title: Arthropods of the Cambrian Period
text: # Arthropods of the Cambrian Period
  
  This list contains extinct arthropod genera from the Cambrian Period of the Paleozoic Era.
  
  ## Artiopoda
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Acanthomeridion | 1989 | Chengjiang Biota |
  
  ## Dinocaridida
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Anomalocaris | 1892 | Emu Bay Shale Maotianshan Shales Burgess Shale |
  | Cucumericrus | 1995 | Chengjiang Biota |
  | Paranomalocaris | 2013 | Wulongqing Formation |
  | Cambroraster | 2019 | Burgess Shale Chengjiang Biota |
  | Hurdia | 1992 | Burgess Shale |
  | Peytoia | 1911 | Burgess Shale Zawiszyn Formation |
  | Ursulinacaris | 2019 | Carrara Formation |
  | Titanokorys | 2018 | Burgess Shale |
  | Pahvantia | 1981 | Wheeler Shale Marjum Formation |
  | Amplectobelua | 1995 | Chengjiang Burgess Shale |
  | Houcaris | 2021 | |
  | Lyrarapax | 2014 | |
  | Caryosyntrips | 2010 | Burgess Shale Wheeler Shale Marjum Formation Valdemiedes Formation |
  | Tamisiocaris | 2010 | Buen Formation Sirius Passet |
  | Opabinia | 1912 | Burgess Shale |
  | Utaurora | 2022 | Wheeler Formation |
  | Kerygmachela | 1993 | Buen Formation Sirius Passet |
  | Laminacaris | 2018 | |
  | Pambdelurion | 1997 | Sirius Passet |
  
  ## Malacostraca
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Hymenocaris | 1853 | Burgess Shale |
  | Odaraia | 1912 | Burgess Shale Greater Phyllopod Bed |
  
  ## Mandibulata
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Branchiocaris | 1929 | Wheeler Shale Burgess Shale |
  | Tokummia | 2017 | Burgess Shale |
  | Canadaspis | 1978 | Burgess Shale Pioche Shale Chengjiang Biota |
  | Waptia | 1912 | Burgess Shale |
  | Pectocaris | 1999 | Maotianshan Shales Burgess Shale |
  | Clypecaris | 1999 | Chengjiang Biota |
  | Pauloterminus | 2002 | Sirius Passet |
  | Perspicaris | 1977 | Burgess Shale |
  | Nereocaris | 2012 | Burgess Shale |
  | Plenocaris | 1912 | Burgess Shale Chengjiang Biota |
  | Ercaicunia | 1999 | Maotianshan Shales |
  | Fibulacaris | 2019 | Burgess Shale |
  | Synophalos | 2009 | Maotianshan Shales |
  
  ## Marrellomorpha
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Marrella | 1912 | Burgess Shale Kaili Formation Balang Formation |
  | Skania | 1931 | Maotianshan Shales Burgess Shale |
  | Primicaris | 2003 | Chengjiang Biota |
  
  ## Maxillopoda
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Heymonsicambria | 1994 | Orsten Fauna |
  | Skara | 1983 | Orsten Fauna |
  | Goticaris | 1990 | Orsten Fauna |
  
  ## Megacheira
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Actaeus | 1970 | Burgess Shale |
  | Alalcomenaeus | 1970 | Burgess Shale Chengjiang Biota Wheeler Shale |
  | Leanchoilia | 1912 | Burgess Shale Chengjiang Biota Spence Shale |
  | Fortiforceps | 1997 | Maotianshan Shales |
  | Yawunik | 2015 | Burgess Shale |
  | Parapeytoia | 1995 | Maotianshan Shales |
  | Jianfengia | 1987 | Maotianshan Shales |
  | Haikoucaris | 2004 | Chengjiang Biota |
  | Kanoshoia | 2020 | Wheeler Formation |
  | Yohoia | 1912 | Burgess Shale |
  
  ## Ostracoda
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Carnarvonia | 1912 | Burgess Shale |
  
  ## Trilobita
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Bathyuriscus | 1873 | Burgess Shale |
  | Olenoides | 1877 | Burgess Shale Marjum Formation |
  | Hanburia | 1916 | Burgess Shale Stephen Formation |
  | Xiuqiella | 1974 | |
  | Acontheus | 1851 | |
  | Hemirhodon | 1937 | |
  | Thoracocare | 1974 | Spence Shale Formation |
  | Abakania | 1973 | Sanashtykgol Formation |
  | Zacanthoides | 1888 | Burgess Shale |
  | Changaspis | 1961 | |
  | Ogygopsis | 1889 | Burgess Shale |
  | Oryctocephalus | 1886 | Burgess Shale |
  | Chancia | 1924 | Burgess Shale |
  | Ehmaniella | 1951 | Burgess Shale |
  | Elrathia | 1924 | Wheeler Formation Burgess Shale |
  | Ellipsocephalus | 1883 | Słowiec Sandstone Formation |
  | Phantaspis | 2020 | Mantou Formation |
  | Aojia | 1935 | |
  | Bailiaspis | 1936 | Whitesands Bay Formation Jbel Lmgaysmat Formation Manuels River Formation Maentwrog Formations |
  | Solenoparia | 1935 | |
  | Westergaardites | 1937 | |
  | Yunnanocephalus | 1936 | Chengjiang Biota |
  | Asaphiscus | 1873 | |
  | Olenus | 1827 | |
  | Quitacetra | 1967 | |
  | Ejinaspis | 1986 | |
  | Asthenopsis | 1939 | |
  | Agraulos | 1847 | Menevia Formation Jince Formation Manuels River Formation |
  | Coosella | 1936 | |
  | Norwoodia | 1916 | |
  | Tricrepicephalus | 1935 | Chefu Formation Deadwood Formation Little Elk Creek |
  | Meniscopsia | 2011 | |
  | Acrocephalella | 1963 | |
  | Cambroproteus | 2015 | |
  | Cedaria | 1924 | Weeks Formation |
  | Conocoryphe | 1847 | |
  | Galeaspis | 1983 | |
  | Genevievella | 1936 | Mungerbar Formation |
  | Meteoraspis | 1890 | Limestone Formation |
  | Validaspis | 1991 | |
  | Palaeolenus | 1912 | |
  | Ptychoparia | 1847 | |
  | Kootenia | 1939 | Burgess Shale Emu Bay Shale |
  | Misszhouia | 1985 | Chengjiang Biota |
  | Naraoia | 1912 | Burgess Shale Stephen Formation Chengjiang biota Formation of Southern Ontario Balang Formation Emu Bay Shale |
  | Estaingia | 1964 | |
  | Bathynotus | 1860 | |
  | Bajanaspis | 1978 | Edreisk Formation |
  | Kleptothule | 1995 | Sirius Passet |
  | Acadoparadoxides | 1957 | |
  | Paradoxides | 1822 | Duda Formation |
  | Eoredlichia | 1950 | Chengjiang Biota |
  | Profallotaspis | 2002 | |
  | Lochmanolenellus | 1952 | Poleta Formation |
  | Olenellus | 1861 | Pioche Shale Carrara Formation Kinzers Formation |
  | Archaeaspis | 1965 | Campito Formation |
  | Parapaokannia | 1980 | |
  | Plutonides | 1985 | Purley Shale Formation |
  | Redlichia | 1902 | |
  | Repinaella | 1965 | Pestrotsvet Formation |
  | Ushbaspis | 1965 | Shabakty Formation |
  | Balcoracania | 1970 | Balcoracana Creek |
  | Emuella | 1970 | Emu Bay |
  | Megapharanaspis | 2006 | |
  | Wutingaspis | 1944 | |
  | Mesolenellus | 1993 | Sofiekammen Formation |
  | Olenelloides | 1894 | |
  | Biceratops | 1971 | Pioche Shale |
  | Bradyfallotaspis | 1972 | |
  | Eopeachella | 2009 | Delamar Mountains |
  | Laudonia | 1956 | |
  | Mummaspis | 1992 | Mural Formation |
  | Nephrolenellus | 1993 | Pyramid Shale Member Carrara Formation |
  | Nevadella | 1936 | Poleta Formation Sekwi Formation |
  | Paranephrolenellus | 2007 | |
  | Wanneria | 1910 | Kinzers Formation |
  | Bristolia | 1956 | Carrara Formation |
  | Acadoparadoxides | 1957 | |
  | Emigrantia | 1910 | |
  | Bolbolenellus | 1993 | |
  | Buenellus | 1988 | Sirius Passet |
  | Callavia | 1890 | Brigus Formation |
  | Chengjiangaspis | 1980 | Maotianshan Shales |
  | Cirquella | 1993 | |
  | Daguinaspis | 1950 | Amouslek Formation |
  | Fallotaspis | 1953 | |
  | Holmia | 1890 | Sparagmite Formation |
  | Hydrocephalus | 1846 | |
  | Fritzolenellus | 1998 | |
  | Mesonacis | 1859 | Forteau Formation |
  | Xystridura | 1936 | |
  | Peachella | 1910 | |
  | Richterops | 1954 | |
  | Archaeagnostus | 1939 | |
  | Agnostus | 1822 | Marjum Biota Linyi Lagerstätte Orsten Fauna |
  | Itagnostus | 1979 | |
  | Peronopsis | 1847 | |
  | Acmarhachis | 1938 | |
  | Acadagnostus | 1939 | |
  | Agnostotes | 1963 | |
  | Lejopyge | 1828 | |
  | Glyptagnostus | 1936 | |
  | Egyngolia | 1980 | |
  | Jinghediscus | 1985 | |
  | Pagetia | 1916 | Burgess Shale |
  | Acimetopus | 1966 | |
  | Pentagnostus | 1940 | |
  | Calodiscus | 1935 | Schodack Formation |
  | Mallagnostus | 1935 | |
  | Runcinodiscus | 1976 | |
  | Serrodiscus | 1941 | Brigus Formation |
  | Weymouthia | 1913 | |
  | Analox | 1966 | |
  | Litometopus | 1966 | |
  | Meniscuchus | 1975 | |
  | Ninadiscus | 1980 | Egyngolskaya Suite |
  | Oodiscus | 1966 | |
  | Semadiscus | 1978 | |
  | Stigmadiscus | 1966 | |
  | Parapagetia | 1964 | |
  | Tannudiscus | 1959 | Purley Shale Formation |
  | Chelediscus | 1966 | |
  | Bathydiscus | 1966 | |
  | Cobboldites | 1943 | |
  | Acidiscus | 1966 | |
  | Buenaspis | 1999 | Sirius Passet |
  | Condylopyge | 1847 | |
  | Delgadella | 1908 | Tyuser Formation |
  | Dicerodiscus | 1954 | |
  | Pleuroctenium | 1847 | |
  | Ptychagnostus | 1909 | |
  | Bolboparia | 1966 | |
  | Tchernyshevioides | 1975 | |
  | Toragnostus | 1988 | |
  | Tsunyidiscus | 1966 | |
  | Birmanites | 1934 | |
  | Kendallina | 1953 | Bison Creek Formation |
  | Dikelocephalus | 1852 | |
  | Orygmaspis | 1938 | Morgan Creek Member |
  | Cheirurus | 1845 | |
  | Cindarella | 1996 | Chengjiang Biota |
  | Helmetia | 1912 | Burgess Shale Jince Formation |
  | Tegopelte | 1975 | Burgess Shale Kicking Horse Shale |
  | Retifacies | 1989 | Maotianshan Shales |
  | Schmalenseeia | 1903 | |
  | Burlingia | 1908 | |
  | Entomaspis | 1931 | |
  | Neodrepanura | 2006 | Maotianshan Shales |
  
  ## Incertae sedis
  
  | Genus | Year of Discovery | Location of Fossils |
  | - | - | - |
  | Kiisortoqia | 2010 | Sirius Passet |
  | Vladicaris | 1995 | |
  | Wujicaris | 2010 | Orsten Fauna |
  | Protocaris | 1884 | Parker Formation |
  | Burgessia | 1912 | Burgess Shale |
  | Liangshanella | 1997 | Chengjiang Biota Burgess Shale |
  | Kunmingella | 1912 | Maotianshan Shales |
  | Comptaluta | 1968 | Maotianshan Shales |
  | Jiucunella | 1991 | Maotianshan Shales |
  | Kunyangella | 1965 | Maotianshan Shales |
  | Cyclopites | 1951 | |
  | Mongolitubulus | 1997 | |
  | Habelia | 1912 | Burgess Shale |
  | Khankaspis | 1969 | Snegurovka Formation |
  | Isoxys | 1890 | Burgess Shale Chengjiang Biota Sirius Passet |
  | Emeraldella | 1912 | Burgess Shale Wheeler Shale |
  | Molaria | 1912 | Burgess Shale Sirius Passet |
  | Sidneyia | 1911 | Burgess Shale Maotianshan Shales |
  | Bailongia | 2021 | Maotianshan Shales |
  | Luohuilinella | 2012 | Chengjiang Biota |
  | Panlongia | 2006 | Guanshan Fauna |
  | Pygmaclypeatus | 2000 | Chengjiang Biota |
  | Sinoburius | 1991 | Chengjiang Biota |
  | Retifacies | 1989 | Chengjiang Biota |
  | Strabops | 1901 | |
  | Paleomerus | 1956 | |
  | Parapaleomerus | 1999 | Chengjiang Biota |
  | Sanctacaris | 1988 | Burgess Shale |
  | Mollisonia | | Chengjiang Biota Burgess Shale Langston Formation Wheeler Shale |
  | Thelxiope | 1975 | Burgess Shale Wheeler Shale Fezouata Formation Linyi Lagerstätte |
  | Glypharthrus | 1939 | McKay Group Clay Creek |
  | Urokodia | 1989 | Maotianshan Shales |
  | Fuxianhuia | 1987 | Chengjiang Biota |
  | Alacaris | 2018 | |
  | Chengjiangocaris | 1991 | |
  | Guangweicaris | 2007 | |
  | Tuzoia | 1912 | Burgess Shale Emu Bay Shale Rockslide Formation Spence Shale Wheeler Formation Marjum Formation Kinzers Formation |
  | Klausmuelleria | 2003 | Orsten Fauna |
  | Kylinxia | 2020 | Maotianshan Shales |
  | Aaveqaspis | 2009 | Sirius Passet |
  | Worthenella | 1911 | Burgess Shale |
  | Kodymirus | 1965 | Paseky Shale |
  | Mosineia | 2010 | Blackberry Hill |
  | Cambropachycope | 1990 | Orsten Fauna |
  | Bushizheia | 2020 | Chiungchussu |
  | Combinivalvula | 1987 | Chengjiang Biota |
  | Diplopyge | 1999 | Chengjiang Biota |
  | Dongshanocaris | 1999 | Chengjiang Biota |
  | Ercaia | 2001 | Chengjiang Biota |
  | Jianshania | 1999 | Chengjiang Biota |
  | Pseudoiulia | 1998 | Chengjiang Biota |
  | Priscansermarinus | 1981 | Burgess Shale |
  | Kootenichela | 2013 | |
  | Forfexicaris | 1999 | Maotianshan Shales |
  | Kockurus | 1995 | |
  | Occacaris | 1999 | Maotianshan Shales |
  | Sarotrocercus | 1981 | Burgess Shale |
  