He received a B.S. degree in engineering physics from Westmont College in 1997, an M.S. degree in civil engineering from UCLA in 1999, and a Ph.D. degree from Caltech in 2003. He is the recipient of a 2007 NSF CAREER award. Klug's primary scientific background is in continuum and computational modeling of the mechanics of solids and structures. He has particular experience in the development of numerical methods for modeling thin beam- and shell-like structures, and in the application of those methods to multi-physics problems in biology, from the macroscale to the nanoscale. Some of his current topics of study include mechanics and assembly of viruses, mechanics of biomembranes, mechanics of motor-driven semiflexible cytoskeletal networks, and electromechanics of the heart.