He received Bachelor’s Degrees in Biology and Anthropology from the University of California, San Diego and completed his Ph.D. in Biological Sciences in the lab of Dr. George Somero at Stanford University. Dr. Lockwood was an NIH postdoctoral fellow in the lab of Dr. Kristi Montooth at Indiana University. His lab works at the interface of physiology and evolutionary biology to elucidate the mechanisms that constrain or facilitate adaptation to environmental change.