He obtained his PhD from the Department of Clinical and Experimental Medicine, University of Perugia, Italy, working on the pharmacological regulation of the immune response. In 2005, he became a Research Assistant at Section of Pharmacology, University of Perugia, Italy. In 2007, he started a Career Development Fellow at the MRC Toxicology Unit, Leicester, UK, and then was appointed as Senior Investigator until 2014. In 2014 he was also a visiting scientist, at The Campbell Family Institute for Breast Cancer Research, Tak W. Mak laboratory, Toronto, Canada. In the last 5 years, his main research interests was studying the role of p73, a p53 family member, in neurodevelopment and neurotoxicity using mouse model.