He has studied the social behaviour and evolution of various animals, including praying mantids, spiders, scorpions, termites, and a group of stalk-eyed flies that lives in SE Asia. He has studied the evolution and genetics of mate competition, mate choice, cooperation and altruism, and sexual cannibalism. When he moved to Singapore, he became enamoured with the recently returned smooth-coated otters, and has been studying them ever since.