 1842
        * Antarctiola Straneo, 1977
        * Chaetauchenium Tschitscherine, 1900
        * Feroniola Tschitscherine, 1900
        * Kuschelinus Straneo, 1963
        * Metius Curtis, 1839
        * Parhypates Motschulsky, 1866
     * Sottotribù Molopina Bonelli, 1810
        * Abacops Tschitscherine, 1902
        * Abax Bonelli, 1810
        * Aristochroa Tschitscherine, 1898
        * Aristochroodes Marcillac, 1993
        * Camptoscelis Dejean, 1828
        * Cedrorum Borges & Serrano, 1993
        * Chalcochrous Chaudoir, 1838
        * Eucamptognathus Chaudoir, 1837
        * Eudromus Klug, 1835
        * Henrotius Jeannel, 1953
        * Lesticus Dejean, 1828
        * Molopidius Jeannel, 1942
        * Molopinus Jeannel, 1948
        * Molops Bonelli, 1810
        * Myas Sturm, 1826: 171
        * Nirmala Andrewes, 1930
        * Orthomus Chaudoir, 1838
        * Oscadytes Lagar Mascaro, 1975
        * Pedius Motschulsky, 1850
        * Percus Bonelli, 1810
        * Peyrieraselus Deuve, 1981
        * Sthenocranion Tschitscherine, 1899
        * Styracoderus Chaudoir, 1874
        * Teratotarsa Tschitscherine, 1893
        * Trigonognatha Motschulsky, 1858
        * Tropidocerus Chaudoir, 1878
        * Zariquieya Jeannel, 1924
     * Sottotribù Pterostichina sensu stricto
        * Abacoleptus Fauvel, 1903
        * Abacomorphus Chaudoir, 1878
        * Abacophrastus Will, 2011
        * Acanthoferonia Moore, 1965
        * Allotriopus Bates, 1882
        * Analoma Darlington, 1971
        * Aulacopodus Britton, 1940
        * Basilewskya Straneo, 1948
        * Castelnaudia Tschitscherine, 1890
        * Catadromus W.S. Macleay, 1825
        * Conchitella Moore, 1962
        * Cophosomorpha Tschitscherine, 1890
        * Cratoferonia Tschitscherine, 1902
        * Cratogaster Blanchard, 1843
        * Cyclotrachelus Chaudoir, 1838
        * Cyrtoderus Hope, 1842
        * Darodilia Castelnau, 1867
        * Delinius Westwood, 1864
        * Euryabax Fauvel, 1903
        * Euryaptus Bates, 1892
        * Gastrellarius Casey, 1918
        * Gourlayia Britton, 1964
        * Harpostomus Chaudoir, 1856
        * Holcaspis Chaudoir, 1865
        * Leiradira Castelnau, 1867
        * Liopasa Tschitscherine, 1901
        * Lophoglossus LeConte, 1852
        * Loxodactylus Chaudoir, 1865
        * Loxogenius Sloane, 1907
        * Mecynognathus W.J.MacLeay, 1873
        * Megadromus Motschulsky, 1865
        * Neoferonia Britton, 1940
        * Nesites Andrewes, 1939
        * Notabax Moore, 1976
        * Notolestus Sloane, 1894
        * Notonomus Chaudoir, 1862
        * Nurus Motschulsky, 1866
        * Oberthueria Vuillet, 1911
        * Ogmophora Tschitscherine, 1896
        * Onawea Johns, 2007
        * Pachymorphus Chaudoir, 1838
        * Paniestichus Will, 2011
        * Paranurus Tschitscherine, 1901
        * Pareuryaptus Dubault, Lassalle & Roux, 2008
        * Percolaus Bates, 1882
        * Piesmus LeConte, 1852
        * Platycaelus Blanchard, 1843
        * Platysmodes Fauvel, 1903
        * Plocamostethus Britton, 1940
        * Poecilinus Jeannel, 1948
        * Poecilus Bonelli, 1810
        * Psegmatopterus Chaudoir, 1878
        * Pseudoceneus Tschitscherine, 1891
        * Pterostichus Bonelli, 1810
        * Rhabdotus Chaudoir, 1865
        * Rhytiferonia Darlington, 1962
        * Rhytisternus Chaudoir, 1865
        * Sarticus Motschulsky, 1865
        * Secatophus Castelnau, 1867
        * Setalidius Chaudoir, 1878
        * Speluncarius Reitter, 1886
        * Speomolops Patrizi, 1955
        * Sphodrosomus Perroud & Montrouzier, 1864
        * Stereocerus Kirby, 1837
        * Steropanus Fairmaire, 1888
        * Stomis Clairville, 1806
        * Straneostichus Sciaky, 1994
        * Taphoxomimus Straneo, 1975
        * Tapinopterus Schaum, 1858
        * Teropha Castelnau, 1867
        * Trichosternus Chaudoir, 1865
        * Trigonotoma Dejean, 1828
        * Wahlbergiana Bousquet, 2002
        * Xenion Tschitscherine, 1902
        * Zeopoecilus Sharp, 1886
  * Tribù Zabrini Bonelli, 1810
     * Sottotribù Amarina C. Zimmermann, 1831
        * Amara Bonelli, 1810
        * Pseudamara Lindroth, 1968
     * Sottotribù Zabrina Bonelli, 1810
        * Zabrus Clairville, 1806
  