He received his PhD in immunology from the All India Institute of Medical Sciences, New Delhi, India. After completing his post-doctoral fellowships at Yale University and Mount Sinai Medical Center, he joined Columbia University faculty as an Assistant Professor in the Department of Medicine. In addition, he also held joint appointments as an Associate Director of Vascular Biology Research Program and an Assistant Professor of Neurosurgery at St. Luke’s Roosevelt Hospital Center in New York before joining the University of Pittsburgh.