Keeling Center for Comparative Medicine and Research. Dr. Schapiro earned his Ph.D. from the University of California at Davis, after receiving his B.A. from Johns Hopkins University. He completed a postdoctoral research fellowship at the Caribbean Primate Research Center of the University of Puerto Rico and joined the Department of Veterinary Sciences at the Keeling Center in 1989. In 2009, Dr. Schapiro was designated an Honorary Professor in the Department of Experimental Medicine at the University of Copenhagen, Denmark.