He received his B.S. in Biology from St Lawrence University, his M.S. in Plant Biology from Indiana University, and Ph.D. in Cell, Molecular & Developmental Biology from the Sackler School at Tufts University. He completed postdoctoral training at the University of Rochester and Tufts School of Medicine. His major research interests focus on the eye with emphasis on the aqueous humor outflow pathway and its role in intraocular pressure (IOP) regulation and glaucoma.