title: Orthogonius
text: # Orthogonius
  
  Orthogonius is a genus of beetles in the family Carabidae, containing the following species:
  
  * Orthogonius aberlenci Tian & Deuve, 2016 (Laos)
  * Orthogonius acrogonus (Wiedemann, 1819) (Malaysia, Indonesia, and Borneo)
  * Orthogonius acutangulus Chaudoir, 1878 (Sri Lanka)
  * Orthogonius adoriae Tian & Deuve, 2006 (India)
  * Orthogonius aemulus Péringuey, 1896 (Zimbabwe, Botswana, and Namibia)
  * Orthogonius alternans (Wiedemann, 1823) (Indomalaya)
  * Orthogonius alutaceus Quedenfeldt, 1883 (Africa)
  * Orthogonius andrewesi Emden, 1928 (Philippines)
  * Orthogonius andrewesianus Tian & Deuve, 2006 (Malaysia, Singapore, and Indonesia)
  * Orthogonius angkor Tian & Deuve, 2006 (Cambodia)
  * Orthogonius angkorensis Tian & Deuve, 2008 (Cambodia)
  * Orthogonius angustus Chaudoir, 1872 (Thailand, Vietnam, and Malaysia)
  * Orthogonius annamensis Jedlicka, 1965 (Vietnam)
  * Orthogonius annamicola Tian & Deuve, 2006 (Vietnam)
  * Orthogonius annemariae Tian & Deuve, 2006 (India)
  * Orthogonius assamensis Tian & Deuve, 2006 (India)
  * Orthogonius assamicola Tian & Deuve, 2006 (India)
  * Orthogonius assmuthi Wasmann, 1920 (India)
  * Orthogonius baconi Chaudoir, 1872 (India and Myanmar)
  * Orthogonius baconioides Tian & Sabu, 2009 (India)
  * Orthogonius balthasari Jedlicka, 1935 (Philippines)
  * Orthogonius batesi Tian & Deuve, 2003 (Sri Lanka)
  * Orthogonius batesianus Tian & Deuve, 2007
  * Orthogonius batesioides Tian & Deuve, 2006 (India)
  * Orthogonius baumi Jedlicka, 1955 (Malaysia)
  * Orthogonius becqueti Burgeon, 1937 (Democratic Republic of the Congo and Burundi)
  * Orthogonius bellus Tian & Deuve, 2016 (Vietnam)
  * Orthogonius bicolor Jedlicka, 1965 (Vietnam)
  * Orthogonius borneoensis Tian & Deuve, 2006 (Indonesia and Borneo)
  * Orthogonius brancuccii Tian & Deuve, 2006 (Thailand)
  * Orthogonius brevilabris Kolbe, 1889 (Democratic Republic of the Congo and Angola)
  * Orthogonius brevithorax Dejean, 1825 (Africa)
  * Orthogonius bruggei Tian & Deuve, 2006 (Malaysia)
  * Orthogonius buquetii Chaudoir, 1850 (Sierra Leone and Ivory Coast)
  * Orthogonius burckhardti Tian & Deuve, 2006
  * Orthogonius burmanensis Tian & Deuve, 2006 (Myanmar)
  * Orthogonius caffer Boheman, 1848 (Zimbabwe and South Africa)
  * Orthogonius cambodgensis Tian & Deuve, 2006 (Cambodia)
  * Orthogonius canaraensis Tian & Deuve, 2006 (India)
  * Orthogonius canaraicola Tian & Deuve, 2006 (India)
  * Orthogonius capucinus Boheman, 1848 (Zimbabwe and South Africa)
  * Orthogonius carbonicolor Tian & Deuve, 2006 (Myanmar)
  * Orthogonius carinatus Tian & Deuve, 2013 (Laos)
  * Orthogonius celebesicus Tian & Deuve, 2006 (Indonesia)
  * Orthogonius ceylanicus Tian & Deuve, 2016 (Sri Lanka)
  * Orthogonius chaudoiri Tian & Deuve, 2001 (Vietnam)
  * Orthogonius cheni Tian & Deuve, 2001 (China)
  * Orthogonius chiangensis Tian & Deuve, 2006 (Thailand)
  * Orthogonius clarkii Murray, 1858 (Africa)
  * Orthogonius confusus Tian & Deuve, 2006 (Malaysia)
  * Orthogonius constanti Tian & Deuve, 2006 (Cambodia)
  * Orthogonius constrictus Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius coomani Tian & Deuve, 2006 (Vietnam)
  * Orthogonius coomanioides Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius coracinus Kolbe, 1895 (Uganda, Rwanda, Burundi, and Tanzania)
  * Orthogonius crassicrus Chaudoir, 1872 (Indonesia and Borneo)
  * Orthogonius cruralis Putzeys in Chaudoir, 1872
  * Orthogonius curvatus Tian & Deuve, 2006 (India)
  * Orthogonius cyclothorax Tian & Deuve, 2007 (Singapore)
  * Orthogonius davidi Chaudoir, 1878 (China and Taiwan)
  * Orthogonius deletus Schmidt-Goebel, 1846 (China, India, Southeast Asia)
  * Orthogonius deuvei Tian & Kirschenhofer, 2013 (Vietnam)
  * Orthogonius dispar Bates, 1892 (Myanmar)
  * Orthogonius dohertyi Tian & Deuve, 2006 (Malaysia)
  * Orthogonius dongnanya Tian & Deuve, 2006 (Singapore, Indonesia, and Borneo)
  * Orthogonius doriae Putzeys in Chaudoir, 1872 (Indonesia and Borneo)
  * Orthogonius drescheri Liebke, 1937 (Indonesia)
  * Orthogonius drumonti Tian & Deuve, 2006 (Indonesia)
  * Orthogonius duboisi Tian & Deuve, 2006 (China)
  * Orthogonius duplicatus (Wiedemann, 1819) (Indomalaya)
  * Orthogonius dupuisi Basilewsky, 1948 (Zambia)
  * Orthogonius dupuisianus Tian & Deuve, 2010 (Indonesia)
  * Orthogonius dureli Tian & Deuve, 2005 (Bhutan and India)
  * Orthogonius edentatus Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
  * Orthogonius elisabethanus Burgeon, 1937 (Democratic Republic of the Congo)
  * Orthogonius emarginatus Tian & Deuve, 2007 (Singapore)
  * Orthogonius equimarginalis Tian & Deuve, 2006 (Indonesia)
  * Orthogonius euthyphallus Tian & Deuve, 2013 (Laos and Vietnam)
  * Orthogonius fairmairei Tian & Deuve, 2006 (Singapore)
  * Orthogonius feai Tian & Deuve, 2007
  * Orthogonius femoralis Chaudoir, 1848 (Sri Lanka and India)
  * Orthogonius flavipes Deuve, 2004 (Indonesia)
  * Orthogonius flavus Jedlicka, 1964 (India)
  * Orthogonius foveiclypeus Tian & Deuve, 2004 (Indonesia)
  * Orthogonius fryi Tian & Deuve, 2006 (Myanmar)
  * Orthogonius fugax Chaudoir, 1872 (Sri Lanka)
  * Orthogonius gracililamella Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius gracilipes Tian & Deuve, 2006 (India)
  * Orthogonius gracilis Tian & Deuve, 2006 (India)
  * Orthogonius grootaerti Tian & Deuve, 2006 (Thailand)
  * Orthogonius hageni Oberthür, 1883 (Indonesia)
  * Orthogonius himalaya Tian & Deuve, 2006 (Bhutan and India)
  * Orthogonius himalayicus Tian & Deuve, 2005 (Bhutan and India)
  * Orthogonius hirtulus Tian & Deuve, 2006 (Indonesia)
  * Orthogonius hirtus Chaudoir, 1872 (Malaysia and Indonesia)
  * Orthogonius hopei Gray, 1832 (Indomalaya)
  * Orthogonius huananoides Tian & Deuve, 2006 (Thailand and Vietnam)
  * Orthogonius huananus Tian & Deuve, 2001 (China)
  * Orthogonius hypocrita Chaudoir, 1872 (Indonesia and Philippines)
  * Orthogonius hypocritoides Jedlicka, 1935 (Philippines)
  * Orthogonius impunctipennis Quedenfeldt, 1883 (Democratic Republic of the Congo and Angola)
  * Orthogonius indicus Tian & Deuve, 2006 (India)
  * Orthogonius indophilus Tian & Deuve, 2006 (India)
  * Orthogonius inexpectatus Tian & Deuve, 2006 (India)
  * Orthogonius inops Chaudoir, 1872 (Indonesia)
  * Orthogonius insularis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
  * Orthogonius intermedius Chaudoir, 1872 (Indonesia)
  * Orthogonius javanus Tian & Deuve, 2006 (Taiwan and Indonesia)
  * Orthogonius jianfengling Tian & Deuve, 2001 (China)
  * Orthogonius kalimantanensis Tian & Deuve, 2006 (Indonesia and Borneo)
  * Orthogonius kapiriensis Burgeon, 1937 (Democratic Republic of the Congo)
  * Orthogonius katangensis Burgeon, 1937 (Democratic Republic of the Congo)
  * Orthogonius kickeli Kolbe, 1895 (Namibia)
  * Orthogonius kirirom Tian & Deuve, 2008 (Thailand and Cambodia)
  * Orthogonius klickai Jedlicka, 1935 (Philippines)
  * Orthogonius kubani Tian & Deuve, 2006 (Thailand)
  * Orthogonius kumatai Habu, 1979 (Nepal)
  * Orthogonius lancangjiang Tian & Deuve, 2006 (Laos)
  * Orthogonius lao Tian & Deuve, 2006 (Laos)
  * Orthogonius latreillei Tian & Deuve, 2006 (India)
  * Orthogonius latus Hope, 1842 (Africa)
  * Orthogonius ledouxi Tian & Deuve, 2006 (Vietnam)
  * Orthogonius legrandi Tian & Deuve, 2006 (Malaysia)
  * Orthogonius lieftincki Andrewes, 1936 (Indonesia)
  * Orthogonius limbourgi Tian & Deuve, 2016 (Vietnam)
  * Orthogonius loeicus Tian & Deuve, 2006 (Thailand)
  * Orthogonius loeiensis Tian & Deuve, 2006 (Thailand)
  * Orthogonius longicornis Chaudoir, 1872 (Sri Lanka and Thailand)
  * Orthogonius longilamella Tian & Deuve, 2008
  * Orthogonius longipennis Hope, 1842
  * Orthogonius longiphallus Tian & Deuve, 2005 (Bhutan)
  * Orthogonius lucidus Bates, 1891 (India)
  * Orthogonius luzonicus Chaudoir, 1872 (Philippines)
  * Orthogonius macrophthalmus Tian & Deuve, 2013 (Vietnam)
  * Orthogonius maindronianus Tian & Deuve, 2006 (India)
  * Orthogonius makiling Tian & Deuve, 2006 (Philippines)
  * Orthogonius malacca Tian & Deuve, 2006 (Malaysia)
  * Orthogonius malaisei Andrewes, 1947 (Myanmar)
  * Orthogonius malaya Tian & Deuve, 2006 (Malaysia)
  * Orthogonius malaysiaensis Tian & Deuve, 2006 (Malaysia)
  * Orthogonius medanensis Tian & Deuve, 2006 (Indonesia)
  * Orthogonius meghalayaensis Tian & Deuve, 2016 (India)
  * Orthogonius mekongensis Tian & Deuve, 2006 (Laos)
  * Orthogonius melanipes Tian & Deuve, 2006 (India)
  * Orthogonius mellyi (Chaudoir, 1850) (Bangladesh, India, and Myanmar)
  * Orthogonius minimus Tian & Deuve, 2006 (India)
  * Orthogonius mniszechi Chaudoir, 1872 (India, Malaysia, Indonesia, and Borneo)
  * Orthogonius moestus Chaudoir, 1872
  * Orthogonius monolophus Andrewes, 1931 (Indonesia and Borneo)
  * Orthogonius montanus Tian & Deuve, 2006 (Myanmar)
  * Orthogonius morvani Tian & Deuve, 2003 (Thailand)
  * Orthogonius morvanianus Tian & Deuve, 2016 (Thailand)
  * Orthogonius mouhoti Chaudoir, 1872 (Southeast Asia)
  * Orthogonius myanmarensis Tian & Deuve, 2006 (Myanmar)
  * Orthogonius nahaeo Tian & Deuve, 2006 (Thailand)
  * Orthogonius nepalensis Habu, 1979 (Nepal)
  * Orthogonius nepalicus Tian & Deuve, 2006 (Nepal)
  * Orthogonius niger Jedlicka, 1935 (Philippines)
  * Orthogonius nigripes Tian & Deuve, 2001 (China and Vietnam)
  * Orthogonius nongfaensis Tian & Kirschenhofer, 2013 (Laos)
  * Orthogonius nyassicus Kolbe, 1895
  * Orthogonius obliquepes Tian & Deuve, 2006 (Philippines)
  * Orthogonius opacus Schmidt-Goebel, 1846 (Indomalaya)
  * Orthogonius orphnodes Andrewes, 1930 (Indonesia)
  * Orthogonius ovatulus Tian & Deuve, 2003 (Sri Lanka)
  * Orthogonius ovatus Chaudoir, 1878 (Indonesia)
  * Orthogonius pachlatkoi Tian & Deuve, 2006 (Thailand)
  * Orthogonius palangbangensis Tian & Deuve, 2006 (Indonesia)
  * Orthogonius panghongae Tian & Deuve, 2006 (China)
  * Orthogonius pangi Tian & Deuve, 2006 (Thailand and Vietnam)
  * Orthogonius parallelus Chaudoir, 1872 (Sri Lanka)
  * Orthogonius parastantoni Tian & Deuve, 2006 (Sri Lanka)
  * Orthogonius parasuturalis Tian & Deuve, 2006 (Indonesia)
  * Orthogonius parcepunctatus Kolbe, 1895 (Tanzania)
  * Orthogonius paris Tian & Deuve, 2006 (Thailand)
  * Orthogonius parvus Chaudoir, 1872 (India)
  * Orthogonius pectinatus Tian & Deuve, 2016 (Thailand and Vietnam)
  * Orthogonius perakensis Tian & Deuve, 2006 (Malaysia)
  * Orthogonius perakicus Tian & Deuve, 2007 (Malaysia)
  * Orthogonius perpuncticollis Burgeon, 1937 (Africa)
  * Orthogonius petiolaris Tian & Deuve, 2006 (India)
  * Orthogonius piceus Chaudoir, 1872 (Malaysia and Indonesia)
  * Orthogonius picilabris (W.S.MacLeay, 1825) (Malaysia, Indonesia, and Borneo)
  * Orthogonius picipennis Chaudoir, 1872 (Cambodia and Vietnam)
  * Orthogonius pinguis Murray, 1858 (Nigeria)
  * Orthogonius pinophilus Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius planiger (Walker, 1858) (Sri Lanka)
  * Orthogonius plebius Tian & Deuve, 2006 (Sri Lanka)
  * Orthogonius poggii Tian & Deuve, 2007
  * Orthogonius politior Tian & Deuve, 2016 (Laos)
  * Orthogonius politus Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
  * Orthogonius pollinctor Basilewsky, 1948 (Democratic Republic of the Congo)
  * Orthogonius pradieri Chaudoir, 1872
  * Orthogonius pseudochaudoiri Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius pseudocheni Tian & Deuve, 2006 (Vietnam)
  * Orthogonius pseudolongicornis Tian & Deuve, 2006 (Southeast Asia)
  * Orthogonius pseudomouhoti Tian & Deuve, 2008 (Cambodia)
  * Orthogonius punctatellus Tian & Deuve, 2006 (Indonesia)
  * Orthogonius punctulatus Chaudoir, 1872 (India)
  * Orthogonius punctum Tian & Deuve, 2016 (Thailand and Vietnam)
  * Orthogonius putzeysi Tian & Deuve, 2006 (Sri Lanka)
  * Orthogonius quadrisetosus Tian & Deuve, 2006 (Malaysia)
  * Orthogonius rhodesianus Csiki, 1932
  * Orthogonius rhodesiensis Lorenz, 1998 (Zambia)
  * Orthogonius ribbei Tian & Deuve, 2003 (Indonesia)
  * Orthogonius rogueti Tian & Deuve, 2016 (India)
  * Orthogonius rotundatus Tian & Deuve, 2006
  * Orthogonius rufiventris Bates, 1892 (Myanmar)
  * Orthogonius rufotestaceus G.Müller, 1941 (Ethiopia)
  * Orthogonius sabahensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
  * Orthogonius sabahicus Tian & Deuve, 2016 (Malaysia, Indonesia, and Borneo)
  * Orthogonius salvazai Tian & Deuve, 2006 (Laos)
  * Orthogonius sarawakensis Tian & Deuve, 2006 (Malaysia, Indonesia, and Borneo)
  * Orthogonius saundersi Andrewes, 1926 (Malaysia and Singapore)
  * Orthogonius schaumi Chaudoir, 1872 (Sri Lanka and Indonesia)
  * Orthogonius schaumioides Tian & Deuve, 2006 (India)
  * Orthogonius schauteni Tian & Deuve, 2006 (Malaysia)
  * Orthogonius schmidtgoebeli Chaudoir, 1872 (Myanmar)
  * Orthogonius senegalensis Dejean, 1831 (Africa)
  * Orthogonius setosopalpiger Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius siamensis Tian & Deuve, 2006 (Thailand)
  * Orthogonius similaris Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius simplicatus Tian & Deuve, 2006 (Indonesia)
  * Orthogonius singaporensis Tian & Deuve, 2006 (Singapore)
  * Orthogonius sinuatiphallus Tian & Deuve, 2001 (China)
  * Orthogonius smetsi Tian & Deuve, 2006 (Cambodia)
  * Orthogonius solidicornis Tian & Deuve, 2003 (China)
  * Orthogonius solidus Tian & Deuve, 2006 (India)
  * Orthogonius spinosus Burgeon, 1937 (Africa)
  * Orthogonius srilankaicus Tian & Deuve, 2003 (Sri Lanka)
  * Orthogonius stantoni Tian & Deuve, 2006 (India)
  * Orthogonius sterbai Jedlicka, 1935 (Philippines)
  * Orthogonius stygius Andrewes, 1930 (Malaysia and Indonesia)
  * Orthogonius sulawesicus Tian & Deuve, 2003 (Indonesia)
  * Orthogonius sulcatoides Tian & Deuve, 2006 (India, Myanmar, and Malaysia)
  * Orthogonius sulcatus Schmidt-Goebel, 1846 (Myanmar and Cambodia)
  * Orthogonius sumatraicus Tian & Deuve, 2006 (Indonesia)
  * Orthogonius suturalis Chaudoir, 1872 (Malaysia, Indonesia, and Borneo)
  * Orthogonius taghavianae Tian; Deuve & Felix, 2012 (Thailand)
  * Orthogonius taiwanicus Tian & Deuve, 2006 (Taiwan)
  * Orthogonius tangsen Tian & Deuve, 2006 (India)
  * Orthogonius tenasserimensis Tian & Deuve, 2006 (Myanmar)
  * Orthogonius termiticola Wasmann, 1902 (Malaysia)
  * Orthogonius thaicus Tian & Deuve, 2003 (Thailand)
  * Orthogonius thaiensis Tian & Deuve, 2006 (Thailand)
  * Orthogonius thailandensis Tian & Deuve, 2006 (Thailand)
  * Orthogonius thoracicus Gestro, 1875
  * Orthogonius tikekee Tian & Deuve, 2006 (Myanmar)
  * Orthogonius tonkinensis Tian & Deuve, 2006 (Vietnam)
  * Orthogonius tonkinicus Tian & Deuve, 2006 (Vietnam)
  * Orthogonius travancore Tian & Deuve, 2006 (India)
  * Orthogonius tricarinatus Burgeon, 1937 (Africa)
  * Orthogonius uncipennis Tian & Deuve, 2006 (Nepal)
  * Orthogonius vari Tian; Deuve & Felix, 2012 (Thailand and Cambodia)
  * Orthogonius variabilis Tian; Deuve & Felix, 2012 (China and Thailand)
  * Orthogonius virgulatus Andrewes, 1931 (Indonesia and Borneo)
  * Orthogonius wallardiensis Tian & Deuve, 2006 (India)
  * Orthogonius wrasei Tian & Deuve, 2016 (Myanmar)
  * Orthogonius xanthomerus L.Redtenbacher, 1868 (China)
  * Orthogonius yoga Tian & Deuve, 2006 (India)
  * Orthogonius yunnanensis Tian & Deuve, 2001 (China)
  