Dr. Graham received a BS in Chemical Engineering and French from Washington University in St. Louis in 2001. He received a PhD in Chemical Engineering from the California Institute of Technology in 2007 for research into the crosstalk between intercellular and intracellular signaling pathways under the guidance of Anand Asthagiri. He then did postdoctoral training at the University of California, Los Angeles, in the laboratory of Thomas Graeber, where he investigated the interactions between kinase signaling and tumor metabolism.