He received an AB summa cum laude in biochemical sciences from Harvard in 2003 for his thesis on the statistical mechanics of protein design. Upon earning a Rhodes Scholarship to St. John's College, Oxford, Jeremy spent the following two years at the Rudolf Peierls Centre for Theoretical Physics studying the effects of stochastic noise on morphogen gradient patterning in embryonic development. Subsequently, as a Hertz Fellow at Stanford University, and also partly at Israel's Weizmann Institute of Science, he completed his doctorate in physics carrying out theoretical and computational studies of protein folding in vivo and in vitro.