He earned his Ph.D. from Pennsylvania State University College of Medicine and completed postdoctoral fellowships at the University of Alabama at Birmingham and Johns Hopkins University School of Medicine.Dr. Riddle’s research focuses on Wnt/b-catenin signaling in musculoskeletal development and in response to mechanical, hormonal, and other anabolic signals. His prior work examined skeletal mechanotransduction—the process in which mechanical signals are translated to biochemical signals.He is a member of the American Society for Bone & Mineral Research and the Endocrine Society. His work has been recognized with a 2011-2012 Musculoskeletal Pilot and Feasibility Grant from the JHU Center for Musculoskeletal Research and is supported by grants from the Veterans Administration and the National Institutes of Health.