Meinig School of Biomedical Engineering at Cornell University. He obtained his Ph.D. at Cornell, studying the photophysics of photosynthesis using time-resolved laser spectroscopy. He later worked as a Research Associate in Applied and Engineering Physics under Watt Webb in the area of laser-scanning microscopy, during which time he was involved in the development of multiphoton microscopy at Cornell. His current research focuses on instrumentation and application developments in the areas of laser-scanning microscopy, fluorescence lifetime imaging (FLIM), super-resolution microscopy and single molecule level analysis of biological processes, which his lab applies to studies in cancer biology and transcriptional control. Core 2 Leader