title: List of Copris species
text: # List of Copris species
  
  These species belong to Copris, a genus of dung beetles in the family Scarabaeidae.
  
  ## Copris species
  
  * Copris acutidens Motschulsky, 1860
  * Copris afgoi Cambefort, 1992
  * Copris agnus Sharp, 1875
  * Copris algol Nguyen-phung, 1989
  * Copris amabilis Kolbe, 1914
  * Copris amyntor Klug, 1855
  * Copris anceus (Olivier, 1789)
  * Copris andrewesi Waterhouse, 1891
  * Copris angusticornis Arrow, 1933
  * Copris angustus Nguyen-phung & Cambefort, 1986
  * Copris anomiopseoides Boucomont, 1924
  * Copris antares Ferreira, 1958
  * Copris arcturus Gillet, 1907
  * Copris ares Zelenka, 1993
  * Copris arimotoi Ochi & Araya, 1992
  * Copris arizonensis Schaeffer, 1906
  * Copris armatus Harold, 1869
  * Copris armeniacus Faldermann, 1835
  * Copris armiger Gillet, 1910
  * Copris arrowi Felsche, 1910
  * Copris aspericollis Gillet, 1910
  * Copris atropolitus Gillet, 1910
  * Copris bacchus Thunberg, 1787
  * Copris barclayi Ochi, Kon & Bai, 2009
  * Copris basilewskyi Ferreira, 1962
  * Copris basipunctatus Balthasar, 1942
  * Copris bellator Chevrolat, 1844
  * Copris bengalensis Gillet, 1911
  * Copris bidens Kolbe, 1893
  * Copris binodis
  * Copris bootes Klug, 1855
  * Copris boucardi Harold, 1869
  * Copris bovinus Gillet, 1908
  * Copris brachypterus Nomura, 1964
  * Copris brittoni Ferreira, 1962
  * Copris caelatus (Fabricius, 1794)
  * Copris caliginosus Kohlmann, Cano & Delgado, 2003
  * Copris cambeforti Nguyen-phung, 1988
  * Copris cambodiensis Ochi, Kon & Kawahara, 2008
  * Copris camerunus Felsche, 1904
  * Copris caobangensis Bui, Dumack & Bonkowski, 2018
  * Copris capensis Waterhouse, 1891
  * Copris cariniceps Felsche, 1910
  * Copris carinicus Gillet, 1910
  * Copris carmelita Fabricius, 1801
  * Copris cassius Péringuey, 1901
  * Copris celebensis Lansberge, 1886
  * Copris cheni Ochi, Kon & Bai, 2007
  * Copris chimalapensis Mora-Aguilar & Delgado, 2015
  * Copris ciliatus
  * Copris colmanti Gillet, 1908
  * Copris complexus Nguyen-phung, 1989
  * Copris compressipennis Gillet, 1910
  * Copris confucius Harold, 1877
  * Copris coriarius Gillet, 1907
  * Copris cornifrons Boheman, 1860
  * Copris corniger Sahlberg, 1823
  * Copris corpulentus Gillet, 1910
  * Copris costaricensis Gahan, 1894
  * Copris costariensis Matthews
  * Copris crassus Deschodt & Davis, 2015
  * Copris cribratus Gillet, 1927
  * Copris cribricollis Gillet, 1910
  * Copris curlettii Moretto, 2011
  * Copris davidis (Deyrolle, 1878)
  * Copris davisi Nguyen-phung & Cambefort, 1986
  * Copris davisoni Waterhouse, 1891
  * Copris decellei Cambefort, 1992
  * Copris delicatus Arrow, 1931
  * Copris denticulatus Nguyen-phung, 1988
  * Copris diversus Waterhouse, 1891
  * Copris draco Arrow, 1906
  * Copris dracunculus Fereira, 1959
  * Copris druidum Heer, 1862
  * Copris dudleyi Cambefort, 1992
  * Copris eburneus Cambefort, 1992
  * Copris elpheneroides Felsche, 1910
  * Copris elphenor Klug, 1855
  * Copris erratus Lansberge, 1886
  * Copris evanidus Klug, 1855
  * Copris excisus Waterhouse, 1891
  * Copris fallaciosus Gillet, 1907
  * Copris fallax Felsche, 1910
  * Copris felschei Reitter, 1892
  * Copris fidius (Olivier, 1789)
  * Copris frankenbergerianus Balthasar, 1958
  * Copris freyi Ferreira, 1961
  * Copris fricator (Fabricius, 1787)
  * Copris fukiensis Balthasar, 1952
  * Copris furciceps Felsche, 1910
  * Copris garambae Cambefort, 1992
  * Copris gazellarum Gillet, 1918
  * Copris gibbulus Lansberge, 1886
  * Copris gilleti Kolbe, 1907
  * Copris gladiator Arrow, 1933
  * Copris gopheri Hubbard, 1894
  * Copris gracilis Waterhouse, 1891
  * Copris graueri Kolbe, 1914
  * Copris halffteri Matthews, 1959
  * Copris harrisi Waterhouse, 1891
  * Copris hartli Frey, 1975
  * Copris hatayamai Ochi & Araya, 1992
  * Copris hispanus (Linnaeus, 1764)
  * Copris howdeni Matthews & Halffter, 1959
  * Copris humilis Gillet, 1908
  * Copris hybridus Gillet, 1910
  * Copris igualensis Warner, 1990
  * Copris illotus Balthasar, 1942
  * Copris imitans Felsche, 1910
  * Copris inaequabilis Zhang, 1997
  * Copris incertus Say, 1835
  * Copris inemarginatus Blatchley, 1918
  * Copris inhalatus Quedenfeldt, 1884
  * Copris insidiosus Péringuey, 1901
  * Copris integer Reiche, 1847
  * Copris interioris Kolbe, 1897
  * Copris iris Sharp, 1875
  * Copris jacchoides Nguyen-phung & Cambefort, 1987
  * Copris jacchus (Fabricius, 1775)
  * Copris jahi Nguyen-phung & Cambefort, 1986
  * Copris johkii Ochi & Kon, 2014
  * Copris juanmorai Mora-Aguilar & Delgado, 2015
  * Copris jucundus Gillet, 1932
  * Copris kachinensis Ochi, Kon & Kawahara, 2008
  * Copris kartlinus Kabakov, 1988
  * Copris kasagii Ochi & Kon, 1996
  * Copris kasanka Josso & Prévost, 2016
  * Copris katangae Cambefort & Nguyen-phung, 1996
  * Copris keralensis Gill, 1986
  * Copris kiuchii Masumoto, 1989
  * Copris klapperichi Balthasar, 1942
  * Copris klugi Harold, 1869
  * Copris kusakabei Ochi, Kon & Kawahara, 2005
  * Copris laeviceps Harold, 1862
  * Copris laevigatus Gillet, 1927
  * Copris laiiformis Nguyen-phung & Cambefort, 1987
  * Copris laioides Boucomont, 1932
  * Copris laius Harold, 1868
  * Copris lannathai Hanboonsong & Masumoto & Ochi, 2003
  * Copris leakeyorum Paulian, 1976
  * Copris lecontei Matthews, 1962
  * Copris leruei Josso & Prévost, 2017
  * Copris lugubris Boheman, 1858
  * Copris lunaris [fr] (Linnaeus, 1758)
  * Copris macacus Lansberge, 1886
  * Copris macclevei Warner, 1990
  * Copris macer Péringuey, 1901
  * Copris maesi Ratcliffe, 1998
  * Copris magicus Harold, 1881
  * Copris manni Ochi, Kon & Bai, 2009
  * Copris marcus Gillet, 1921
  * Copris martinae Nguyen-phung, 1988
  * Copris martinezi Matthews and Halffter, 1968
  * Copris matthewsi Delgado & Kohlmann, 2001
  * Copris medogensis Zhang, 1988
  * Copris megaceratoides Waterhouse, 1891
  * Copris megasoma Matthews & Halffter, 1959
  * Copris mesacanthus Harold, 1878
  * Copris mexicanus Matthews & Halffter, 1959
  * Copris minutus (Drury, 1773)
  * Copris misellus Péringuey, 1901
  * Copris moechus LeConte, 1854
  * Copris moffartsi Gillet, 1907
  * Copris mongkhoni Hanboonsong & Masumoto & Ochi, 2003
  * Copris montivagus Gillet, 1908
  * Copris morphaeus Gillet, 1932
  * Copris mourgliai Zelenka, 1993
  * Copris neglectus Moxey, 1963
  * Copris neotridens Génier & Prévost, 2017
  * Copris nepos Gillet, 1908
  * Copris nevinsoni Waterhouse, 1891
  * Copris novaki Zelenka, 1992
  * Copris nubilosus Kohlmann & Cano & Delgado, 2003
  * Copris numa Lansberge, 1886
  * Copris obenbergeri Balthasar, 1933
  * Copris obesus Boheman, 1857
  * Copris ochus (Motschulsky, 1860)
  * Copris orion Klug, 1835
  * Copris orphanus Guérin-Méneville, 1847
  * Copris parapecuarius Ochi, Kon & Kawahara, 2008
  * Copris pauliani Nguyen-phung & Cambefort, 1986
  * Copris pecuarius Lewis, 1884
  * Copris pedarioides Lansberge, 1886
  * Copris phungae Cambefort, 1992
  * Copris phylax Gillet, 1908
  * Copris poggii Ochi & Kon, 2005
  * Copris potanini Semenov, 1891
  * Copris prevosti Josso, 2011
  * Copris pristinus Pierce, 1946
  * Copris pseudobootes Ferreira, 1962
  * Copris pseudochus Ochi & Kon, 2004
  * Copris pseudomoffartsi Moretto, 2011
  * Copris pseudosinicus Balthasar, 1958
  * Copris pueli Mollandin De Boissy, 1905
  * Copris punctatus Gillet, 1910
  * Copris puncticollis Boheman, 1857
  * Copris punctipennis Boucomont, 1914
  * Copris punctulatus Wiedemann, 1823
  * Copris punjabensis Gillet, 1921
  * Copris quasilaevigatus Ochi, Kon & Bai, 2007
  * Copris ramosiceps Gillet, 1921
  * Copris rebouchei Harold, 1869
  * Copris remotus Leconte, 1866
  * Copris renwarti Nguyen-phung & Cambefort, 1986
  * Copris repertus Walker, 1858
  * Copris ribbei Lansberge, 1886
  * Copris riekoae Ochi & Kon, 2003
  * Copris ritsemae Harold, 1875
  * Copris rugosus Gillet, 1908
  * Copris ruricola Balthasar, 1933
  * Copris sabinus Gillet, 1910
  * Copris sacontala Redtenbacher, 1848
  * Copris sallei Harold, 1869
  * Copris sarpedon Harold, 1868
  * Copris saundersi Harold, 1869
  * Copris schoolmeestersi Ochi & Kon, 2010
  * Copris scorpio Zelenka, 1993
  * Copris serius Nguyen-phung, 1987
  * Copris sexdentatus Thunberg, 1818
  * Copris siamensis Gillet, 1910
  * Copris siangensis Biswas, 1980
  * Copris sierrensis Matthews, 1961
  * Copris signatus Walker, 1858
  * Copris simonettai Cambefort, 1992
  * Copris simulator Nguyen-phung & Cambefort, 1986
  * Copris singularis Nguyen-phung & Cambefort, 1986
  * Copris sinicus Hope, 1842
  * Copris sinon (Olivier, 1789)
  * Copris sodalis Walker, 1858
  * Copris sonensis Bui, Dumack & Bonkowski, 2018
  * Copris sorex Balthasar, 1942
  * Copris sphaeropterus Harold, 1877
  * Copris spinator Harold, 1881
  * Copris subdolus Balthasar, 1958
  * Copris subpunctatus Gillet, 1910
  * Copris subsidens Péringuey, 1901
  * Copris subterraneus Heer, 1862
  * Copris sumatrensis Gillet, 1921
  * Copris surdus Arrow, 1931
  * Copris szechouanicus Balthasar, 1958
  * Copris tetraodon Gillet, 1910
  * Copris tridentatus Solis & Kohlmann, 2003
  * Copris tripartitus Waterhouse, 1875
  * Copris truncatus Felsche, 1901
  * Copris tsukamotoi Ochi, Kon & Kawahara, 2008
  * Copris typhoeus Gerstaecker, 1884
  * Copris uenoi Hanboonsong & Masumoto & Ochi, 2003
  * Copris umbilicatus Abeille De Perrin, 1901
  * Copris urus Boheman, 1857
  * Copris usambaricus Gillet, 1908
  * Copris vankhaii Nguyen-phung, 1988
  * Copris victorini Boheman, 1857
  * Copris vietnamicus Kabakov, 1994
  * Copris vilhenai Ferreira, 1962
  * Copris vrydaghi Ferreira, 1962
  * Copris warneri Mccleve & Kohlmann, 2005
  * Copris wiesei Kolbe, 1914
  * Copris yangi Ochi, Kon & Bai, 2007
  * Copris youngai Balthasar, 1967
  * Copris zhangi Ochi, Kon & Bai, 2009
  