His group studies how internal variables are calculated by the the Drosophila nervous system and how these variables influence behavior. Recent work has focused on several specific neural computations. For example, his group has studied how the fly brain internally subtracts one sensory signal from a set of neurons that carry multiple related signals. His group has also been characterizing the circuit architecture for how fly brains update their internal sense of orientation when they turn left or right in the dark. These results seek to provide a detailed, circuit-level understanding of how brains internally store and update variables related to spatial orientation and navigation.