He is also a member of Stem Cell Institute and Interdisciplinary Biosciences Program at Columbia University. He received his Ph.D. from MIT in 2002. His current research interests focus on dynamic studies of molecules in live cells. His lab extends the expertise in superresolution imaging and computational modeling and simulation to address dynamic problems in biology. Specifically, the research areas can be categorized into the following three directions: (1) dynamics of signaling activities in primary cilia of mammalian cells; (2) mechanobiology of induced pluripotent stem cells; and (3) structure-function relationship of molecular motors. .