He holds a Ph.D. from Arizona State University (advisor, Philip Hedrick) and B.S. from the University of Washington (advisor, Scott Edwards). Dan did postdoctoral training with Michael Hammer at the University of Arizona and John Wakeley and Richard Lewontin at Harvard University. Dan has worked on many genomic systems, from Drosophila speciation, to human population genetics, to MHC evolution in vertebrates, often developing innovative computational methods to address complex biological questions. At Lummei Analytics, Dan serves as the head programmer, database developer, and computer systems administrator. He automates large-scale NGS analysis, performs simulations, and conducts statistical analyses.