title: Phyllophaga tristis
text: # Phyllophaga tristis
  
  Phyllophaga tristis, the tristis complex, is a species of scarab beetle in the family Scarabaeidae.
  
  ## Subspecies
  
  These three subspecies belong to the species Phyllophaga tristis:
  
  * Phyllophaga tristis amplicornis Reinhard, 1939
  * Phyllophaga tristis suttonana Reinhard, 1939
  * Phyllophaga tristis tristis (Fabricius, 1781)
  