title: Columella (botany)
text: # Columella (botany)
  
  Columella (in plants) is an axis of sterile tissue which passes through the center of the spore-case of mosses. In fungi it refers to a centrally vacuolated part of a hypha, bearing spores. The word finds analogous usage in myxomycetes.
  
  The term columella is also used to refer to story 1 to story 4 (S1 – S4) cells in the root cap, located apically of the quiescent centre.
  
  