He received his Bachelor’s degree in 1982 and his Master’s degree in 1989 in animal science from Jilin Agricultural University, China. In 1996, he received his Ph.D. degree in Animal Molecular and Cell Biology from the University of Florida. He completed his postdoctoral training in Powell Gene Therapy Center at University of Florida in 2001. He has received 1999 Young Investigator Fellowship Award of the Alpha One Foundation, and 2009-2011 University of Florida Research Foundation (UFRF) Professorship Award.