title: Helius (fly)
text: # Helius (fly)
  
  Helius is a genus of crane fly in the family Limoniidae.
  
  ## Distribution
  
  Worldwide, but the Australasian and Oriental regions are the most species rich.
  
  ## Species
  
  * Subgenus Eurhamphidia Alexander, 1915
  
     * H. abnormalis (Brunetti, 1918)
  
     * H. ata Alexander, 1931
  
     * H. atroapicalis Alexander, 1978
  
     * H. auranticolor Alexander, 1936
  
     * H. connectus Edwards, 1928
  
     * H. diacanthus Alexander, 1931
  
     * H. fuscofemoratus Alexander, 1931
  
     * H. glabristylatus Alexander, 1931
  
     * H. indivisus Alexander, 1931
  
     * H. inelegans (Alexander, 1928)
  
     * H. invenustipes (Alexander, 1930)
  
     * H. melanosoma Alexander, 1936
  
     * H. mimicans Edwards, 1933
  
     * H. mirus Edwards, 1926
  
     * H. monticola (Edwards, 1919)
  
     * H. mouensis Alexander, 1948
  
     * H. nigrofemoratus (Alexander, 1930)
  
     * H. niveitarsis (Skuse, 1890)
  
     * H. pallens Edwards, 1933
  
     * H. perelegans Alexander, 1930
  
     * H. perlongatus Alexander, 1978
  
     * H. scabiosus Alexander, 1932
  
     * H. vitiensis Alexander, 1956
  * Subgenus Helius Lepeletier & Serville, 1828
  
     * H. acanthostyla Alexander, 1944
  
     * H. aciferus Alexander, 1937
  
     * H. aka Alexander, 1972
  
     * H. albitarsis (Osten Sacken, 1888)
  
     * H. albogeniculatus Alexander, 1936
  
     * H. amplus Edwards, 1933
  
     * H. anaemicus Alexander, 1932
  
     * H. anamalaiensis Alexander, 1967
  
     * H. angustalbus Alexander, 1953
  
     * H. aphrophilus Alexander, 1948
  
     * H. apicalis (Alexander, 1915)
  
     * H. apoensis Alexander, 1931
  
     * H. apophysalis Alexander, 1967
  
     * H. araucariae Alexander, 1945
  
     * H. arcuarius Alexander, 1929
  
     * H. argyrosternus Alexander, 1930
  
     * H. arunachalus Alexander, 1975
  
     * H. attenuatus Alexander, 1929
  
     * H. barbatus Edwards, 1921
  
     * H. bicolor Edwards, 1933
  
     * H. bifurcus Alexander, 1956
  
     * H. bitergatus Alexander, 1950
  
     * H. boops Alexander, 1942
  
     * H. brachyphallus Alexander, 1956
  
     * H. brevioricornis (Alexander, 1920)
  
     * H. brevisector Alexander, 1956
  
     * H. brunneus Byers, 1981
  
     * H. cacoxenus (Alexander, 1920)
  
     * H. calviensis Edwards, 1928
  
     * H. capensis (Alexander, 1917)
  
     * H. capniopterus Alexander, 1945
  
     * H. catreus Alexander, 1967
  
     * H. cavernicolus Alexander, 1961
  
     * H. chikurinensis Alexander, 1930
  
     * H. chintoo Theischinger, 1994
  
     * H. communis (Skuse, 1890)
  
     * H. comoreanus Alexander, 1959
  
     * H. compactus Alexander, 1967
  
     * H. copiosus Alexander, 1935
  
     * H. corniger Savchenko, 1983
  
     * H. costofimbriatus Alexander, 1930
  
     * H. costosetosus Alexander, 1932
  
     * H. creper Alexander, 1926
  
     * H. ctenonycha Alexander, 1938
  
     * H. dafla Alexander, 1972
  
     * H. darlingtonae Welch & Gelhaus, 1994
  
     * H. destitutus Alexander, 1971
  
     * H. devinctus Alexander, 1932
  
     * H. distinervis Alexander, 1940
  
     * H. dolichorhynchus Edwards, 1933
  
     * H. eremnophallus Alexander, 1978
  
     * H. euryphallus Alexander, 1960
  
     * H. fasciventris Edwards, 1926
  
     * H. ferruginosus (Brunetti, 1912)
  
     * H. flavidibasis Alexander, 1975
  
     * H. flavipes (Macquart, 1855)
  
     * H. flavitarsis (Alexander, 1920)
  
     * H. flavus (Walker, 1856)
  
     * H. fragosus Alexander, 1937
  
     * H. franckianus Alexander, 1940
  
     * H. fratellus (Brunetti, 1918)
  
     * H. fulvithorax (Skuse, 1890)
  
     * H. fumicosta Edwards, 1928
  
     * H. fuscoangustus Alexander, 1967
  
     * H. garcianus Alexander, 1972
  
     * H. gibbifer Savchenko, 1981
  
     * H. gorokanus Alexander, 1962
  
     * H. gracillimus Alexander, 1938
  
     * H. graphipterus Alexander, 1954
  
     * H. haemorrhoidalis Alexander, 1937
  
     * H. harrisi (Alexander, 1923)
  
     * H. hatschbachi Alexander, 1954
  
     * H. hispanicus Lackschewitz, 1928
  
     * H. hova Alexander, 1955
  
     * H. impensus Alexander, 1967
  
     * H. imperfectus (Alexander, 1920)
  
     * H. inconspicuus (Brunetti, 1912)
  
     * H. ineptus Alexander, 1938
  
     * H. infirmus Alexander, 1932
  
     * H. invariegatus Alexander, 1938
  
     * H. iris (Alexander, 1920)
  
     * H. kambangani (de Meijere, 1913)
  
     * H. kambanganoides Alexander, 1931
  
     * H. larotypa Alexander, 1926
  
     * H. lectus Alexander, 1936
  
     * H. leucoplaca Alexander, 1936
  
     * H. lienpingensis Alexander, 1945
  
     * H. liguliferus Alexander, 1978
  
     * H. liliputanus Alexander, 1929
  
     * H. lobuliferus Alexander, 1938
  
     * H. longinervis Edwards, 1928
  
     * H. longirostris (Meigen, 1818)
  
     * H. luachimoensis Alexander, 1963
  
     * H. luniger (Riedel, 1914)
  
     * H. mainensis (Alexander, 1916)
  
     * H. malagasicus Alexander, 1955
  
     * H. manueli Alexander, 1972
  
     * H. medleri Alexander, 1976
  
     * H. melanophallus Alexander, 1938
  
     * H. mesorhynchus Alexander, 1933
  
     * H. micracanthus Alexander, 1945
  
     * H. mirabilis (Alexander, 1914)
  
     * H. mirandus (Alexander, 1921)
  
     * H. morosus (Alexander, 1920)
  
     * H. multivolus Alexander, 1945
  
     * H. murreensis Alexander, 1960
  
     * H. myersiellus Alexander, 1935
  
     * H. nawaianus Alexander, 1929
  
     * H. neocaledonicus Alexander, 1945
  
     * H. nigricapella Alexander, 1938
  
     * H. nigriceps (Edwards, 1916)
  
     * H. nipponensis (Alexander, 1913)
  
     * H. numenius Alexander, 1967
  
     * H. obliteratus (Alexander, 1920)
  
     * H. obsoletus (Alexander, 1920)
  
     * H. oxystylus Alexander, 1967
  
     * H. pallidipes Alexander, 1926
  
     * H. pallidissimus Alexander, 1930
  
     * H. pallirostris Edwards, 1921
  
     * H. panamensis Alexander, 1934
  
     * H. paramorosus Alexander, 1949
  
     * H. parvidens Alexander, 1944
  
     * H. patens Edwards, 1933
  
     * H. pauperculus (Alexander, 1921)
  
     * H. pavoninus Alexander, 1938
  
     * H. perflavens Alexander, 1964
  
     * H. perpallidus Alexander, 1942
  
     * H. pervenustus Alexander, 1953
  
     * H. phasmatis Alexander, 1945
  
     * H. pictus Edwards, 1926
  
     * H. plebeius Alexander, 1946
  
     * H. pluto Alexander, 1932
  
     * H. polionotus Alexander, 1938
  
     * H. procerus Alexander, 1931
  
     * H. productellus Alexander, 1944
  
     * H. protumidus Alexander, 1978
  
     * H. quadrifidus Alexander, 1926
  
     * H. quadrivena Alexander, 1971
  
     * H. rectispinus Alexander, 1947
  
     * H. rectus Alexander, 1945
  
     * H. regius Alexander, 1944
  
     * H. rostratus Alexander, 1956
  
     * H. rubicundus (Alexander, 1922)
  
     * H. rufescens (Edwards, 1916)
  
     * H. rufithorax Alexander, 1928
  
     * H. sanguinolentus (Alexander, 1921)
  
     * H. schildi Alexander, 1945
  
     * H. selectus Alexander, 1936
  
     * H. serenus Alexander, 1967
  
     * H. setigerus Alexander, 1980
  
     * H. sigillatus Alexander, 1964
  
     * H. stenorhynchus Alexander, 1954
  
     * H. stolidus Alexander, 1948
  
     * H. subanaemicus Alexander, 1964
  
     * H. subarcuarius Alexander, 1934
  
     * H. subfasciatus Alexander, 1924
  
     * H. submorosus (Alexander, 1921)
  
     * H. subobsoletus (Alexander, 1921)
  
     * H. subpauperculus Alexander, 1975
  
     * H. tantalus Alexander, 1967
  
     * H. tanyrhinus Alexander, 1964
  
     * H. taos Alexander, 1967
  
     * H. tenuirostris Alexander, 1924
  
     * H. tenuistylus Alexander, 1929
  
     * H. tetracradus Alexander, 1967
  
     * H. tienmuanus Alexander, 1940
  
     * H. tonaleah Theischinger, 1994
  
     * H. trianguliferus Alexander, 1931
  
     * H. tshinta Theischinger, 1994
  
     * H. unicolor (Brunetti, 1912)
  
     * H. uniformis (Alexander, 1914)
  
     * H. venustus (Skuse, 1890)
  
     * H. verticillatus Alexander, 1967
  
     * H. yindi Theischinger, 1994
  * Subgenus Idiohelius Alexander, 1948
  
     * H. melanolithus Alexander, 1953
  
     * H. mirificus Alexander, 1953
  
     * H. pentaneura Alexander, 1948
  * Subgenus Mammuthonasus Theischinger, 1994
  
     * H. allunga Theischinger, 1994
  * Subgenus Prohelius Alexander, 1975
  
     * H. dugaldi Alexander, 1945
  
     * H. edwardsianus Alexander, 1956
  
     * H. fulani Alexander, 1975
  * Subgenus Rhamphidina Alexander, 1920
  
     * H. camerounensis (Alexander, 1920)
  * Subgenus Rhamphidioides Alexander, 1920
  
     * H. alluaudi (Riedel, 1914)
  
     * H. venustissimus (Alexander, 1920)
  * Subgenus Rhampholimnobia Alexander, 1915
  
     * H. bigeminatus Alexander, 1962
  
     * H. brevinasus Alexander, 1934
  
     * H. deentoo Theischinger, 1994
  
     * H. diffusus Alexander, 1941
  
     * H. fenestratus Alexander, 1973
  
     * H. gracilirostris Alexander, 1962
  
     * H. guttulinus Alexander, 1960
  
     * H. japenensis Alexander, 1948
  
     * H. matilei Hynes, 1993
  
     * H. mesolineatus Alexander, 1962
  
     * H. nimbus Alexander, 1941
  
     * H. papuanus Alexander, 1934
  
     * H. reticularis (Alexander, 1915)
  
     * H. simulator Alexander, 1962
  
     * H. spinteris Alexander, 1960
  
     * H. subreticulatus Alexander, 1962
  * Subgenus Rhyncholimonia Alexander, 1964
  
     * H. dicroneurus Alexander, 1964
  