Dr. Stein’s area of research is on the impact of Wnt signaling abnormalities in cancer development and progression, with specific interest in liver cancer, melanoma, and osteosarcoma. As a veterinary oncologist, he is interested in using spontaneously-occurring cancer in domestic animals as a model for human cancers, especially for osteosarcoma and malignant melanoma. Dr. Stein completed his DVM at Iowa State University, his PhD and medical oncology residency at UW-Madison