title: Aphodius
text: # Aphodius
  
  Aphodius is a genus of beetles in the family Scarabaeidae. In most species both the adults and larvae are coprophagous (dung feeding) although some species have herbivorous or saprophagous larvae. Aphodius species typically dominate dung beetle communities in north temperate ecosystems. Most species are functionally classified as endocoprids, also known as dwellers, because the larvae live and feed within the dung pat itself.
  
  ## Species
  
  These 44 species belong to the genus Aphodius, including 21 extinct species.
  
  * Aphodius beloni Mulsant & Godart, 1879
  * Aphodius calichromus Balthasar, 1932
  * Aphodius cardinalis Reitter, 1892
  * Aphodius clypeatus Fischer von Waldheim, 1821
  * Aphodius coniugatus (Panzer, 1795)
  * Aphodius corallifer Koshantschikov, 1913
  * Aphodius crux Wiedemann, 1823
  * Aphodius elegans Allibert, 1847
  * Aphodius fasciger Harold, 1881
  * Aphodius fimetarius (Linnaeus, 1758)
  * Aphodius foetidus (Herbst, 1783)
  * Aphodius guangdongensis Maté, 2008
  * Aphodius irregularis Westwood, 1839
  * Aphodius marginatus Fischer von Waldheim, 1842
  * Aphodius micros Walker, 1871
  * Aphodius minatorius Péringuey, 1908
  * Aphodius pedellus (Degeer, 1774)
  * Aphodius pedrosi Wollaston, 1854
  * Aphodius plasoni Käufel, 1914
  * Aphodius reginae Král, 1997
  * Aphodius spadix Schmidt, 1916
  * Aphodius swaneticus Reitter, 1892
  * Aphodius thoracicus Fischer von Waldheim, 1842
  * † Aphodius aboriginalis Wickham, 1912
  * † Aphodius anteactus Krell, 2000
  * † Aphodius bosniaskii Handlirsch, 1907
  * † Aphodius brevipennis Heer, 1862
  * † Aphodius charauxi Piton, 1940
  * † Aphodius cretaceous Nikolajev, 2008
  * † Aphodius florissantensis Wickham, 1911
  * † Aphodius granarioides Wickham, 1913
  * † Aphodius helvolus Statz, 1952
  * † Aphodius inundatus Wickham, 1914
  * † Aphodius krantzi Heyden & Heyden, 1866
  * † Aphodius laminicola Wickham, 1910
  * † Aphodius mediaevus Wickham, 1914
  * † Aphodius meyeri Heer, 1847
  * † Aphodius praeemptor Wickham, 1913
  * † Aphodius precursor Horn, 1876
  * † Aphodius schlickumi Statz, 1952
  * † Aphodius senex Wickham, 1914
  * † Aphodius shoshonis Wickham, 1912
  * † Aphodius theobaldi Krell, 2000
  * † Aphodius vectis Krell, 2019
  
  The species of the following genera were formerly classified in Aphodius:
  
      Acrossus Mulsant, 1842
      Agoliinus Schmidt, 1913
      Alloblackburneus Bordat, 2009
      Ballucus Gordon & Skelley, 2007
      Blackburneus Schmidt, 1913
      Calamosternus Motschulsky, 1859
      Caligodorus Gordon & Skelley, 2007
      Cephalocyclus Dellacasa et al., 1998
      Chilothorax Motschulsky, 1859
      Cinacanthus Schmidt, 1913
      Coelotrachelus Schmidt, 1913
      Colobopterus Mulsant, 1842
      Cryptoscatomaseter Gordon & Skelley, 2007
      Dellacasiellus Gordon & Skelley, 2007
      Dialytodius Gordon & Skelley, 2007
      Drepanocanthoides Schmidt, 1913
      Eupleurus Mulsant, 1842
      Flaviellus Gordon & Skelley, 2007
      Geomyphilus Gordon & Skelley, 2007
      Haroldiellus Gordon & Skelley, 2007
      Hornosus Dellacasa, Dellacasa & Gordon, 2015
      Irrasinus Gordon & Skelley, 2007
      Labarrus Mulsant & Rey, 1869
      Lechorodius Gordon & Skelley, 2007
      Liothorax Motschulsky, 1859
      Luxolinus Gordon & Skelley, 2007
      Maculaphodius Gordon & Skelley, 2007
      Melinopterus Mulsant, 1842
      Mendidius Harold, 1868
      Merogyrus Gordon & Skelley, 2007
      Neotrichonotulus Dellacasa et al., 2004
      Nialaphodius Kolbe, 1908
      Orodaliscoides Schmidt, 1913
      Oscarinus Gordon & Skelley, 2007
      Otophorus Mulsant, 1842
      Oxyomoides Dellacasa et al., 2016
      Pardalosus Gordon & Skelley, 2007
      Phaeaphodius Reitter, 1892
      Planolinellus Dellacasa & Dellacasa, 2005
      Planolinoides Dellacasa & Dellacasa, 2005
      Planolinus Mulsant & Rey, 1869
      Pseudagolius Schmidt, 1913
      Rugaphodius Gordon & Skelley, 2007
      Scabrostomus Gordon & Skelley, 2007
      Schaefferellus Gordon & Skelley, 2007
      Setodius Gordon & Skelley, 2007
      Stenotothorax Schmidt, 1913
      Strigodius Gordon & Skelley, 2007
      Tetraclipeoides Schmidt, 1913
      Teuchestes Mulsant, 1842
      Trichaphodioides Paulian, 1942
      Trichonotulus Bedel, 1911
  