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A CHECKLIST OF 
THE MILLIPEDS OF 
MEXICO AND CENTRAL AMERICA 


For sale by the Superintendent of Documents, U.S. Government Printing Office 
Washington, D.C. 20402 - Price 70 cents (paper cover) 


UNITED STATES NATIONAL MUSEUM BULLETIN 266 


A Checklist of 
the Millipeds of 


Mexico and Central America 


H. F. LOOMIS 


Honorary Fellow, Department of 
Entomology, Smithsonian Institution 


Research Associate, Florida State 
Collection of Arthropods 





SMITHSONIAN INSTITUTION PRESS 
CITY OF WASHINGTON 
1968 


Publications of the United States National Museum 


The scientific publications of the United States National Museum in- 
clude two series, Proceedings of the United States National Museum and 
United States National Museum Bulletin. 

In these series are published original articles and monographs dealing 
with the collections and work of the Museum and setting forth newly ac- 
quired facts in the field of anthropology, biology, geology, history, and 
technology. Copies of each publication are distributed to libraries and 
scientific organizations and to specialists and others interested in the 
various subjects. 

The Proceedings, begun in 1878, are intended for the publication, in 
separate form, of shorter papers. These are gathered in volumes, octavo 
in size, with the publication date of each paper recorded in the table of 
contents of the volume. 

In the Bulletin series, the first of which was issued in 1875, appear 
longer, separate publications consisting of monographs (occasionally 
in several parts) and volumes in which are collected works on related 
subjects. Bulletins are either octavo or quarto in size, depending on the 
needs of the presentation. Since 1902, papers relating to the botanical 
collections of the Museum have been published in the Bulletin series 
under the heading Contributions from the United States National Herbarium. 

This work forms number 266 of the Bulletin series. 


Frank A. Taylor 


Director, United States National Museum 


U.S. GOVERNMENT PRINTING OFFICE 
WASHINGTON : 1968 


iv 


Contents 


Introduction 


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INTRODUCTION 


The milliped fauna of Mexico and Central America is very poorly 
known. Records of it are scattered through a large number of papers, 
only two of which apply to the area in its entirety. The first of these 
was by Henri Saussure and Alois Humbert, in 1872, and contained a 
catalog which was rather brief, needless to say, of the centipeds and 
millipeds then known from North and South America. The second com- 
prehensive publication was by R. I. Pocock, on the same groups in 
Mexico and Central America, appearing inthe Biologia Centrali- Americana 
from 1903 to 1910. Not only did this paper list all the previously known 
milliped species of these regions, many with supplemental descriptions, 
illustrations, and new locality records, but it added illustrated descrip- 
tions of 87 new ones and, for the first time, included complete keys to 
the various classification categories then in use. This paper today 
remains the outstanding systematic work on the Diplopoda of the area 
included in this checklist. 

Since 1910 only four other noteworthy papers have been published, 
all on smaller portions of the area. 1.--The Millipeds of Central America, 
R. V. Chamberlin, in 1922, recataloged the known species and added 
many new ones, but did not include any Mexican forms. 2.--On Mexican 
Millipeds, same author, in 1943, cataloged the Mexican species he had 
previously described and addednew ones. 3.--Myriapoden aus El Salvador, 
O. Kraus, 1954, cataloged previously known species, with new locality 
records, and described new ones. 4.--The Millipeds of Panama, H. F. 
Loomis, 1964, cataloged previously known species, with new locality 
records, described new ones, and gave keys to all categories as well as 
a bibliography of papers relating to the Panamanian species. This is the 
only extensive bibliography that has appeared on Mexican or Central 
American Diplopoda. Besides these four papers and the two previously 
mentioned, a total of more than 200 smaller ones apply to the area. 

Today this scattered literature contains descriptions and records of 
over 750 more or less well-known species, but this number is only a 
small part of the undiscovered ones inhabiting this vast warm-temperate 
to tropical area. 

A major reason for the backward state of our knowledge of the millipeds 
of this region, or most of its parts, probably is that none of the eight 
countries ever has produced a recognized diplopod taxonomist or any 
outstanding collectors, other than Drs. O. Bolivar and F. Bonet, of 
Mexico City, on whose collection Chamberlin based his Mexican paper 
in 1943. Thus foreign taxonomists and collectors have been the principal 


1 


2 U.S. NATIONAL MUSEUM BULLETIN 266 


contributors to the advances that have been made in our understanding 
and classification of this group. 

The present checklist has been prepared to overcome some of the 
difficulties of studying the millipeds of these countries, as presented 
by the dispersed literature and lack of bibliographies to it. In general, 
this checklist follows the arrangement of the admirable Checklist of the 
Millipeds of North America, by R. V. Chamberlin and R. L. Hoffman, and 
adopts the basic classification system they proposed on pages 7 and 8, 
except as altered later by the junior author to combine the orders Spiros- 
treptida and Cambalida, with the latter reduced to a suborder. As in that 
checklist, keys to all classification categories through families have 
been included here. 

Citations throughout this checklist, except where they do not apply to 
the area involved, have been reduced to the last name of the author, 
followed by the year of publication. Where two or more publications by 
the same author occurred in the same year, each is indicated by a letter 
added to the date, i.e., 1935a, 1935b, etc. Full citations for these abbre- 
viations are given in the bibliography which lists the references appli- 
cable to the class Diplopoda in Mexico and Central America. 


Where a change or changes have occurred in the use of the original 
name of a genus or species, such changes are cited in chronological 
order under the current name. In a few cases this results in making it 
appear that the last author changed the immediately preceding name, 
whereas his change may have been in relation to a still earlier one. 
When it is essential to determine this point, examination of the last 
publication involved usually will settle the matter. 

A few new synonyms and other nomenclatorial changes have been 
presented on the basis of my personal knowledge or of information com- 
municated by Drs. N. B. Causey and R. L. Hoffman, credit for such 
information being shown in each case. 

The depository of the holotype specimen of each species is named 
wherever possible. Dr. Hoffman graciously assumed the task of supplying 
the name of the depository of all species where the author could not do 
so. His familiarity with milliped types seen during past visits to Furopean 
museums greatly aided in this and the many names he supplied are all 
followed by his parenthesized last initial. 

Because so few of the species are known from more than one or two 
localities, no general statements on distribution have been given. Instead, 
type and paratype localities are shown, and all other published localities 
are listed, with the name of the recorder and the place of record indi- 
cated as for other bibliographic citations. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 3 


Special thanks are due Dr. R. L. Hoffman for reviewing manuscript 
of this checklist, for his participation in naming type depositories, and 
for help in numerous other ways. Grateful acknowledgment is made to 
Dr. N. B. Causey for her encouragement and aid in various ways, and to 
Dr. Ralph E. Crabill, Jr., for suggesting improvements in the checklist 
and for providing bibliographic and other data not available to me. 


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MILLIPEDS OF MEXICO AND CENTRAL AMERICA 5 


CHECKLIST 


Class DIPLOPODA 
KEY TO THE SUBCLASSES OF DIPLOPODA 


1. Body soft; segments few, 9 or 10, with dense tufts of peculiarly 
shaped setae and a brush of very long setae in last segment; males 
PROM APOMOPOUOS cscs mi cic eis lelnieisieis s sics =) shale © PSELAPHOGNATHA 

Body usually well chitinized; segments 14 to very many; setae sim- 
ple, if present, not in dense tufts or brushes; males with gono- 


Faia etter nia tess Snel © (erate c) a )insial einer eis’ = sene ayes ois) ciahel = ene CHILOGNATHA 


Subclass PSELAPHOGNATHA 
Order POLYXENIDA 


KEY TO THE FAMILIES OF POLYXENIDA 


1. Eyes present; last antennal joint much smaller than 7th joint; seg- 
ments with two transverse rows of bristles along posterior 
tesa ea escatn alec oh em, wd n'/0) ek eihay a) allatre (aire eUelle) a(x 5 e)\siey0 @ aves s POLY XENIDAE 

Eyes absent; last antennal joint approximating 7th joint in length; 
segments with one transverse row of longer bristles along posterior 


RR fatten esa ict lin ellerraiiwt oie /al karen ah aiiatiel “cl ctor, Gilal ove evra LOPHOPROCTIDAE 


Family LOPHOPROCTIDAE 
Genus ALLOPROCTINUS Jeeke! 1963b, p. 156. 


Alloproctus Silvestri 1948b, p. 217, preoccupied. 

TYPE SPECIES: Alloproctus guineensis Silvestri, by original desig- 
nation. 
Alloproctinus anisorhabdus Conde and Terver 1964, p. 19. 

TYPE: Conde Collection, Univ. of Nancy. 

TYPE LOCALITY: Guatemala. 


Genus LOPHOPROCTINUS Silvestri 1948b, p. 217. 


TYPE SPECIES: Lophoproctinus inferus Silvestri, by original desig- 
nation. 
Lophoproctinus diversunguis Silvestri 1949, p. 3. 

TYPE: Depository not known. 

TYPE LOCALITY: Orizaba, Veracruz, Mexico. 
Lophoproctinus mexicanus Silvestri 1949, p. 1. 

TYPE: Depository not known. 

TYPE LOCALITY: Jalapa, Veracruz, Mexico. 


6 U.S. NATIONAL MUSEUM BULLETIN 266 


Lophoproctinus notandus Silvestri 1949, p. 4. 
TYPE: Location unknown. 
TYPE LOCALITY: San Francisco, Veracruz, Mexico. 


Family POLYXENIDAE 
Genus BARROXENUS Chamberlin 1940a, p. 3. 
TYPE SPECIES: Barroxenus panamanus Chamberlin, by original des- 


ignation. 
Barroxenus panamanus Chamberlin 1940a, p. 3. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone, Panama. 


Genus POLYXENUS Latreille 1802, p. 45. 


TYPE SPECIES: Scolopendra lagura Linnaeus, by monotypy. 
Polyxenus poecilus Chamberlin 1923a, p. 400. 

TYPE: Mus. Calif. Acad. Sci., no. 1256. 

TYPE LOCALITY: Isla Santa Inéz Sur, Golfo de California, Baja Cal- 
ifornia, Mexico. 

PARATYPE LOCALITIES: Puerto Bellandra, Isla Carmen; Isla Mon- 


serrate, Golfo de California, Baja California, Mexico. 


Subclass CHILOGNATHA 
KEY TO SUPERORDERS OF CHILOGNATHA 


1. Gonopods of male modified from one or both pairs of legs at caudal 
end of body; legs of 7thsegment not modified ....... PENTAZONIA 
Gonopods of male modified from at least one pair of legs of 7th seg- 
ment. 
2. First legs of 7th segment of male not modified; eight pairs of legs 
IN PON EOL POHOD Od Siete sjalcrevereraiols celelalelsisierelainiepas COLOBOGNATHA 
First legs of 7th segment of male modified into gonopods; seven 
pairs of legs in front of gonopods......... HELMINTHOMORPHA 


Superorder PENTAZONIA 
KEY TO ORDERS OF PENTAZONIA 


1. Body short, composed of 14 to 16 segments, and contractible into a 
Das sieve sais s 0,2)0 cro jahe natal sgekate: shel ehetetelelheretddMan dentate Rete GLOMERIDA 
Body elongate, composed of 19 to 21 segments, not contractible in- 


Hol a badly 5 ox. ci anc:csctlotet Matete. ste foveete petals atctateteNetavele GLOMERIDESMIDA 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA a 


Order GLOMERIDA 
Family GLOMERIDAE 
Genus GLOMERIS Latreille 1802, p. 44, 


TYPE SPECIES: Oniscus marginatus Villers, by monotypy? 
Glomeris boneti Chamberlin 1943b, p. 69. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Parque Nacional de Zempoala, Morelos, Mexico. 
Causey 1964c, and Hoffman in letter of November 1964 have expressed 
the opinion that this species probably belongs in the genus Glomeroides. 


Genus GLOMEROIDES Chamberlin 1922, p. 60. 
TYPE SPECIES: Glomeroides centralis Chamberlin by original desig- 


nation. 
Glomeroides caecus Causey 1964c, p. 65. 

TYPE: Mus. Comp. Zool. 

TYPE AND PARATYPE LOCALITY: Sétano de Huitzmolotitla, about 
2 km. southwest of Tlamaya, which is 10 km. northeast of Xilitla, San 
Luis Potosi, Mexico. 
Glomeroides centralis Chamberlin 1922, p. 60. 

TYPE? U.S. Nat. Mus: no: 856: 

TYPE LOCALITY: Trece Aguas, Guatemala. 
Glomeroides promiscus Causey 1964c, p. 65. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Sétano de Gémez Farfas, 5 km. east of Gémez 
Farias, Tamaulipas, Mexico. 

PARATYPE LOCALITIES: Unnamed caves nos. 3 and 7, Rancho de 
Cielo, elev. 1150m., 16 km. northwest of Gémez Fartas, Tamaulipas, 
Mexico. 


Order GLOMERIDESMIDA 
Family GLOMERIDESMIDAE 
Genus GLOMERIDESMUS Gervais 1844a, p. 61. 


TYPE SPECIES: Glomeridesmus porcellus Gervais, by monotypy. 
Glomeridesmus barricolens Chamberlin 1940a, p. 4. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone, Panama. 
Glomeridesmus bicolor Loomis 1964a, p. 11. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro, Panama. 

PARATYPE LOCALITY: El Valle, Coclé, Panama. 


8 U.S. NATIONAL MUSEUM BULLETIN 266 


Glomeridesmus centralis Chamberlin 1922, p. 61. 

TYPE: U.S. Nat. Mus., no. 862. , 

TYPE LOCALITY: Actel4, Guatemala. 
Glomeridesmus circularis Loomis 1964a, p. 12. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqu’, Panama. 

PARATYPE LOCALITY: Almirante, Bocas del Toro, Panama. 
Glomeridesmus latus Loomis 1961, p. 78. 

TYPE: U.S. Nat. Mus., no. 2655. 

TYPE LOCALITY: Cerro Campana, Panama. 
Glomeridesmus parvior Chamberlin 1940a, p. 4. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone, Panama. 

OTHER LOCALITY: Pitta area, Canal Zone, Panama, Loomis 1961. 
Glomeridesmus rotundatus Loomis 1964a, p. 13. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro, 


Panama. 


Superorder HELMINTHOMORPHA 
KEY TO ORDERS OF HELMINTHOMORPHA 


1. Body composed of 18 to 22 segments; sternites, pleurites, and ter- 
gites completely fused without trace of sutures; tergites usually 
produced laterally into keels; eyes always absent; second legs of 
7th segment of male not modified as gonopods ..... POLYDESMIDA 
Body composed of 30 or more (rarely 20, 26, or 28) segments; the 
body rings less complete, the sternites being either free or set off 
by distinct sutures; eyes usually present, sometimes lost in cave 
BOE HIS wile fee yelie vol Oe orisl sale ai ast siege lai eile eae aie) irae) sta folchal a tckale) siete ed tee nent 2 
2. Terminal segment of body with 1 to 3 pairs of spinnerets; either one 
or both pairs of legs of 7th segment of males modified into gono- 
pods; sternites free from the pleurotergites, the latter usually with 
Wome tse bales (ate) ecs ee cnioieis auetaueiauscaerelaie nya) cise (eaten een CHORDEUMIDA 
Terminal segment of body without spinnerets; either both pairs of 
legs of 7th segment modified as gonopods, or one pair missing; 
sternites usually fused with pleurotergites ........ccscccceces 3 
3. Stipites of gnathochilarium broadly in contact behind the laminae 
Wim gialless farsa ge o0s) ose +e -ierete oseyar sein aiesose iota, are -ore atee Oe aens JULIDA 
Stipites of gnathochilarium separated for their entire length by the 
mentum and laminae linguales) << s.yof:. cjecae «share cieleinlebaetrionie ee ee 4 

4. Third segment closed ventrally, the 5th segment with a single pair 


Of Ne psig o:c:s: a igiahuhentre o, eters rch sete brea steerer a keke cose tes SPIROBOLIDA 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 2 


Third segment open ventrally, the 4th and following segments closed; 
5th segment with two pairs of legs .......... SPIROSTREPTIDA 


Order POLYDESMIDA 
KEY TO SUBORDERS OF POLYDESMIDA 


1. Opening of sternal gonopod socket more or less constricted between 
the gonopods, the coxae of which are entirely free of each 
Bata a athrnsoe. jara eh Srisen vel wifndalinde tej alten) wysicaye PARADOXOSOMATIDEA 

Opening of sternal gonopod socket not medially constricted; coxae 
of gonopods connected to each other by a ligament or pseudoster- 
nite, or else broadly in contact along the median surfaces. 


POLYDESMIDEA 


Suborder POLYDESMIDEA 


KEY TO MEXICAN AND CENTRAL AMERICAN 
FAMILIES OF POLYDESMIDEA 


1. Body capable of being tightly rolled into a sphere or a thick, later- 
ally compressed disk ........ eet aia airalist el otiat eo) ots fel nied el sianet fee dean wise 2 
Body incapable of being tightly rolled 2.666 0.060 acc coc ca veins cles 5 

2. Pores not on the lateral keels but located high on the sides of the 
dorsum; some of the anterior keels with special flanges on inner 

Sr Ventre | side se... 5.5.0.<% ss’ Srenedetege iets tener etoians DORSOPORIDAE 
Pores, if present, always on the lateral keels; ventral surface of 
keels without special flanges...... ete Regie eee ea telat atl mato lel « 3 

3. Surface of segments strongly hispid, irregularly roughened, tubercu- 
lar, or a combination of these; supplementary margin of segments 
fimbriate or crenate; gonopods with large subglobose coxae par- 
tially enclosing the distal elements .......... CYRTODESMIDAE 
Surface of segments seldom conspicuously hispid or generally great- 

ly roughened, at most with a few broad seried tubercles; supple- 
mentary margin unbroken or lacking; distal elements of gonopods 
rising at or near the apex of the relatively small non-inclusive 

EP ORAE inp iw sin.n ois, «.0' «2 e4sjere wieles\e' Seieis ofan ese aisieynis ymin iene atin a hake 4 

4. Posterior portion of segments usually sharply marked off into elon- 
gate rectangular areas; pores obviously present at or near middle 

Saas eee a Sella! afer eta oi e's bie, o1'afnn wl cette ia oda sbasgs - ONISCODESMIDAE 
Segments lacking posterior rectangular areas; pores absent or minute 
and obscure, partially hidden by overlapping metazonite of preced- 

MM SER MCE cn enn nie > oc evap aie x neue a muna, eaehae SPHAERIODESMIDAE 

5. Segment 1 with anterior margin expanded forward over head, its mar- 


gin divided into rounded lobes, serrations, or subquadrate areas .6 


10 


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U.S. NATIONAL MUSEUM BULLETIN 266 


Segment 1 with anterior margin slightly or not expanded forward and 
not divided into lobes, serrations, or quadrate areaS ...-seeeee no 
Pores opening through special tubercles on or near the outer mar- 
gin of the keels ......seeeeeeescees eiereiei=tet= = STYLODESMIDAE 
Pores not on special tubercles but opening through channels in outer 
margin of keels or from their upper surface ...... avettellet aol sient 
Anterior border of segment 1 and outer margins of keels transparent, 
with opaque intrusions from inner surface ..... CRYPTODESMIDAE 
Margins of segment | and of keels opaque, similar to inner surface .8 
Outer and posterior margins of keels with prominent seta-bearing 
Teeth. st. a: Seen eee de oa shes cea te cetee SE RIDON LOD SM uouarE 
Outer and posterior margins of keels usually somewhat lobed but not 
prominently serrate.....esccececcecseeseees CH YTODESMIDAE 
Seminal canal of male gonopod widening into a distinct vesicle or 
ampulla near its distal extremity ......+e+.2++++ POLYDESMIDAE 
Seminal canal not distally widened or modified as above ........10 
Lateral keels with pores on ventral surface; outer margins developed 
into two or three large acute lobes; outer margin of segment 1 lobed 
somewhat, like keels sri< ie ctcliasts soe esivionlasielg. | RIDONTOMIDAE 
Keels with pores on outer edge or dorsal surface; outer margins not 
developed into large lobes; segment 1 with outer margins not spe- 
Giallly lobed! ‘leis oe ois /e ein elie ara ale etel eleieleier ele? exatereatete chore ete tallaieem el 
Body shape suggestive of Polydesmus; number of segments 18-21; 
living animals very active............«! RICHOPOLYDESMIDAE 
Body shape not suggestive of Polydesmus; segments 20, 19 in males 
of Hypsiloporus (Comodesmidae); animals rather sluggish......1é 
Body small, 10 mm. or less long; sides of segment 2 extending be- 
low those of segments 1 and 3; dorsum setose; coxae of gonopods 
large and enclosing much of distal joints......... COMODESMIDAE 
Size medium to large; segment 2 not extending below segments 1 and 
3; dorsum rarely with conspicuous setae; coxae of gonopods not 
enlatged or enclosing distal) joantsies + ete s/s) elatelale)o}ale el niateisielataiaiale 
Pores on dorsal surface of keels which have rather thin, unrimmed 
outer margins that are irregularly dentate or tubercular. 
PLAT YRHACIDAE 
Pores opening from more or less thickened outer rim of the keels 
which may have several small teeth but is not otherwise irregular 
OL UUDETCUL AL. ein! <\'a! ais! a\is'51 4, siellsiiove oy's).o) syst elelrt eal </s) sfolein elmicitsleloleieneterenenen eee 
Last segment broadly rounded behind............++.-H®#URYURIDAE 
Past sepment comicdlaat rape kre etstl-Valerese( vets sia oes miei tele etateretarenatentlee 
Stigmata displaced anteriorly above the legs so that the posterior 
stigma is located above the anterior coxa, and the anterior stigma 
is placed in the transverse segmental constriction in contact with 
THE. PYOZOMILE ccc oc socio sic's oleles 0 etiam sre eo td Olal om © PA a Tennant 


Stigmata not displaced, occupying the normal position ..........16 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 11 


16.Gonopod coxa without solenite; telopodite with large oval or elon- 
gate cavity in mesal face containing many setae. 
RHACHODESMIDAE 
Gonopod coxa with solenite which projects into a narrow groove at 
ee tee PAROLE pics’ w winfaia a's oie is tales wliche mis uw ck miele aia ean Sw nel 7 
17. Usually broad, compact forms, the W/L ratio usually 18% or greater, 
body widest near middle in males, near caudal end in females; 
sensory cones of antennae not separated or paired; 6th antennal 
joint without an external sensory pit; solenite of gonopod coxa 
originating from a circular socket well removed from distal edge 
of coxa; prefemora of walking legs normally with acute distal 
SEEING usta lento) Se) a(n! © albania Bin)'e. nie,9 5 2 oie eceeee- XYSTODESMIDAE 
Narrower forms, typically with reduced and well-separated keels, the 
W/L ratio usually 18% or less, body usually widest near head in 
both sexes; sensory cones of antennae usually separated into two 
pairs in larger forms; 6th joint with a prominent external distal 
sensory pit; solenite of gonopod coxa located in a notch on distal 
edge of coxa; prefemora of walking legs normally without acute 


SABES 2) olen eure sien alee coc cees Siu wets es 8 wie’ CHELODESMIDAE 


Family CHELODESMIDAE 
Genus ALOCODESMUS Silvestri 1896a, p. 5. 


TYPE SPECIES: Alocodesmus angustatus Silvestri, by monotypy. 
Alocodesmus angustatus Silvestri 1896a, p. 5. 

Alocodesmus dromeus Chamberlin 1922, p. 48; Loomis 1964a, p. 16. 

TYPE: Torino Mus. 

TYPE LOCALITY: Given as Darien, but Carl 1914, p. 897, says the 
holotype came from Punta Sabana, Darien Prov., Panama. 
Alocodesmus silvestri (Attems). 

Leptodesmus (Desmoleptus) silvestri Attems 1931, p. 23; Loomis 

1964a, p. 130. 
TYPE: Male and female syntypes in Naturh. Mus. Wien. (H) 
TYPE LOCALITY: Panama. 


Genus ATYLOPHOR Chamberlin 1922, p. 49. 
TYPE SPECIES: Atylophor rafaelanus Chamberlin, by original desig- 


nation. 
Atylophor rafaelanus Chamberlin 1922, p. 50. 

TYPE: U.S. Nat. Mus., no. 849. 

TYPE LOCALITY: San Rafael, Guatemala. There are two localities 
by this name, the smaller is about 15 km. west of Guatemala City, the 
other about 30 km. southwest of Jalapa, and it is not known which is the 
type locality. 


273-687 O - 68 - 2 


12 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus CHONDRODESMUS Silvestri 1897b, p. 13. 


Dirhabdophallus Pocock 1909, p. 161; Carl 1914, p. 886. 

TYPE SPECIES: Chondrodesmus armatus Silvestri, by monotypy. 
Chondrodesmus acanthurus (Peters). 

Polydesmus (Oxyurus) acanthurus Peters 1864, p. 532; Carl 1914, 

p. 972. 

TYPE: Probably Zool. Mus. Berlin. 

TYPE LOCALITY: ‘‘Veragua’’ (probably Veraguas Prov.), Panama. 
Chondrodesmus acuticollis Attems 1931, p. 47. 

TYPE: Depository not given. 

TYPE LOCALITY: Acajutla, El Salvador. 

OTHER LOCALITIES: Bebedero, Costa Rica (Attems 1933). In 7 de- 
partments of E] Salvador (Kraus 1954b). 

Chondrodesmus alidens Chamberlin 1922, p. 46. 

TYPE: US. Nat. Mus. no. oo. 

T YPE LOCALITY: La Ceiba, Honduras. 
Chondrodesmus allenae Loomis 1959p, p. 2. 

TYPE: U.S. Nat. Mus., no. 2666. 

TYPE LOCALITY: Escuela Agricola Panamericana, Zamorano, near 
Tegucigalpa, Honduras. 

Original spelling of the species name was ‘‘alleni’’ but R. L. Hoffman 
pointed out that since the species was named for Mrs. Paul Allen the 
name should be feminine. 

Chondrodesmus atrophus Chamberlin 1925, p. 42. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Ft. Sherman, Canal Zone. 

OTHER LOCALITIES: Monte Lirio, Canal Zone; Juan Dias, Panama 
Prov., Panama. 

Chondrodesmus ensiger (Pocock). 

Dirhabdophallus ensiger Pocock 1909, p. 165; Attems 1938, p. 86. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 

PARATYPE LOCALITY: Xautipa, Guerrero, Mexico. 

Chondrodesmus euliotus Chamberlin 1933, p. 21. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Chitaria, Costa Rica. 
Chondrodesmus falciphallus Chamberlin 1933, p. 20. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Parismina, Costa Rica. 

Chondrodesmus frauenfeldianus (Humbert and Saussure). 

Polydesmus frawenfeldianus Humbert and Saussure 1870, p. 172. 

Leptodesmus frauenfeldianus Silvestri 1896a, p.5; Carl 1914, p. 972. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Colombia. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 13 


OTHER LOCALITIES: Punta Sabana; forests of the rivers Cianati and 
Lara, Darien Prov., Panama (Silvestri 1896a). 
Chondrodesmus granosus (Carl). 

Leptodesmus plataleus granosus Car] 1902, p. 602. 

D irhabdophallus granosus Pocock 1909, p.164; Chamberlin 1922, p.47. 

T YPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: San José, Costa Rica. 

OTHER LOCALITIES: La Uruca (Pocock 1909); Hamburg Farm, Limén, 
Costa Rica. 
Chondrodesmus hoffmanni (Peters). 

Rhacophorus hoffmanni Peters 1864, p. 537. 

Dirhabdophallus (?) hoffmanni Pocock 1909, p. 165; Chamberlin 1922, 

p. 48. 

TYPE: Zool. Mus. Berlin (H) 

EYPE LOCALITY : Costa. Rica: 
Chondrodesmus montanus (Pocock). 

Dirhabdophallus montanus Pocock 1909, p. 163; Chamberlin 1922, p. 47. 

TYPE: British Mus. 

TYPE LOCALITY: Volcan de Agua, Guatemala. 

OTHER LOCALITIES: Five localities in 3 departments of El Sal- 
vador (Kraus 1954b). 
Chondrodesmus murphyi Chamberlin 1946, p. 147. 

TYPE: American Mus. Nat. Hist. 

TYPE LOCALITY: Isla San José, Pearl Archipelago, Panama. 
Chondrodesmus nannus Chamberlin 1943b, p. 58. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chilpancingo, Guerrero, Mexico. 
Chondrodesmus orientalis Kraus 1954b, p. 309. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Conchagua, Dept. La Union, EF] Salvador. 

PARATYPE LOCALITIES: Same, and La Union, Dept. La Union; 
Laguna Aramuaca and Laguna de Jocotal, Dept. San Miguel, E1 Salvador. 
Chondrodesmus panamenus Chamberlin 1922, p. 46. 

Chondrodesmus panamenus Chamberlin 1925, p. 42. 

Chondrodesmus chamberlini Hoffman 1950b, p. 70; Loomis 1964a, p. 16. 

TYPE: Mus. Comp. Zool., cat. no. 5023. 

TYPE AND PARATYPE LOCALITY: Panama. 

OTHER LOCALITY: Barro Colorado Island, Canal Zone (Loomis 1961). 
Chondrodesmus pittieri Loomis 1961, p. 83. 

TYPE: U.S. Nat. Mus., no. 2635. 

TYPE LOCALITY: Puerto Obaldia, Panama; (probably Coclé rather 
than San Blas Province). 
Chondrodesmus rodriguezi (Brolemann). 

L eptodesmus rodriguezi Brolemann 1900, p. 103. 


14 U.S. NATIONAL MUSEUM BULLETIN 266 


Dirhabdophallus rodriguezi Pocock 1909, p. 164; Chamberlin 1922, p. 47. 

C hondrodesmus nicaraguae Chamberlin 1924, p. 174; Loomis 1950, p. 165. 

TYPE: Mus. Nat. Hist. Natur. Paris (H) 

TYPE LOCALITY: Guatemala. 

OTHER LOCALITIES: San José, and Surubres, near San Mateo, Costa 
Rica (Brolemann 1905); Purulha, Guatemala (Pocock 1909); Nicaragua?, 
collected in bananas from Nicaragua at Port of New York (Chamberlin 
1924). 

Chondrodesmus sabachanus Chamberlin 1938a, p. 174. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Sabacha Cave, Tekax, Yucatan, Mexico. 

OTHER LOCALITY: Mugeres Island, Yucatan, Mexico (Chamberlin 
1952a). Probably Isla Mujeres, near Puerto Juarez, Yucatan, Mexico. 
Chondrodesmus singularis Chamberlin 1922, p. 44. 

TYPE: Mus. Comp. Zool., cat. no. 5022. 

TYPE LOCALITY: Puerto Limén, Costa Rica. 

The specimen from Puerto Limén, Costa Rica, that Attems 1901, p.85, 
identified as his South American Leptodesmus carinovatus (see Pocock 
1909, p.165, and Jeekel 1963a, p.127) more probably was C. singularis. 
Chondrodesmus spatulatus (Pocock). 

Dirhabdophallus spatulatus Pocock 1909, p.165; Attems 1938, p. 86. 

TYPE: British Mus. 

T YPE LOCALITY: Omilteme, Guerrero, Mexico. 

Chondrodesmus tuberculifer Chamberlin 1922, p. 45. 

TYPE: U.S. Nat. Mus., no. 847. 

TYPE LOCALITY: La Ceiba, Honduras. 


Genus ETHOPHALLUS Chamberlin 1933, p. 19, 
TYPE SPECIES: Ethophallus cervantes Chamberlin, by original des- 


ignation. 
Ethophallus cervantes Chamberlin 1933, p. 20. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Cervantes, Costa Rica. 


Genus EUTYPORACHIS Pocock 1909, p. 168. 
TYPE SPECIES: Eutyporachis tessellatus Pocock, by original desig- 


nation. 
Eutyporachis oltramarei (Carl). 
Leptodesmus oltramarei Carl 1902, p. 600. 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE LOCALITY: Guatemala. 
Eutyporachis tessellatus Pocock 1909, p. 169. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Senahu, Alta Verapaz, Guate- 


mala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 15 


Genus ISIDRONA Attems 1933, p. 267. 
TYPE SPECIES: /sidrona forficula Attems, by monotypy. 
Isidrona forficula Attems 1933, p. 268. 
T YPE: Natur. Mus. Wien. (H) 
TYPE LOCALITY: San Isidro, Costa Rica. 


Genus PHYLACTOPHALLUS Pocock 1909, p. 166. 


Allarithmus Attems 1933, p. 265. New synonymy, suggested by R.L. 
Hoffman. 

TYPE SPECIES: Phylactophallus stenomerus Pocock, by original des- 
ignation. 
Phylactophallus parvulus (Attems). 

Allarithmus parvulus Attems 1933, p. 265. 

T YPE: Natur. Mus. Wien. (H) 

T YPE LOCALITY: Irazu, 2,200 m., Costa Rica. 

PARATYPE LOCALITIES: LaCarpintera and San Isidro, Costa Rica. 
Phylactophallus stenomerus Pocock 1909, p. 166. 

TYPE: British Mus. 

TYPE LOCALITY: Irazu, Costa Rica. 


Genus PROLETUS Attems 1931, p. 36. 
TYPE SPECIES: Proletus foetidus Attems, by monotypy. 
Proletus foetidus Attems 1931, p. 36. 
TYPE: Natur. Mus. Wien. (H) 
TYPE LOCALITY: Panama. 


Genus SCHISTIDES Chamberlin 1922, p. 51. 
TYPE SPECIES: Schistides atopophallus Chamberlin, by original des- 


ignation. 

Schistides atopophallus Chamberlin 1922, p. 51. 
T YPE: U.S. Nat. Mus., no. 850. 
TYPE LOCALITY: San Juan Puebla, Honduras. 


Genus SOLAENORHABDUS Hoffman 1950b, p. 70. 
TYPE SPECIES: Cyclorhabdus contortus Brolemann, by original des- 


ignation. 
Solaenorhabdus contortus (Brolemann). 

Cyclorhabdus contortus Brolemann 1900, p. 98. 

Leptodesmus contortus Car|1902, p.607; Hoffman 1950b, p. 70. 

Habrodesmus uncinatus Kraus 1954b, p. 307; new synonymy suggested 
by R. L. Hoffman and approved by O. Kraus. 

TYPE: Mus. Nat. Hist. Natur., Paris (H) 

TYPE LOCALITY: Guatemala. 

OTHER LOCALITY: Finca San Jorge, near Santa Ana, Dept. Santa 
Ana, El Salvador (Kraus 1954). 


16 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus TRICHOMORPHA Silvestri 1897c, p. 11. 


Typophallus Chamberlin 1940a, p. 10: Loomis 1964a, p. 17. 

Desmacrides Chamberlin 1940a, p. 11; Loomis 1964a, p. 17. 

TYPE SPECIES: Trichomorpha elegans Silvestri, by monotypy. 
Trichomorpha capillata Loomis 1964a, p. 17. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: West of Finca Palo Santo, near Nueva California, 
Chiriqui Prov., Panama. 

PARATYPE LOCALITIES: Barca area and Casita Alta, Finca Lerida, 
near Boquete, Chiriqui Prov.; Chilibrillo Cave, near Chilibre, Panama 
Prov., Panama. 


Trichomorpha evidens (Chamberlin). 

Typophallus evidens Chamberlin 1940a, p. 11; Loomis 1964a, p.18. 

Trichomorpha fratrellus Chamberlin 1947c, p.64; Loomis 1964a, p. 18. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

OTHER LOCALITIES: Summit, Canal Zone (Chamberlin 1947c); ‘‘Pa- 
cific Side,’’ and Barro Colorado Island, Canal Zone (Loomis 1961). 
Trichomorpha folium (Brolemann). New Combination. 

Leptodesmus folium Brolemann 1903, p. 139. 

Camptomorpha folium Attems 1938, p. 73. 

TYPE: Mus. Nat. Hist. Natur., Paris (H) 

TYPE LOCALITY: Cocos Island, Costa Rica. 

Trichomorpha nidicola Chamberlin 1925, p. 42. 

Trichomorpha extrema Chamberlin 1925, p. 43; Loomis 1964a, p.18. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Barro Colorado, Canal Zone. 

OTHER LOCALITY: Same (Loomis 1961). 

Trichomorpha panamica Chamberlin 1925, p. 43. 

T richomorpha recta Chamberlin 1925, p. 43; Loomis 1964a, p. 19. 

Desmacrides dichrus Chamberlin 1940a, p.12; Loomis 1964a, p.19. 

T richomorpha kraussi Chamberlin 1947c, p.63; Loomis 1961, p. 84. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 

OTHER LOCALITIES: Same (Chamberlin 1940a, Loomis 1961); Sum- 
mit (Chamberlin 1947c); Frijoles, and Piffa area (Loomis 1961), Canal 
Zone. El Valle, Coclé Prov., and Cerro Campana, Panama Prov., Pana- 
ma (Loomis 1961). 

Trichomorpha tacarcuna Loomis 1964b, p. 182. 
TYPE: U.S. Nat. Mus., no. 2983. 
TYPE AND PARATYPE LOCALITY: Rio Tacarcuna, elev. 1,900 ft., 


Darien Prov., Panama. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 17 


Family CHYTODESMIDAE 
Genus DOCODESMIELLA Loomis 1961, p. 80. 
TYPE SPECIES: Docodesmiella insularis Loomis, by original desig- 


nation. 
Docodesmiella insularis Loomis 1961, p. 81. 

TYPE: U.S. Nat. Mus., no. 2634. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 
Docodesmiella magna Loomis 1964a, p. 20. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui Prov., Pan- 
ama. 

PARATYPE LOCALITIES: Same, and Cerro Punta, Chiriqui Prov.; 
Almirante, Bocas del Toro Prov., Panama. 
Docodesmiella spathulata Loomis 1964a, p. 21. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: ‘‘Casita Alta,’’ Finca Lerida, near Boquete, Chi- 
riqut Prov., Panama. 

PARATYPE LOCALITIES: Same, and other nearby areas; near Nueva 


California, Chiriqui Prov., Panama. 


Genus PTYXODESMUS Loomis 1964a, p. 24. 
TYPE SPECIES: Ptyxodesmus spiculatus Loomis, by original desig- 


nation. 
Ptyxodesmus spiculatus Loomis 1964a, p. 24. 
T YPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Cerro Azul, near ‘‘Goofy Lake,’’ Panama Prov., 


Panama. 
Genus TRICHODESMIELLA Loomis 1964a, p. 25. 
TYPE SPECIES: Trichodesmiella multiloba Loomis, by original des- 


ignation. 
Trichodesmiella multiloba Loomis 1964a, p. 26. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui Prov., Pan- 
ama. 


PARATYPE LOCALITIES: Same and near by. 


Family COMODESMIDAE 
Genus HYPSILOPORUS Loomis 1961, p. 104. 


TYPE SPECIES: Hypsiloporus proclivis Loomis, by original designa- 


tion. 


18 U.S. NATIONAL MUSEUM BULLETIN 266 


Hypsiloporus erosus Loomis 1964a, p. 27. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: El Valle, Coclé Prov., Panama. 
Hypsiloporus montanus Loomis 1964a, p. 28. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: ‘‘Casita Alta,’’ Finca Lerida, near Boquete, Chi- 
riqui Prov., Panama. 

PARATYPE LOCALITIES: Same, and Cerro Punta; Finca Palo Santo, 
near Nueva California, Chiriqui Prov., Panama. 
Hypsiloporus proclivis Loomis 1961, p. 105. 

TYPE: U.S. Nat. Mus., no. 2645. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

OTHER LOCALITIES: Same, and Madden Lake Dam, Panama Prov., 
Panama. 
Hypsiloporus serratus Loomis 1964a, p. 29. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 


Panama. 
Genus INODESMUS Cook 1896, p. 25. 


Lasiodesmus Silvestri 1908, p. 575; Loomis 1934, p. 64. 

TYPE SPECIES: /nodesmus jamaicensis Cook, by monotypy. 
Inodesmus globulosus Loomis 1964a, p. 30. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Las Cumbres, Panama Prov., 


Panama. 


Family CRYPTODESMIDAE 
Genus DIAPHANACME Loomis 1964a, p. 31. 


TYPE SPECIES: Diaphanacme latus Loomis, by original designation. 
Diaphanacme latus Loomis 1964a, p. 32. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 

PARATYPE LOCALITIES: Same, and Almirante, Bocas del Toro Prov., 


Panama. 


Family CYRTODESMIDAE 
Genus CRYPTURODESMUS Silvestri 1897a, p. 1. 


Katantodesmus Attems 1900, p. 385; Pocock 1909, p. 116. 

TYPE SPECIES: Crypturodesmus targioni Silvestri, by monotypy. 
Crypturodesmus targioni Silvestri 1897a, p. 1. 

TYPE: Mus. Zool. Torino. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 19 


TYPE LOCALITY: Mexico. 

In the copy of Silvestri’s paper (loc. cit.), which he sent me, he had 
canceled ‘‘Messico’’ in the title and in the species habitat as well, and 
had written “‘Brasile’’ in both places. No other collections of the spe- 
cies have been reported and it seems most likely that it is truly Brazil- 
ian. 


Genus CYRTODESMUS Gervais 1847, p. 92. 


Oncodesmus Cook 1896, p. 28. 

Cyliocyrtus Cook 1898, p. 458, in part. 

T rigonostylus Brolemann 1898a, p. 273, in part. 

Peltedo Chamberlin 1940a, p. 8; Loomis 1964a, p. 34. 

Oncodesmoides Kraus 1954a, p. 41; Loomis 1964a, p. 34. 

TYPE SPECIES: Polydesmus velutinus Gervais and Goudot, by mono- 
typy- 

Cyrtodesmus archimedes (Chamberlin). 

Peltedo archimedes Chamberlin 1940a, p. 8. 

Oncodesmoides angulatus Loomis 1961, p. 87; Loomis 1964a, p. 37. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 

OTHER LOCALITIES: Canal Zone: Fort Sherman; Frijoles; Pifva area 
(Loomis 1961). Panama: Almirante, Bocas del Toro Prov.; El Valle, 
Coclé Prov.; Juan Dias, Panama Prov. (Loomis 1961). 

Cyrtodesmus bicolor Loomis 1964a, p. 37. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: West of Finca Palo Santo, near 
Nueva California, Chiriqui Prov., Panama. 

Cyrtodesmus bifurcus Loomis 1964a, p. 38. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 
Panama. 

Cyrtodesmus confluentus Loomis 1964a, p. 40. 

TYPE: U.S. Nat. Mus., no. 2801. 

TYPE LOCALITY: Changuinola district, Bocas del Toro Prov., Pan- 
ama. 

Cyrtodesmus coronatus Loomis 1964a, p. 42 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 
Panama. 

Cyrtodesmus dentatus Loomis 1964a, p. 43. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui Prov., Pan- 


ama. 


20 U.S. NATIONAL MUSEUM BULLETIN 266 


Cyrtodesmus depilis Loomis 1964a, p. 44. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Finca Lerida, near Boquete, 
Chiriqut Prov., Panama. 

Cyrtodesmus granosus (Gervais and Goudot). 

Polydesmus granosus Gervais and Goudot 1844, p. 28. 

Cyrtodesmus granosus Gervais 1847, p. 93. 

Oncodesmus granosus Cook 1896, p. 28. 

TYPE: British Mus., but not found in recent years. 

TYPE LOCALITY: Colombia. 

Although reported from Darien Prov., Panama, it is doubtful that the 
identification was correct or that the species can be reidentified (see 
Loomis 1964a, p. 34). 

Cyrtodesmus hispidulosus Loomis 1964a, p. 46. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Palo Santo, near Nueva California, Chiriqut 
Prov., Panama. 

PARATYPE LOCALITIES: Ten mi. south of El Volcan, and at Finca 
Lerida, near Boquete, Chiriqur Prov., Panama. 

Cyrtodesmus lyrapes Loomis 1964a, p. 47. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 

PARATYPE LOCALITIES: Same, and west of Finca Palo Santo, near 
Nueva California, Chirigqui Prov., Panama. 

Cyrtodesmus macrosetosus Loomis 1964a, p. 48. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui Prov., Pan- 
ama. 

PARATYPE LOCALITIES: Same and adjacent regions. 

Cyrtodesmus mundus Loomis 1964a, p. 49. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui. Prov., Pan- 
ama. 


PARATYPE LOCALITIES: Same and adjacent regions. 


Family DORSOPORIDAE 
Genus DORSOPORUS Loomis 1958, p. 236. 


TYPE SPECIES: Dorsoporus barroensis Loomis, by original designa- 
tion. 
Dorsoporus barroensis Loomis 1958, p. 237. 

TYPE2U-S. Nat: Mus., no. 2511. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 21 


Family EURYURIDAE 
Genus AMPLINUS Attems 1899, p. 281. 


Pycnotropis Carl 1914, p. 932. 

Seminellogon Chamberlin 1933, p. 18. 

Phinotropis, Ptyxogon and Amydrinus Chamberlin 1941le, pp. 499-500. 

(These are Peruvian genera.) 

Sigmogonotropis Hoffman 1951, p. 240; Jeekel 1963a, p. 73. 

TYPE SPECIES: Pachyurus (Amplinus) kalonotus Attems, by designa- 
tion of Pocock 1909. 
Amplinus cerroazulensis (Hoffman). 

Seminellogon cerroazulensis Hoffman 1954a, p. 54; Jeekel1963a, p.75. 

TYPE: U.S. Nat. Mus., no. 2100. 

TYPE LOCALITY: Cerro Azul, near ‘‘Goofy Lake,’’ Panama Prov., 
Panama. 

PARATYPE LOCALITIES: Boquete, and El Volcan, Chiriqut Prov., 
Panama. 
Amplinus chitarianus (Chamberlin). 

Seminellogon chitarianus Chamberlin 1933, p. 19; Jeekel 1963a, p. 75. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chitaria, Costa Rica. 
Amplinus latzeli (Attems). 

Pycnotropis latzeli Attems 1931, p. 79; Jeekel 1963a, p. 75. 

TYPE: Naturh. Mus. Wien (H). 

TYPE LOCALITY: Panama. 
Amplinus magnus (Loomis). New combination. 

Seminellogon magnus Loomis 1964a, p. 54. 

T YPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 
Amplinus panamicus (Chamberlin). 

Aphelidesmus panamicus Chamberlin 1925, p. 41. 

Seminellogon panamicus Loomis 1961, p. 86. 

Sigmogonotropis serratus Hoffman 1951, p. 240; Jeekel 1963a, p. 75. 

Seminellogon serratus Loomis 1961, p. 86. New synonymy, suggested 

by R. L. Hoffman. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Headwaters of Rio Chinilla, Canal Zone. 

OTHER LOCALITIES: Barro Colorado Island; Gorgona; Monte Lirio; 
Pifta area, Canal Zone (Loomis 1961). Cana, Darien Prov. (Hoffman 1951); 
Cerro Campana, Panama Prov. (Loomis 1961), Panama. 


Genus APHELIDESMUS Brolemann 1898b, p. 322. 


Trachelorachis Silvestri 1898a, p. 5. 
Trachelacanthe Berg 1899, p. 77. 


22 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE SPECIES: Aphelidesmus hermaphroditus Brolemann, by mono- 
typy- 

Aphelidesmus calverti Chamberlin 1914, p. 191. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: La Emilia, Costa Rica. 

OTHER LOCALITIES: Puerto Limén (Chamberlin 1922), and Paris- 
mina, Costa Rica (Chamberlin 1933). 

Aphelidesmus glaphyros (Attems). 

Euryurus glaphyros Attems 1900, p. 279; Pocock 1909, p. 158. 

TYPE: Naturh. Mus. Wien. 

TYPE LOCALITY: Costa Rica. 

OTHER LOCALITIES: Carrillo; Cuesta del Tablazo; and Cariblanco, 
Costa Rica (Brolemann 1905). 

Aphelidesmus intermedius Chamberlin 1914, p. 192. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: Juan Vifias, Costa Rica. 

OTHER LOCALITY: San Jose, Costa Rica (Chamberlin 1933), in which 
paper he listed the type locality as La Emilia, actually the type locality 
of his A. calverti. 

Aphelidesmus taurinus Loomis 1964a, p. 51. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 


Panama. 


Genus POLYLEPISCUS Pocock 1909, p. 154. 


Euplinus Chamberlin 1952a, p. 578; Hoffman 1954a, p. 52. 

TYPE SPECIES: Polylepiscus stolli Pocock, by original designation. 
Polylepiscus actaeon Pocock 1909, p. 155. 

TYPE: British Mus. 

TYPE LOCALITY: Guatemala. 
Polylepiscus burgeri Causey 1954, p. 65. 

TYPE: Illinois St. Nat. Hist. Surv., Urbana. 

TYPE LOCALITY: Seventeen mi. south of Simojovel, Chiapas, Mex- 
ico. 

PARATYPE LOCALITY: Fifteen mi. south of Pichucalco, Chiapas, 
Mexico. 
Polylepiscus campanulae Hoffman 1962, p. 137. 

TYPE: U.S. Nat. Mus., no. 2776-D617. 

TYPE AND PARATYPE LOCALITY: Seven mi. east of San Cristébal 
de los Casas, on Rt. 190, Chiapas, Mexico. 
Polylepiscus furcifer Pocock 1909, p. 156. 

Euplinus volcanicola Chamberlin 1952a, p.580; Hoffman 1954a, p. 53. 

TYPE: British Mus. 

TYPE LOCALITY: Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 23 


OTHER LOCALITIES: Volcan Tajumulco, and El Porudnir, San Mar- 
cos, Guatemala (Chamberlin 1952a). 
Polylepiscus heterosculptus (Carl). 

Pachyurus heterosculptus Carl 1902, p. 635; Pocock 1909, prlii, 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Guatemala. 

Polylepiscus heterosculptus pococki Hoffman 1954a, p. 53. 

TYPE: U.S. Nat. Mus., no. 2099, 

TYPE LOCALITY: Tumbala, Chiapas, Mexico. 

Polylepiscus major (Chamberlin). 

Aphelidesmus major Chamberlin 1952a, p. 581. New combination, sug- 
gested by R. L. Hoffman, who stated “‘there is no doubt it belongs 
in Polylepiscus as the 3rd species without polygonal areas.”’ 

TYPE: Chicago Mus. Nat. Hist. 

TYPE LOCALITY: Escobas, opposite Puerto Barrios, Guatemala. 

Polylepiscus stolli Pocock 1909, p. 155. 
TYPE: British Mus. 
TYPE LOCALITY: Cholhuitz, Guatemala. 
Polylepiscus trimaculatus Hoffman 1954a, p. 53. 
TYPE: U.S. Nat. Mus., no. 2098. 
TYPE AND PARATYPE LOCALITY: Sepacuite, Guatemala. 


Genus PSEUDAMPLINUS Hoffman 1954a, p. 51. 
TYPE SPECIES: Amplinus orphnius Chamberlin, by original designa- 


tion. 
Pseudamplinus areatus (Pocock). 
Amplinus areatus Pocock 1909, p. 151. 
Orthomorpha areatus Cook 1911c, p. 627; Hoffman 1954a, petals 
T YPE: British Mus. 
TYPE LOCALITY: Retalhuleu, Guatemala. 
P seudamplinus armatus (Pocock). 
Amplinus armatus Pocock 1909, p. 153. 
Orthomorpha armatus Cook 1911c, p. 627; Hoffman 1954a, p. 51. 
TYPE: British Mus. 
TYPE LOCALITY: Mexico. 
Pseudamplinus constrictus (Chamberlin). New combination. 
Amplinus constrictus Chamberlin 1953b, p. 67. 
TYPE: Chicago Mus. Nat. Hist. 
TYPE AND PARATYPE LOCALITY: West of San Pedro Sula, Honduras. 
Pseudamplinus convexus (Carl). 
Pachyurus convexus Carl 1902, p. 633. 
Amplinus convexus Pocock 1909, p. 150. 
Orthomorpha convexus Cook 1911c, p. 628; Hoffman 1954a, p. 51. 
TYPE: Mus. Hist. Nat. Geneve (H) 


24 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE LOCALITY: Costa Rica. 

OTHER LOCALITY: Parismina, Costa Rica (Chamberlin 1933). 
Pseudamplinus crenus (Chamberlin). 

Amplinus crenus Chamberlin 1943b, p. 56; Hoffman 1954a, p. 51. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Near Fortin, Veracruz, Mexico. Probably Fortin 
de las Flores. 

PARATYPE LOCALITY: Same, and Puroje, Veracruz, Mexico. 
Pseudamplinus erichsoni (Brandt). 

Polydesmus erichsoni Brandt 1841, p. 135. 

Polydesmus (Paradesmus) erichsoni Saussure 1860, p. 293: 

Polydesmus (Euryurus) erichsoni Peters 1864, p. 542. 

Amplinus erichsoni Pocock 1909, p. 152. 

Orthomorpha erichsoni Cook 1911c, p. 627; Hoffman 1954a, p. 51. 

TYPE: Zool. Mus. Berlin (H) 

TYPE LOCALITY: Mexico. 

Pseudamplinus eutypus (Chamberlin). 

Amplinus eutypus Chamberlin 1952a, p. 573; Hoffman 1954a, p. 51. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: El Fortin, Veracruz, Mexico. 
Probably Fortin de las Flores. 

Chamberlin founded this species on three females with no mention be- 
ing made of any male characters. However, he refers to Figure 35, which 
is of a male gonopod! Figure 36 is said to be of a new species which he 
does not describe. It is possible that both figures are of this undescribed 
species. 

Pseudamplinus flavicornis (Pocock). 

Amplinus flavicornis Pocock 1909, p. 151. 

Orthomorpha flavicornis Cook 1911c, p. 627; Hoffman 1954a, p. 51. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Central America. 

OTHER LOCALITY: Palenque, Chiapas, Mexico (Hoffman 1951). 
Pseudamplinus intermittens (Causey). 

Amplinus intermittens Causey 1954, p. 64; Hoffman 1954a, p. 51. 

TYPE: Illinois St. Nat. Hist. Surv., Urbana. 

TYPE LOCALITY: Fifteen mi. south of Pichucalco, Chiapas, Mexico. 
Pseudamplinus klugi (Brandt). 

Polydesmus klugi Brandt 1841, p. 133. 

Polydesmus (Paradesmus) klugi Saussure 1860, p. 293. 

Polydesmus (Euryurus) klugi Peters 1864, p. 541. 

Polydesmus (Pachyurus) klugi Saussure and Humbert 1872, p. 27. 

Pachyurus klugi Karsch 1881, p. 37. 

Polydesmus (Paradesmus) picteti Saussure 1859, p. 325. 

Amplinus klugi Pocock 1909, p. 152; Hoffman 1954a, p. 51. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 25 


TYPE: Zool. Mus. Berlin 

TYPE LOCALITY: Alvarado, Veracruz, Mexico. 

OTHER LOCALITIES: Cordoba; Orizaba; Panuco; Anahuac; entire east 
slope of plateau (Saussure 1860); Guanajuato; Jalapa (Pocock 1909); 
Chapulhuacan, Hidalgo (Chamberlin 1943b); La Ruina de Ventura, near 
Santa Rosa, Isthmus of Tehuantepec, Mexico (Chamberlin 1947b). 
Pseudamplinus leon (Chamberlin). 

Amplinus leon Chamberlin 1952a, p. 574; Hoffman 1954a, Deol. 

TYPE: Chicago Mus. Nat. Hist. 

TYPE LOCALITY: Monterrey, Nuevo Ledn, Mexico. 

Pseudamplinus manni (Chamberlin). 

Amplinus manni Chamberlin 1922, p. 41; Hoffman 1954a, p. 51. 

TYPE: U.S. Nat. Mus., no. 844. 

TYPE LOCALITY: La Ceiba, Honduras 

PARATYPE LOCALITY: Lombardia, Honduras. 

Pseudamplinus mimus (Chamberlin). 

Amplinus mimus Chamberlin 1953b, p. 68. 

TYPE: Chicago Mus. Nat. Hist. 

TYPE LOCALITY: West of San Pedro Sula, Honduras. 
Pseudamplinus niteus (Chamberlin). 

Amplinus niteus Chamberlin 1922, p. 43; Hoffman 1954a, p. 51. 

TYPE: U.S. Nat. Mus., no. 846. 

TYPE LOCALITY: Basin of San Juan River, Costa Rica. 
Pseudamplinus nitidus (Brolemann). 

Platyrrhacus nitidus Brolemann 1900, p. 97. 

Pachyurus nitidus Car] 1902, p. 638. 

Amplinus nitidus Pocock 1909, p. 149. 

Orthomorpha nitidus Cook 1911c, p.627; Hoffman 1954a, p. 51. 

TY PE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Guatemala. 

OTHER LOCALITY: See under next species. 

Pseudamplinus orphnius (Chamberlin). 

Amplinus orphnius Chamberlin 1922, p. 42; Hoffman 1954a, p. 51. 

TYPE: U.S. Nat. Mus., no. 845. 

TYPE LOCALITY: La Ceiba, Honduras. 

PARATYPE LOCALITY: Livingston, Guatemala. 

OTHER LOCALITIES: Escobas, Dept. Izabal, Guatemala (Chamberlin 
1952a). Middlesex, British Honduras; Rio Santa Ana, Honduras; La Jolla 
Grande, Dept. Chimaltenango, Guatemala (Hoffman 1951). Specimens from 
last locality reidentified by Hoffman as P. nitidus. 

Pseudamplinus palicaudatus (Attems). 

Pachyurus palicaudatus Attems 1901, p. 98. 

Amplinus palicaudatus Pocock 1909, p. 150. 

Orthomorpha palicaudatus Cook 1911c, p. 627; Hoffman 1954a, p. 51. 


26 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE: Zool. Mus. Hamburg. 

TYPE AND PARATYPE LOCALITY: La Joya, Chiapas, Mexico. 

OTHER LOCALITIES: Quezaltenango; Cholhuitz, Guatemala (Pocock 
1909). Depts. Chaltenango and Santa Ana, El Salvador (Kraus 1954b). 
Pseudamplinus pococki (Cook). 

Orthomorpha pococki Cook 1911c, p. 627; Hoffman 1954a, p. 51. 

Amplinus klugi Pocock 1909, p. 152 (in part). 

TYPE: British Mus. 

TYPE LOCALITY: Jalapa, Mexico. 

PARATYPE LOCALITY: Pocock (1909) saw a specimen from Guana- 
juato at the same time he assigned the above type specimen to Amplinus 
klugi but whether or not it is the true klugi or pococki is not known. 
Pseudamplinus schmidti (Chamberlin). 

Amplinus schmidti Chamberlin 1952a, p. 576; Hoffman 1954a, p. 51. 

TYPE: Chicago Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Pseudamplinus serratus (Kraus). New combination. 

Amplinus serratus Kraus 1954b, p. 311. 

TYPE: Senckenberg Mus., Frankfurt. 


TYPE LOCALITY: Rain forest of Monte Cristo, Metapan Mts., Dept. 


Santa Ana, E1 Salvador. 
PARATYPE LOCALITIES: Same, and apex of Monte Cristo. 
Pseudamplinus tajumulco (Chamberlin). 
Amplinus tajumulco Chamberlin 1952a, p. 576; Hoffman 1954a, p. 51. 
TYPE: Chicago Mus, Nat. Hist. 
TYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Pseudamplinus tapachulae (Chamberlin). 
Amplinus tapachulae Chamberlin 1943b, p. 57. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Tapachula, Chiapas, Mexico. 
Pseudamplinus triramus (Pocock). 
Amplinus triramus Pocock 1909, p. 153. 
Orthomorpha triramus Cook 1911c, p. 627; Hoffman 1954a, p. 51. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Pseudamplinus vergelanus (Chamberlin). New combination. 
Amplinus vergelanus Chamberlin 1943b, p. 57. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: El Vergel, Chiapas, Mexico. 
Pseudamplinus xelitlus (Chamberlin). 
Amplinus xelitlus Chamberlin 1943b, p. 57; Hoffman 1954a, p. 51. 
TYPE: Collection of R. V. Chamberlin. 


ico. 


TYPE AND PARATYPE LOCALITY: Xilitla, San Luis Potost, Mex-, 








MILLIPEDS OF MEXICO AND CENTRAL AMERICA oT 


OTHER LOCALITY: Valles, San Luis Potosf, Mexico (Causey 1954). 
Pseudamplinus zuniulus (Chamberlin). 

Amplinus zuniulus Chamberlin 1952a, p. 577; Hoffman 1954a, p. 51. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Pinca (Pica?) Monte Cristo, 


VolcA4n Zunil, Guatemala. 


Family HOLISTOPHALLIDAE 


This family, erected by Silvestri when he described the single in- 
cluded genus, is being reviewed by R. L. Hoffman, who has added, in 
his treatment, the other genera which appear here. 


Genus DUOPORUS Cook 1901, p. 403. 


TYPE SPECIES: Duoporus barretti Cook, by original designation. 
Duoporus barretti Cook 1901, p. 404. 

TYPE: Probably U.S. Nat. Mus. 

TYPE AND PARATYPE LOCALITY: Cuernavaca, Morelos, Mexico. 


Genus ELCARMENIA Kraus 1954, p. 328. 


TYPE SPECIES: Elcarmenia engelhardi Kraus, by monotypy. 
Elearmenia engelhardi Kraus 1954b, p. 328. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Finca E] Carmen, San Vincente 


volcanos, Dept. San Vincente, FE] Salvador. 
Genus HOLISTOPHALLUS Silvestri 1909, p. 2. 


TYPE SPECIES: Holistophallus peregrinus Silvestri, by original des- 
ignation. 
Holistophallus peregrinus Silvestri 1909, p. 4. 

TYPE: Torino Mus. 

TYPE LOCALITY: Tabasco, Mexico. 

OTHER LOCALITIES: Quirigua; Cacao Trece Aguas, Guatemala (Cham- 
berlin 1922). Progreso, San Juan Pueblo, Honduras (Chamberlin 1922). 


Genus PAMMICROPHALLUS Pocock 1909, p. 183. 


TYPE SPECIES: Pammicrophallus ornatus Pocock, by original desig- 
nation. 
Pammicrophallus ornatus Pocock 1909, p. 184. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Pammicrophallus pasaquinae Kraus 1954b, p. 327. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: At km. 198 on highway from 
La Union after Pasaquina, Dept. La Union, El Salvador. 


273-687 O - 68 - 3 


28 U.S. NATIONAL MUSEUM BULLETIN 266 


Pammicrophallus pictus Pocock 1909, p. 185. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 


Genus SYNTHODESMUS Chamberlin 1922, p. 54. 
TYPE SPECIES: Synthodesmus simulans Chamberlin, by original des- 


ignation. 

Synthodesmus simulans Chamberlin 1922, p. 54. 
TYPE: U.S. Nat. Mus. no. 853. 
TYPE LOCALITY: San Rafael, Guatemala. 


Genus TUNODESMUS Chamberlin 1922, p. 52. 
TYPE SPECIES: Tunodesmus orthogonus Chamberlin, by original des- 


ignation. 

Tunodesmus laminiger Chamberlin 1922, p. 53. 
TYPE? U.S. Nat. Mus.,moz 852. 
TYPE LOCALITY: San Rafael, Guatemala. 

T unodesmus orthogonus Chamberlin 1922, p. 53. 
TYPE: U.S. Nat. Mus., no. 851. 
TYPE LOCALITY: San Rafael, Guatemala. 


Genus ZEUCTODESMUS Pocock 1909, p. 185. 
TYPE SPECIES: Zeuctodesmus caeruleus Pocock, by original desig- 


nation, 
Zeuctodesmus caeruleus Pocock 1909, p. 186. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Amula, Guerrero, Mexico. 
Zeuctodesmus ferrugineus Chamberlin 1943b, p. 54. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Zitacuaro, Michoacan, Mexico. 


Family ONISCODESMIDAE 
Genus BARRODESMUS Chamberlin 1940a, p. 7. 
TYPE SPECIES: Barrodesmus isolatus Chamberlin, by original des- 


ignation. 
Barrodesmus isolatus Chamberlin 1940a, p. 7. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 
Genus BONETESMUS (see bottom of page 111) 


Genus LIGIODESMUS Pocock 1909, p. 114. 


TYPE SPECIES: Ligiodesmus pusillus Pocock, by original designa- 


tion. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 29 


Ligiodesmus pusillus Pocock 1909, p. 114. 
TYPE: British Mus. 
TYPE LOCALITY: Teapa, Tabasco, Mexico. 


Genus LIGNYDESMUS Cook 1896, p. 28. 


Mesesmus Chamberlin 1941a, p. 57. 
Panamadesmus Loomis 1961, p. 89. 
Huanucodesmus Loomis 1961, p. 91. 
TYPE SPECIES: Oniscodesmus rubriceps Peters, by original desig- 
nation. 
Lignydesmus eutypus (Chamberlin). 
Oniscodesmus eutypus Chamberlin 1940a, p. 7. 
Mesesmus eutypus Chamberlin 1941a, p. 57; Loomis 1964a, p. 57. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
Lignydesmus insulanus (Chamberlin). 
M esesmus insulanus Chamberlin 1941a, p.57; Loomis 1964a, p.57. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
Lignydesmus panamanus Loomis 1961, p. 88. 
TYPE: U.S. Nat. Mus., no. 2637. 
TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 
Panama. 
Lignydesmus projectus Loomis 1964a, p. 57. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqur Prov., Pan- 
ama. 
PARATYPE LOCALITY: Same general region. 
Lignydesmus sculptilis (Loomis). 
Panamadesmus sculptilis Loomis 1961, p. 90; Loomis 1964a, p.58. 
TYPE: U.S. Nat. Mus., no. 2638. 
TYPE AND PARATYPE LOCALITY: Pifa area, Canal Zone. 


Family PERIDONTODESMIDAE 
Genus HEXODONTIA Verhoeff 1932, p. 516. 


Trachyphloeus Attems 1933, p. 264. 

Rhexiphloeus Attems 1940, p. 460; Hoffman 1950b, p. 71. 

Parapomus (subgenus of Apomus) Kraus ]1954b, p. 317. New synonymy. 

TYPE SPECIES: Hexodontia cordobana Verhoeff, by original desig- 
nation. 
Hexodontia cordobana Verhoeff 1932, p. 518. 

TYPE: Zool. Mus. Munchen (H). 

TYPE LOCALITY: Cordoba, Veracruz, Mexico. 


30 U.S. NATIONAL MUSEUM BULLETIN 266 


Hexodontia electa (Chamberlin). 

Peridontodesmus electus Chamberlin 1914, p. 188. 

Trachyphloeus electus Attems 1933, p. 265. 

Rhexiphloeus electus Attems 1940, p. 460; Hoffman 1950b, p. 71. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE AND PARATYPE LOCALITY: Laguna, Juan Vinas, Costa Rica. 

OTHER LOCALITIES: Cartago (Chamberlin 1922); Carpintera, Costa 
Rica (Attems 1933). 
Hexodontia morela (Chamberlin). 

Peridontodesmus morelus Chamberlin 1943b, p. 59; Hoffman 1950b, 

pe il. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Chapultepec, near Cuernavaca, 
Morelos, Mexico. 
Hexodontia zilchi (Kraus). 

Apomus (Parapomus) zilchi Kraus 1954b, p. 317, illustr. New combi- 

nation. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Volcan los Naranjos, Dept. Santa Ana, El Salva- 
dor. 

PARATYPE LOCALITIES: Same, and Finca el Paraiso, Santa Tecla, 
Dept. La Libertad; San Salvador, Dept. San Salvador, El Salvador. 

R. L. Hoffman first called my attention to the fact that this species 
actually was a Peridontodesmid. 


Genus KALESMUS Chamberlin 1943b, p. 60. 
TYPE SPECIES: Kalesmus phanus Chamberlin, by original designa- 


tion. 
Kalesmus eutropis Chamberlin 1943b, p. 60. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Fortin, Veracruz, Mexico. Prob- 
ably Fortin de las Flores. 
Kalesmus phanus Chamberlin 1943b, p. 60. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Parque National de Zempoala, 


Morelos, Mexico. 


Genus MADERESMUS Chamberlin 1943b, p. 61. 
TYPE SPECIES: Maderesmus tepoztlanus Chamberlin, by original des- 


ignation, 
Maderesmus tepoztlanus Chamberlin 1943b, p. 61. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Tepoztlan, Morelos, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 31 


Maderesmus hoogstraali (Chamberlin). 
Peridontodesmus hoogstraali Chamberlin 1942a, p. 58; Chamberlin 
1943b, p. 61. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Cerro Tancitaro, Michoacan, 


Mexico. 
Genus PERIDONTODESMUS Silvestri 1896b, p. 197. 
TYPE SPECIES: Polydesmus (Scytonotus) woodianus Humbert and 


Saussure, by original designation. 
Peridontodesmus flagellatus Pocock 1909, p. 136. 
TYPE: British Mus. 
TYPE LOCALITY: Cholhuitz, Guatemala. 
Peridontodesmus hirsutus Pocock 1909, p. 136. 
TYPE: British Mus. 
TYPE LOCALITY: Teapa, Tabasco, Mexico. 
Peridontodesmus medius Chamberlin 1943b, p. 59. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Ixtapan del Oro, Mexico, Mexico. 
P eridontodesmus parvus Chamberlin 1943b, p. 59. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: LaJunta, Villa Juarez, Puebla, Mexico. 
P eridontodesmus purulicus Cook 1911b, p. 470. 
T YPE: U.S. Nat. Mus., no. 809. 
TYPE LOCALITY: Purulha, Baja Verapaz, Guatemala. 
Peridontodesmus woodianus (Humbert and Saussure). 
Polydesmus (Scytonotus ) woodianus Humbert and Saussure 1869, p. 152. 
Polydesmus woodianus Saussure and Humbert 1872, p. 52; Silvestri 
1896b, p. 197. 
TYPE: Mus. Hist. Nat. Geneve (H) 
TYPE LOCALITY: Eastern Cordillera, Mexico. 


Genus PINESMUS Chamberlin 1943b, p. 62. 
TYPE SPECIES: Pinesmus setosus Chamberlin, by original designa- 


tion. 
Pinesmus monticolens Chamberlin 1952a, p. 558. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Volcd4n Tajumulco, Guatemala. 
Pinesmus setosus Chamberlin 1943b, p. 62. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Desierto de los Leones, D. F., 
Mexico. 
Pinesmus viganus Chamberlin 1952a, p. 559. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Las Vigas, Veracruz, Mexico. 


32 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus SIERRESMUS Chamberlin 1943b, p. 60. 
TYPE SPECIES: Sierresmus hidalgonus Chamberlin, by original des- 


ignation. 
Sierresmus hidalgonus Chamberlin 1943b, p. 61. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Chapulhuacan, Hidalgo, Mexico. 


Family PLATYRHACIDAE 


Genus NYSSODESMUS Cook 1896, p. 53. 


TYPE SPECIES: Nyssodesmus alboalatus Cook, by original designa- 
tion. 
Nyssodesmus alboalatus Cook 1896, p. 53. 

TYPE: Zool. Mus. Berlin. 

TYPE LOCALITY: Nicaragua. 
Nyssodesmus antius (Chamberlin). 

Platyrhacus antius Chamberlin 1914, p. 189; Chamberlin 1922, p. 40. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE AND PARATYPE LOCALITY: Juan Vifias, Reventazén Road, 
Costa Rica. 

OTHER LOCALITY: Tilaran, Costa Rica (Attems 1933). 
Nyssodesmus attemsi Hoffman 1960, p. 27. 

TYPE: U.S. Nat. Mus., no. 2468. 

TYPE LOCALITY: Puerto Armuelles, Panama. 
Nyssodesmus bivirgatus (Carl). 

? Polydesmus (Odontodesmus) python Peters 1864, p. 543. 

Platyrrhacus bivirgatus Car] 1902, p. 652; Chamberlin 1922, p. 40. 

TYPE: Mus. Hist. Nat. Geneve (H) 

TYPE LOCALITY: San José, Costa Rica. 

OTHER LOCALITIES: La Palma, and Carrillo, Costa Rica, (Cham- 
berlin 1922); Rio Jiménez, Costa Rica (Chamberlin 1933). 
Nyssodesmus concolor Loomis 1964a, p. 59. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 
Panama. 
Nyssodesmus fraternus (Carl). 

Platyrrhacus fraternus Carl 1902, p. 655; Chamberlin 1922, p. 40. 

Platyrhacus fraternus Hoffman 1953c, p. 256; Jeekel 1963a, p. 89. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: San José, Costa Rica. 

PARATYPE LOCALITY: Limon, Costa Rica. 

OTHER LOCALITIES: La Palma, and Cacht, Costa Rica (Chamberlin 
1922). 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 33 


Nyssodesmus limonensis (Attems). 
Platyrrhacus limonensis Attems 1900, p. 344; Chamberlin 1922, pe 39. 
TYPE: Zool. Mus. Hamburg (H). 
TYPE LOCALITY: Puerto Limon, Costa Rica. 
Nyssodesmus mimus Chamberlin 1922, p. 38. 
TYPE: Mus. Comp. Zool., cat. no. 5020. 
TYPE LOCALITY: Nicaragua. 
Nyssodesmus montivagus (Carl). 
? Platyrhacus mexicanus Attems 1900, p. 348. 
Platyrhacus montivagus Carl 1902, p. 662; Chamberlin 1922, p. 40. 
Nyssodesmus valerii Chamberlin 1933, p. 16; Hoffman 1953c, p. 253. 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE AND PARATYPE LOCALITY: Volcan de Turrialba, Costa Rica. 
OTHER LOCALITIES: Carrillo and La Palma (Pocock 1909); Cacht, 
Reventazén Valley (Chamberlin 1914); La Carpintera (Chamberlin 1933), 
Costa Rica. 
Nyssodesmus nicaraguanus Chamberlin 1922, p. 39. 
TYPE: Acad. Nat. Sci. Philadelphia. 
TYPE LOCALITY: Machucha, Nicaragua. 
Nyssodesmus nigricaudus Chamberlin 1922, p. 37. 
TYPE: U.S. Nat. Mus., no. 843. 
TYPE LOCALITY: Costa Rica. 
Nyssodesmus stenopterus (Brolemann). 
Platyrhacus stenopterus Brolemann 1905, p. 343; Chamberlin 1922, 
p- 40. 
TYPE: Mus. Nat. Hist. Natur., Paris (H). 
TYPE LOCALITY: Rancho Redondo, Costa Rica. 
Nyssodesmus tristani (Pocock). 
Platyrhacus tristani Pocock 1909, p. 141; Chamberlin 1922, p. 39. 
TYPE: British Mus. 
TYPE LOCALITY: La Palma, Costa Rica. 


Genus PLATYRHACUS Koch 1847, p. 58. 


Barydesmus Cook 1896, p. 53; Hoffman 1960, p. 21. 

Aymaresmus Chamberlin 194le, p. 494; Jeekel 1963a, p. 89. 

Dynesmus Chamberlin 1941e, p. 495; Jeekel 1963a, p. 89. 

TYPE SPECIES: Polydesmus scaber Perty, by monotypy. 
Platyrhacus bilineatus (Lucas). 

Polydesmus bilineatus Lucas 1840, p. 523. 

Polydesmus (Stenonia) bilineatus Saussure 1860, p. 532. 

? Platyrhacus bilineatus Attems, 1900, p. 347. 

TYPE: Not known to exist, seems not to be at Paris (H). 

TYPE LOCALITY: Mexico. 

Hoffman 1953c places this species in 
less a South American species (H). 


“‘nomen inquirendum.’” Doubt- 


34 U.S. NATIONAL MUSEUM BULLETIN 266 


P latyrhacus chapini Hoffman 1953c, p. 254. 

TYPE: U.S. Nat. Mus., no. 2079. 

TYPE LOCALITY: Los Siguas, Chiriquit Prov., Panama. 
P latyrhacus gracilipes Loomis 1964a, p. 60. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqu! Prov., Panama. 

PARATYPE LOCALITIES: Same, and Finca Lerida, near Boquete, 
Chiriqui Prov., Panama. 
Platyrhacus mexicanus (Lucas). 

Polydesmus mexicanus Lucas 1840, p. 523. 

Polydesmus (Stenonia) mexicanus Saussure 1860, p.534; Attems 1900, 
p. 348. 

TYPE: Present location unknown (H). 

TYPE LOCALITY: Mexico. 

Hoffman 1953c and Jeekel 1963a consider this species unidentifiable. 
Hoffman now thinks it may be South American. 
Platyrhacus pococki Brolemann 191 1a, p. 14. 

Nyssodesmus pococki Chamberlin 1922, p. 40; Hoffman 1953c, p. 257. 

Platyrhacus stenopterus Pocock 1909, p. 145, not Brolemann; Jeekel 

1963a, p. 89. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Rancho Redondo, Costa Rica. 

OTHER LOCALITIES: Cuesta del Tablazo Costa Rica (Chamberlin 
1922); Volcan Irazu, Costa Rica (Attems 1933). 
Platyrhacus propinquus Carl 1902, p. 665. 

Nyssodesmus propinquus Chamberlin 1922, p. 40; Hoffman 1953c, 

p. 207. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Las Delicias, Santa Clara Prov., Costa Rica. 
Platyrhacus riparius Carl 1902, p. 641. 

Nyssodesmus riparius Chamberlin 1922, p. 40; Hoffman 1953c, p. 257. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Rio General, Pacific slope, 
Costa Rica. 


Genus PSAMMODESMUS Cook 1896, p. 52. 


Ernostyx Chamberlin 1941le, p. 497; Hoffman 1953b, p. 301. 

TYPE SPECIES: Psammodesmus cos Cook, by original designation. 
Psammodesmus remotus Loomis 1964a, p. 62. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 
Psammodesmus schmitti Loomis and Hoffman. In Hoffman 1953b, p. 301. 

TYPE: U.S. Nat. Mus., no. 2016. 

TYPE LOCALITY: Puerto Obaldia, Darien Prov., Panama. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 33 


PARATYPE LOCALITY: Cana, Darien Prov., Panama. 
OTHER LOCALITY: Pequene River, just east of Canal Zone, Panama. 


Genus TIRODESMUS Cook 1896, p. 53. 
TYPE SPECIES: Polydesmus (Stenonia) fimbriatus Peters, by original 


designation. 
Tirodesmus biolleyi (Carl). 

Platyrhacus biolleyi Carl 1902, p. 658. 

Tirodesmus fimbriatus Chamberlin 1922, p. 36 (not Peters). 

Nyssodesmus albomarginis Chamberlin 1933, p. 16; Hoffman 1953c, 

p. 235; Jeekel 1963a, p. 91. 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE LOCALITY: Las Delicias, Costa Rica. 
OTHER LOCALITIES: La Palma and Cariblanco, Costa Rica (Pocock 
1909); San Juan del Norte and Machucha, Nicaragua (Chamberlin 1922), 
Tirodesmus fimbriatus (Peters). 
Polydesmus (Stenonia) fimbriatus Peters 1864, p. 543; Cook 1896, 
p. Oa. 

Platyrrhacus fimbriatus Attems 1899, p. 347; Carl 1914, p.975; Jeekel 
1963a, p. 91. 

TYPE: Location unknown. 

TYPE LOCALITY: Veragua, Panama; probably the Province of Vera- 


guas. 


Family POLYDESMIDAE 
Genus POLYDESMUS Latreille 1802, p. 44. 


TYPE SPECIES: Julus complanatus Linnaeus, by monotypy. 
Polydesmus angustus Latzel 1884, p. 262. 

Polydesmus complanatus angustus Latzel 1884, p. 262. 

Polydesmus verhoeffi angustus Attems 1940, p. 12. 

Polydesmus chapultepecus Chamberlin 1943b, p. 69. New synonymy 

by R. L. Hoffman. 

TYPE: Naturh. Mus. Wien. 

A European species reported from Mexico by Jawlowski 1930. 

OTHER LOCALITIES: Chapultepec Park, D. F., and Llano de Sal- 
azar, Morelos, Mexico (Chamberlin 1943b). 


Family RHACHODESMIDAE 
Genus ACERATOPHALLUS Car! 1902, p. 608. 


TYPE SPECIES: Aceratophallus unicolor Carl, by monotypy. 
Aceratophallus calcehtokanus Chamberlin 1938a, p. 178. 
TYPE: Collection of R. V. Chamberlin. 


36 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE LOCALITY: Xkyc Cave, Calcehtok, Yucatan, Mexico. 

PARATYPE LOCALITIES: Spukil and Sazich Caves, Calcehtok, Yu- 
catan, Mexico. 
Aceratophallus dux Chamberlin 1914, p. 193. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: Juan Vintas, Costa Rica. 

OTHER LOCALITIES: Reported from six departments of El Salvador 
(Kraus 1954b). 
Aceratophallus guatemalae Chamberlin 1947b, p. 31. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE AND PARATYPE LOCALITY: Piedras Negras, Guatemala. 
Aceratophallus hoctunanus Chamberlin 1938a, p. 176. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Hoctun Cave, Hoctun, Yucatan, 
Mexico. 
Aceratophallus lamellifer (Brolemann). 

Aceratophallus unicolor lamellifer Brolemann 1905, p. 346; Pocock 

1909, p. 183. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE AND PARATYPE LOCALITY: San José, Costa Rica. 

OTHER LOCALITIES: Chitaria, and Parismina (Chamberlin 1933); 
Tilardn (Attems 1933), Costa Rica. 
Aceratophallus maya Loomis 1964b, p. 186. 

TYPE: U.S. Nat. Mus., no. 2984. 

TYPE AND PARATYPE LOCALITY: Tikal, Dept. of Petén, Guate- 
mala. 
Aceratophallus oxkutzcabus Chamberlin 1938a, p. 176. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Gongora Cave, Oxkutzcab, Yucatan, Mexico. 

PARATYPE LOCALITIES: Same, and Puz and Xmahit Caves, Tekax, 
Yucatan, Mexico. 
Aceratophallus quadratus Loomis 1961, p. 92. 

TYPE: U.S. Nat. Mus., no. 2639. 

TYPE LOCALITY: Probably Canal Zone. 
Aceratophallus sumucus Chamberlin 1956, p. 158. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Musawas, Waspuc River, Nicaragua. 
Aceratophallus unicolor Carl 1902, p. 609. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: San José, Costa Rica. 

OTHER LOCALITIES: La Uruca (Pocock 1909); El Gallito (Chamber- 
lin 1933), Costa Rica. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA a7 


Genus ACUTANGULUS Attems 1899, p. 409. 


TYPE SPECIES: Polydesmus (Strongylosoma) coccineus Saussure and 
Humbert, by monotypy. 
Acutangulus coccineus (Humbert and Saussure). 

Polydesmus (Tropisoma) coccineus Humbert and Saussure 1869, p. 152. 

Polydesmus (Strongylosoma) coccineus Saussure and Humbert 1872, 
p- 50; Attems 1899, p. 409. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Orizaba, Mexico. 
Acutangulus neglectus Carl 1903, p. 560. 

Polydesmus (Strongylosoma) coccineus Var. Saussure and Humbert 
1672,/p. ol. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Orizaba, Mexico. 


Genus METAPHALLUS Chamberlin 1952a, p. 560. 


TYPE SPECIES: Metaphallus mimus Chamberlin, by original designa- 
tion. 
Metaphallus mimus Chamberlin 1952a, 562. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Chichivac, Guatemala. 


Genus NEOLEPTODESMUS Car! 1903, p. 557. 


Curodesmus Chamberlin 1922, p. 55; Attems 1940, p. 467. 
Euphallus Chamberlin 1952a, p.559. New synonymy by R.L. Hoffman. 
TYPE SPECIES: Polydesmus (Oxyurus) sumichrasti Humbert and 
Saussure, by subsequent designation. 
Neoleptodesmus aztecus (Saussure). 
Polydesmus (Leptodesmus) aztecus Saussure 1859, p. 324. 
Polydesmus (Oxyurus)aztecus Saussure and Humbert 1872, p. 160. 
Leptodesmus aztecus Attems 1899, p. 390; Carl 1903, p. 559. 
Euphallus dybasi Chamberlin 1952a, p. 560. New synonymy by R. L. 
Hoffman. 
TYPE: Mus. Nat. Hist. Geneve. 
TYPE LOCALITY: Volcan de Tuxtla, Mexico. 
PARATYPE LOCALITIES: Plateau of Puebla, and peak of Orizaba, 
Mexico. 
OTHER LOCALITY: El Fortin, Veracruz, Mexico. Probably Fortin de 


las Flores. 


38 U.S. NATIONAL MUSEUM BULLETIN 266 


Neoleptodesmus dispersus Chamberlin 1943b, p. 53. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Puroje Nuevo, Veracruz, Mexico. 

PARATYPE LOCALITIES: Same, and Tepoztl4n and Oaxtepec, Mo- 
relos, Mexico. 
Neoleptodesmus geniculatus Kraus 1954b, p. 325. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Rainforest of Monte Cristo, 
Metapan Mts., Dept. of Santa Ana, FE] Salvador. 
Neoleptodesmus guatemalensis ( Chamberlin). 

Curodesmus guatemalensis Chamberlin 1922, p.55; Attems 1940, p. 469. 

TYPE: U.S. Nat. Mus., no. 854. 

TYPE LOCALITY: San Rafael, Guatemala. 
N eoleptodesmus orizabae (Humbert and Saussure). 

Polydesmus(Oxyurus )orizabae Humbert and Saussure 1869, p.151. 

Leptodesmus orizabae Attems 1899, p. 389; Carl 1903, p. 557. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Orizaba, Mexico. 
N eoleptodesmus salvadorensis Kraus 1954b, p. 322. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: ‘“‘Institute,’”? San Salvador, Dept. San Salvador, 
El Salvador. 

PARATYPE LOCALITIES: Same, and Depts. La Paz, Sonsonate, La 
Libertad, San Vincente, and La Union, El Salvador. 
Neoleptodesmus sinuatus Kraus 1954b, p. 326. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Laguna de las Ninfas, Dept. Ahuachapan, F1 Sal- 
vador. 

PARATYPE LOCALITIES: Same, and Finca EF] Carmen, San Vincente 
Volcanos, Dept. San Vincente, EF] Salvador. 
Neoleptodesmus sumichrasti (Humbert and Saussure). 

Polydesmus (Oxyurus) sumichrasti Humbert and Saussure 1869, p. 151. 

Polydesmus (Oxyurus) intermedius Humbert and Saussure 1869, p.151; 

Carl 1903, p. 558. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Orizaba, Mexico. 
Neoleptodesmus vermiformis (Saussure). 

Polydesmus (Strongylosoma) vermiformis Saussure 1859, p. 326; Carl 

1903, p. 557. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 
Neoleptodesmus vulcani Kraus 1954b, p. 324. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Santa Ana Volcanos, near Buenos Aires, Dept. 
Santa Ana, El Salvador. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 39 


PARATYPE LOCALITIES: Same, and Laguna Verde and Laguna de 
las Ninfas, Dept. Ahuachapan, El Salvador. 


Genus PARARHACHISTES Pocock 1909, p. 177. 
TYPE SPECIES: Pararhachistes elevatus Pocock, by original desig- 


nation. 
Pararhachistes amblus Chamberlin 1942b, p. 13. 

TYPE: Escuela Nat. Mexico City. 

TYPE AND PARATYPE LOCALITY: Cueva de Jutxlahuaca, Colotlipa, 
Guerrero, Mexico. 
Pararhachistes elevatus Pocock 1909, p. 178. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Pararhachistes galeanae Chamberlin 1943b, p. 52. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Laguna de Labradores, Galeana, Nuevo Ledn, 
Mexico. 
Pararhachistes potosinus Chamberlin 1943b, p. 53. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tamazunchale, San Luis Potosi, 
Mexico. 
Pararhachistes vertebratus Pocock 1909, p. 178. 

TYPE: British Mus. 

TYPE LOCALITY: Amula, Guerrero, Mexico. 


Genus PHOROSITUS Chamberlin 1952a, p. 562. 
TYPE SPECIES: Phorositus granulifer Chamberlin, by original desig- 
nation. 
Phorositus granulifer Chamberlin 1952a, p. 562. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 


Genus RHACHIDOMORPHA Saussure 1860, p. 326. 


Microrhachis Car] 1903, p. 556; Pocock 1909, p. 174. 
TYPE SPECIES: Polydesmus (Rhachidomorpha) tarasca Saussure, by 
monotypy. 
Rhachidomorpha adunca (Saussure and Humbert). 
Polydesmus (Rhachidomorpha) aduncus Saussure and Humbert 1872, 
p- 158; Attems 1899, p. 410. 
Polydesmus (Rhachidomorpha) uncinatus Humbert and Saussure 1869, 
p. 152 (uncinatus preoccupied). 
Rhachis (Microrhachis) uncinatus Carl 1903, p. 556; Pocock 1909, 
p. 175. 
TYPE: Mus. Hist. Nat. Geneve (H). 


40 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE LOCALITY: Eastern Cordillera, Mexico. 

OTHER LOCALITIES: Monte Azul, Cerro de Azcamela, Sierra de Agua 
in the Eastern Cordillera, Mexico (Pocock 1909). 
Rhachidomorpha.aurantium Chamberlin 1952a, p. 564. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: El Fortin, Veracruz, Mexico. 
Probably Fortin de las Flores. 

Rhachidomorpha tarasea (Saussure). 
Polydesmus (Rhachidomorpha) tarasca Saussure 1860, p. 327; Attems 
1899, p. 410. 
TYPE: Mus. Hist. Nat. Geneve (H). 
TYPE LOCALITY: Cordoba, Mexico. 
Rhachidomorpha vicinus Chamberlin 1943b, p. 52. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Eighteen mi. west of Perote, Veracruz, Mexico. 

PARATYPE LOCALITY: Same, and 18 mi. north. 


Genus RHACHODESMUS Cook 1895, p. 4. 


Polydesmus (Rachis) Saussure 1859, p. 326 (original spelling). 

Rhachis Attems 1899, p. 326. 

Rhachis Carl 1903, p. 554, new subgenus (preoccupied). 

TYPE SPECIES: Polydesmus (Rachis) viridis Saussure, by monotypy, 
see Cook 1896, p. 6. 
Rhachodesmus viridis (Saussure). 

Polydesmus (Rachis) viridis Saussure 1859, p. 326. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Orizaba, Mexico. 


Genus SAKOPHALLUS Chamberlin 1942a, p. 59. 


TYPE SPECIES: Sakophallus simplex Chamberlin, by original desig- 
nation. 
Sakophallus simplex Chamberlin 1942a, p. 59. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Cerro Tancitaro, Michoacan Mexico. 


Genus STRONGYLODESMUS Saussure 1859, p. 327. 


Ceuthauxus Chamberlin 1942b, p. 12. New synonymy. 

Study of a topotype male of Chamberlin’s C. morelus, and of a new 
species closely related to his C. potosianus, confirm this synonymy. 

TYPE SPECIES: Strongylodesmus cyaneus Saussure, by monotypy. 
Strongylodesmus cruzanus (Chamberlin). New combination. 

Ceuthauxus cruzanus Chamberlin 1943b, p. 51. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Near Fortin, Veracruz, Mexico. Probably Fortin 


de las Flores. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 41 


Strongylodesmus cyaneus Saussure 1859, p. 327. 

Strongylodesmus viridis Peters 1864, p. 547; Saussure and Humbert 

1872, p. 55. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Orizaba, Mexico. 
Strongylodesmus geddesi Pocock 1909, p. 173. 

TYPE: British Mus. 

TYPE LOCALITY: Mexico. 
Strongylodesmus mediator (Chamberlin). New combination. 

Ceuthauxus mediator Chamberlin 1947b, p. 32. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9956. 

TYPE LOCALITY: Pablillo, Nuevo Leén, Mexico. 
Strongylodesmus morelus (Chamberlin). New combination. 

Ceuthauxus morelus Chamberlin 1943b, p. 51. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Tepoztlan, Morelos, Mexico. 
Strongylodesmus nuevus (Chamberlin). New combination. 

Pararhachistes nuevus Chamberlin 1941c, p. 63. 

Ceuthauxus nuevus Chamberlin 1943b, p. 51. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, 
Nuevo Leén, Mexico. 

OTHER LOCALITY: Cieneguillas, Nuevo Ledén, Mexico (Chamberlin 
1947b). 
Strongylodesmus palmitonus (Chamberlin). New combination. 

Ceuthauxus palmitonus Chamberlin 1942b, p. 12. 

TYPE: Escuela Nat. Mexico City. 

TYPE LOCALITY: Gruta de Palmito, Bustamente, Nuevo Leon, Mexico. 
Strongylodesmus potosianus (Chamberlin). New combination. 

Ceuthauxus potosianus Chamberlin 1947b, p. 32. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9957. 

TYPE LOCALITY: Alvarez, San Luis Potosi, Mexico. 


Genus TANCITARES Chamberlin 1942a, p. 58. 
TYPE SPECIES: Tancitares michoacanus Chamberlin, by original des- 


ignation. 
Tancitares michoacanus Chamberlin 1942a, p. 58. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Pedregal, Tancitaro, Michoacan, 


Mexico. 
Genus TEINORHACHIS Loomis 1961, p. 94. 


TYPE SPECIES: Teinorhachis tenuis Loomis, by original designation. 
Teinorhachis tenuis Loomis 1961, p. 95. 


42 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE: U.S. Nat. Mus., no. 2640. 
TYPE AND PARATYPE LOCALITY: “‘Probably Panama.’’ 


Genus TIPHALLUS Chamberlin 1952a, p. 566. 
TYPE SPECIES: Tiphallus aberrans Chamberlin, by original designa- 


tion. 
Tiphallus aberrans Chamberlin 1952a, p. 566. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Tamazunchale, San Luis Potosi, 


Mexico. 


Family SPHAERIODESMIDAE 
Genus COLOBODESMUS Brolemann 1905, p. 347. 


TYPE SPECIES: Colobodesmus biolleyi Brolemann, by monotypy. 
Colobodesmus biolleyi Brolemann 1905, p. 350. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE LOCALITY: San Jose, Costa Rica. 

PARATYPE LOCALITIES: Cacht, and Cariblanco, Costa Rica. 
Colobodesmus cobanus (Chamberlin). New combination suggested by R. L. 

Hoffman. 

Sphaeriodesmus cobanus Chamberlin 1952a, p. 554. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cobdn, Guatemala. 

PARATYPE LOCALITY: Alta Verapaz, Guatemala. 
Colobodesmus triramus Kraus 1954b, p. 320. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Finca la Yoya, km. 48 on highway behind Sonso- 
nate, Dept. Sonsonate, EF] Salvador. 

PARATYPE LOCALITIES: Volcan Izalco, foot of Cerro Verde (Mala 
Cara); Monte Cristo, Metapan Mts., Dept. Santa Ana, El Salvador. 


Genus CYCLODESMUS Humbert and Saussure 1869, p. 149. 


TYPE SPECIES: Cyclodesmus aztecus Humbert and Saussure, by mo- 
notypy. 
Cyclodesmus aztecus Humbert and Saussure 1869, p. 149. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 


Genus CYLIONUS Cook 1898, p. 462. 
TYPE SPECIES: Sphaeriodesmus gracilis Humbert and Saussure, by 


original designation. 
Cylionus constrictus Pocock 1909, p. 127. 
TYPE: British Mus. 
TYPE LOCALITY: Volcan de Agua, Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 43 


Cylionus gracilis (Humbert and Saussure). 
Sphaeriodesmus gracilis Humbert and Saussure 1869, p. 149; Cook 
1898, p. 463. 
TYPE: Mus. Hist. Nat. Geneve (H). 
TYPE LOCALITY: Moyoapan, Eastern Cordillera, Mexico. 
Cylionus kauanus Chamberlin 1938a, p. 182. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Kaua Cave, Kaua, Yucatan, Mexico. 


Genus CYPHODESMUS Peters 1864, p. 530. 


TYPE SPECIES: Oniscodesmus mexicanus Saussure, by monotypy. 
Cyphodesmus mexicanus (Saussure). 

Oniscodesmus mexicanus Saussure 1859, p. 328; Peters 1864, p. 530. 

TYPE: Supposed to be in Paris Mus., Cook 1898. 

TYPE LOCALITY: Cérdoba, Mexico. 
Cyphodesmus hidalgonus Chamberlin 1943b, p. 55. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Chapulhuacan, Hidalgo, Mexico. 


Genus EUSPHAERIODESMUS Brolemann 1916, p. 561. 
TYPE SPECIES: Sphaeriodesmus angustus Pocock, by original desig- 


nation. 
Eusphaeriodesmus angustus (Pocock). 
Sphaeriodesmus angustus Pocock 1909, p.124; Brolemann 1916, p. 561. 
TYPE: British Mus. 
TYPE LOCALITY: Senahu, Alta Verapaz, Guatemala. 
Eusphaeriodesmus prehensor (Pocock). 
Sphaeriodesmus prehensor Pocock 1909, p. 123; Brolemann 1916, p. 606. 
TYPE: British Mus. 
TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
OTHER LOCALITIES: Oaxtepec, and Tepoztlan, Morelos, Mexico 
(Chamberlin 1943b). 
Eusphaeriodesmus robustus (Pocock). 
Sphaeriodesmus robustus Pocock 1909, p. 122. 
TYPE: British Mus. 
TYPE LOCALITY: San Andreas Tuxtla, Veracruz, Mexico. 
Eusphaeriodesmus stylifer (Pocock). 
Sphaeriodesmus stylifer Pocock 1909, p. 124; Brolemann 1916, p. 606. 
TYPE: British Mus. 
TYPE LOCALITY: Irazu, Costa Rica. 


Genus HYBOCESTUS Hoffman 1959, p. 173. 
TYPE SPECIES: Hybocestus octonodus Hoffman, by original desig- 


nation. 


273-687 O- 68 - 4 


44 U.S. NATIONAL MUSEUM BULLETIN 266 


Hybocestus octonodus Hoffman 1959, p. 174. 

TYPE: U.S. Nat. Mus., no. 2594, 

TYPE AND PARATYPE LOCALITY: Coban, Baja Verapaz, Guatemala. 
Hybocestus plagiodon Hoffman 1959, p. 177. 

TYPE: U.S. Nat. Mus., no. 2593. 

TYPE AND PARATYPE LOCALITY: Trece Aguas, Guatemala. 


Genus PERIDYSODESMUS Silvestri 1910, p. 357. 
TYPE SPECIES: Peridysodesmus trifidus Silvestri, by original desig- 


nation. 
Peridysodesmus bifidus Silvestri 1910, p. 358. 
TYPE: Location unknown. 
TYPE LOCALITY: Jalapa, Veracruz, Mexico. 
Peridysodesmus trifidus Silvestri 1910, p. 358. 
TYPE: Location unknown. 
TYPE LOCALITY: Cuernavaca, Morelos, Mexico. 


Genus SPHAERIODESMUS Peters 1864, p. 529. 


Glomeridesmus Saussure 1859, p. 328. Not Gervais. 

TYPE SPECIES: Glomeridesmus mexicanus Saussure, by monotypy. 
Sphaeriodesmus bukowinus Chamberlin 1952a, p. 553. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Bukowina, British Honduras. 

PARATYPE LOCALITY: Double Falls, British Honduras. 
Sphaeriodesmus conformans Chamberlin 1925, p. 44. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

OTHER LOCALITIES: Pifia area, Canal Zone; Alahjuela and Juan 
Dias, Panama Prov., Panama (Loomis 1961). 
Sphaeriodesmus coriaceus Pocock 1909, p. 125. 

TYPE: British Mus. 

TYPE LOCALITY: San Juan, Alta Verapaz, Guatemala. 

OTHER LOCALITY: Volcan Tajumulco, Guatemala (Chamberlin 1952a). 
Sphaeriodesmus digitatus Pocock 1909, p. 124. 

TYPE: British Mus. 

TYPE LOCALITY: Volcan de Agua, Guatemala. 
Sphaeriodesmus griseus Chamberlin 1943b, p. 55. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Volcan Orizaba, Puebla, Mexico. 
Sphaeriodesmus hondurasanus Chamberlin 1922, p. 57. 

TYPE: U.S. Nat. Mus. no. 855. 

TYPE LOCALITY: San Juan Pueblo, Honduras. 

PARATYPE LOCALITIES: Same and Lombardia, Honduras. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 45 


Sphaeriodesmus isolatus Chamberlin 1940a, p. 13. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal Zone. 


Sphaeriodesmus longiramus Kraus 1954b, p. 319. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Finca San Jorge, near Santa Ana, Dept. Santa Ana, 
El Salvador. 


PARATYPE LOCALITIES: Same and Laguna de las Ninfas, Dept. 
Ahuachapan; Finca El] Carmen, Volcdns San Vincente, Dept. San Vin- 
cente, El Salvador. 

Sphaeriodesmus longitubus Loomis 1963, p. 119. 
TYPE: U.S. Nat. Mus. no. 2963. 
TYPE AND PARATYPE LOCALITY: Five mi. southwest of Monterrey, 


on road to Chipinque Mesa, Nuevo Leon, Mexico. 


Sphaeriodesmus medius Carl 1902, p. 675. 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE AND PARATYPE LOCALITY: Guatemala. 


Sphaeriodesmus mexicanus (Saussure). 
Glomeridesmus mexicanus Saussure 1859, p. 328; Peters 1864, p. 529. 
TYPE: Mus. Hist. Nat. Geneve (H). 
TYPE LOCALITY: Cordoba, Mexico. 
OTHER LOCALITY: Veracruz, Mexico (Cook 1898). 


Sphaeriodesmus michoacanus Chamberlin 1942a, p. 57. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Cerro San Miguel, Tancitaro, Michoacan, Mexico. 
Sphaeriodesmus neglectus Carl 1902, p. 676. 

Glomeridesmus mexicanus Saussure 1860, p. 276 (in part). 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Cérdoba, Mexico. Carl gives only 
““Mexico”’ in his description. 
Sphaeriodesmus nodulosus Kraus 1954b, p. 318. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Hacienda Los Planes, Metapan Mts., Dept. Santa 
Ana, El Salvador. 


Sphaeriodesmus oniscus Pocock 1909, p. 122. 
TYPE: British Mus. 
TYPE LOCALITY: San Andreas Tuxtla, Veracruz, Mexico. 
Sphaeriodesmus saussurei Attems 1900, p. 391. 
Sphaeriodesmus mexicanus Saussure and Humbert 1872, p. 21 (in part). 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE LOCALITY: Cerro de Escamela, Eastern Cordillera, Mexico. 


46 U.S. NATIONAL MUSEUM BULLETIN 266 
Genus TAPHRODESMUS Silvestri 1910, p. 357. 
TYPE SPECIES: Taphrodesmus vestitus Silvestri, by original desig- 


nation. 

Taphrodesmus vestitus Silvestri 1910, p. 357. 
TYPE: Lab. Zool., Portici? 
TYPE LOCALITY: Jalapa, Mexico. 


Family STYLODESMIDAE 
Genus AIROCAULON Loomis 1964a, p. 66. 
TYPE SPECIES: Airocaulon humerosum Loomis, by original designa- 


tion. 
Airocaulon humerosum Loomis 1964a, p. 66. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 


Genus APSYMA Chamberlin 1943b, p. 65. 


TYPE SPECIES: Apsyma atopa Chamberlin, by original designation. 
Apsyma atopa Chamberlin 1943b, p. 65. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: San Rafael, Mexico. 


Genus AUGESMUS Chamberlin 1951, p. 77. 
TYPE SPECIES: Cynedesmus varilobatus Loomis, by original desig- 


nation. 
Augesmus trinus (Loomis). 
Cynedesmus trinus Loomis 1959a, p. 5; 1964a, p. 67. 
TYPE: U.S. Nat. Mus., no. 2936. 
TYPE AND PARATYPE LOCALITY: Monte Lirio, Canal Zone. 
OTHER LOCALITIES: Cerro Campana, Panama Prov., Panama (Loomis 
1961); Barro Colorado Island, Canal Zone (Loomis 1964b). 


Genus BOLIVARESMUS Chamberlin 1942b, p. 10. 
TYPE SPECIES: Bolivaresmus sabinus Chamberlin, by original desig- 


nation. 
Bolivaresmus sabinus Chamberlin 1942b, p. 10. 

TYPE: Escuela Nat. Mexico City. 

TYPE LOCALITY: Cueva de los Sabinos, Valles, San Luis Potosi, 
Mexico. 

PARATYPE LOCALITIES: Same and Cueva Chica (probably same gen- 


eral area, Loomis). 
Genus BOTRYDESMUS Loomis 1934, p. 62. 


Prionodesmus Loomis 1959a, p. 7; 1964a, p. 67. 
TYPE SPECIES: Botrydesmus lutosus Loomis, by original designa- 


tion. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 47 


Botrydesmus biramosus Loomis 1964a, p. 68. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 
Botrydesmus conifer Loomis 1964a, p. 68. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: West of Finca Palo Santo, near 
Nueva California, Chiriqui Prov., Panama. 
Botrydesmus cryptus Chamberlin 1940a, p. 6. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
Botrydesmus fulgens (Loomis). 

Prionodesmus fulgens Loomis 1959a, p. 7. 

Botrydesmus coronatus Loomis 1961, p. 96; 1964a, p. 69. 

TYPE: U.S. Nat. Mus., no. 2937. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

PARATYPE LOCALITIES: Same and Cerro Campana, Panama Prov., 
Panama. 

Genus CALYMMODESMUS Car! 1914, p. 959. 


T YPE SPECIES: Calymmodesmus andinus Carl, by monotypy. 
Calymmodesmus biensifer Loomis 1959a, p. 4. 

TYPE: U.S. Nat. Mus., no. 2558. 

TYPE AND PARATYPE LOCALITY: Loma Bonita, 20 km. south of 
Tuxtepic, Oaxaca, Mexico. 
Calymmodesmus falcatus Loomis 1959a, p. 2. 

TYPE: U.S. Nat. Mus., no. 2552. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 
Calymmodesmus formicatus Loomis 1959a, p. 2. 

TYPE: U.S. Nat. Mus., no. 2555. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

PARATYPE LOCALITIES: Same and Ancon, Canal Zone. 
Calymmodesmus gracilis Loomis 1959a, p. 4. 

TYPE: U.S. Nat. Mus., no. 2553. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

PARATYPE LOCALITIES: Same and Madden Dam, Panama Prov., 
Panama. 
Calymmodesmus inquinatus Causey 1960, p. 276. 

TYPE: American Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: Gruta el Salvino, near Cayuga 
and Puerto Barrios, Guatemala. 
Calymmodesmus mexicanus Loomis 1959a, p. 5. 

TYPE: U.S. Nat. Mus., no. 2554. 

TYPE AND PARATYPE LOCALITY: Loma Bonita, 20 km. south of 


Tuxtepic, Oaxaca, Mexico. 


48 U.S. NATIONAL MUSEUM BULLETIN 266 


Calymmodesmus montanus Loomis 1964b, p. 187. 

TYPE: U.S. Nat. Mus., no. 2985. 

TYPE AND PARATYPE LOCALITY: Monteverde, Puntarenas Prov., 
Costa Rica. 
Calymmodesmus rettenmeyeri Loomis 1959a, p. 4. 

TYPE: U.S. Nat. Mus., no. 2556. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

PARATYPE LOCALITIES: Same and Madden Dam, Panama Prov., 
Panama. 
Calymmodesmus sodalis Schubart 1952, p. 438. 

TYPE: Dept. Zool., Sao Paulo (H). 

TYPE AND PARATYPE LOCALITY: Costa Rica. 


Genus CATAPLEURADESMUS Loomis 1964a, p. 71. 
TYPE SPECIES: Catapleuradesmus diadematus Loomis, by original 


designation. 
Catapleuradesmus diadematus Loomis 1964a, p. 72. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 


Panama. 


Genus CERATESMUS Chamberlin 1942b, p. 10. 
TYPE SPECIES: Ceratesmus clarus Chamberlin, by original designa- 


tion. 
Ceratesmus clarus Chamberlin 1942b, p. 10. 
TYPE: Escuela Nat. Mexico City. 
TYPE LOCALITY: Gruta de Atoyac, Veracruz, Mexico. 


Genus CRYPTYMA Chamberlin 1943b, p. 66. 


TYPE SPECIES: Cryptyma lobata Chamberlin, by original designation. 
Cryptyma lobata Chamberlin 1943b, p. 66. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Fortin, Veracruz, Mexico. Probably Fortin de las 


Flores. 


Genus DECAPORODESMUS Kenyon 1899, p. 299, 
Oligodesmus Gill 1899, p. 300; Pocock 1909, p. 133. 
TYPE SPECIES: Decaporodesmus motzoronginis Kenyon, by original 


designation. 

Decaporodesmus motzoronginis Kenyon 1899, p. 299. 
TYPE: Location unknown. Not in U.S. Nat. Mus. (H) 
TYPE LOCALITY: Motzorongo, Veracruz, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 49 


Genus EIRENYMA Chamberlin 1943b, p. 67. 


TYPE SPECIES: Eirenyma munda Chamberlin, by original designation. 
Eirenyma munda Chamberlin 1943b, p. 67. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Atoyac, Guerrero, Mexico. 


Genus GASATOMUS Chamberlin 1923b, p. 417. 
TYPE SPECIES: Gasatomus emersoni Chamberlin, by original desig- 


nation. 
Gasatomus dubius Chamberlin 1940a, p. 5. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 


Genus IDIURODESMUS Silvestri 1924, p. 172. 


TYPE SPECIES: /diurodesmus tristani Silvestri, by monotypy. 
Idiurodesmus tristani Silvestri 1924, p. 175. 

TYPE: Lab. Zool., Portici ? (H). 

TYPE LOCALITY: Turrucares, Costa Rica. 


Genus ILYMA Chamberlin 1941b, p. 24. 


TYPE SPECIES: /lyma orizaba Chamberlin, by original designation. 
Ilyma colotlipa Chamberlin 1942b, p. 9. 

TYPE: Escuela Nat. Mexico City. [Probably still at Univ. Utah (H). ] 

TYPE AND PARATYPE LOCALITY: Cueva de Jutxlahuaca, Colotlipa, 
Guerrero, Mexico. 
Ilyma morela Chamberlin 1943b, p. 63. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Parque National de Zempoala, 
Morelos, Mexico. 
Ilyma orizaba Chamberlin 1941b, p. 24. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Collected at Laredo, Texas, on orchids from Ori- 
zaba, Veracruz, Mexico. 
Ilyma potosina Chamberlin 1943b, p. 64. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: El Plutanito, San Luis Potosi, Mexico. 


Genus KAPYRODESMUS Attems 1940, p. 322. 


Xerodesmus Chamberlin 1923a, p. 403; preoccupied by Cook 1896. 
TYPE SPECIES: Xerodesmus mulegensis Chamberlin, by original des- 


ignation. 


50 U.S. NATIONAL MUSEUM BULLETIN 266 


Kapyrodesmus mulegensis (Chamberlin). 
Xerodesmus mulegensis Chamberlin 1923a, p. 403; Attems 1940, p. 322. 
TYPE: Mus. California Acad. Sci., no. 1260. 
TYPE AND PARATYPE LOCALITY: Mulegé, Baja California, Mexico. 


Genus LOPHODESMUS Pocock 1894, p. 372. 
TYPE SPECIES: Lophodesmus pusillus Pocock, by original designa- 


tion. 
Lophodesmus celatus Pocock 1909, p. 133. 
Cynedesmus celatus Chamberlin 1922, p. 59; 1951, p. 77. 
TYPE: British Mus. 
TYPE LOCALITY: Volcan de Agua, Guatemala. 
OTHER LOCALITY: Joyabaj, Guatemala (Chamberlin 1922). 
Lophodesmus laminatus Pocock 1909, p. 132. 
TYPE: British Mus. 
TYPE LOCALITY: Teapa, Tabasco, Mexico. 
Lophodesmus perparvus Pocock 1909, p. 133. 
Cynedesmus perparvus Chamberlin 1922, p. 60; 1951, p. 77. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Volcan de Agua, Guatemala. 


Genus MYRMECODESMUS Silvestri 1910, p. 359. 
TYPE SPECIES: Myrmecodesmus formicarius Silvestri, by original des- 


ignation. 
Myrmecodesmus formicarius Silvestri 1910, p. 359. 
TYPE: Location unknown (H). 
TYPE LOCALITY: Jalapa, Mexico. 
Myrmecodesmus modestus Silvestri 1911, p. 194. 
TYPE: Location unknown (H). 
TYPE LOCALITY: Jalapa, Mexico. 


Genus NABOTOMUS Chamberlin 1940a, p. 5. 


TYPE SPECIES: Nabotomus delus Chamberlin, by original designation. 
Nabotomus delus Chamberlin 1940a, p. 5. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 


Genus ORTHYMA Chamberlin 1943b, p. 66. 


TYPE SPECIES: Orthyma clara Chamberlin, by original designation. 
Orthyma clara Chamberlin 1943b, p. 66. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Fortin, Veracruz, Mexico. Probably Fortin de las 


Flores. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA SI 


Genus PORATIA Cook and Cook 1894, p. 238. 


Tidopterus Chamberlin 1923b, p. 420; Loomis 1961, p. 97. 

Dominicodesmus Chamberlin 1923c, p. 189; Loomis 1961, p. 97. 

TYPE SPECIES: Scytonotus digitatus Porat, by original designation. 
Poratia digitata (Porat). 

Scytonotus digitatus Porat 1889b, p. 24; Cook and Cook 1894, p. 238. 

Poratia heterotuberculata Carl 1902, p. 667; Silvestri 1923, p. 375. 

Dominicodesmus panamicus Chamberlin 1940a, p. 6; Loomis 1961, p. 98. 

TYPE: Naturh. Mus., Stockholm ? (H) 

TYPE LOCALITY: Sweden, probably introduced into greenhouse. 

OTHER LOCALITIES: Frijoles, Gamboa, and Monte Lirio, Canal Zone 
(Loomis 1961); Barro Colorado Island, Canal Zone (Loomis 1964a); Costa 
Rica (Silvestri 1923). 
Poratia fossata Loomis 1964a, p. 74. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 
Panama. 
Poratia granulofrons (Chamberlin). 

Treseolobus granulofrons Chamberlin 1918a, p. 221. 

Dominicodesmus geophilus Chamberlin 1923c, p. 189; Loomis 1934, 
p- 90. 

Psochodesmus granulofrons Loomis 1934, p. 55; 1961, p. 98. 

TYPE: Mus. Comp. Zool. 

TYPE AND PARATYPE LOCALITY: Jacmel, Haiti. 

OTHER LOCALITIES: Barro Colorado Island, and Piffia area, Canal 
Zone (Loomis 1961); Taboga Island (Loomis 1961), and Chilibrillo Cave, 
near Chilibre (Loomis 1964a), Panama Prov., Panama. 


Genus PROSOPODESMUS Silvestri 1910, p. 360. 


Homodesmus Chamberlin 1918a, p. 222; Loomis 1950, p. 166. 

TYPE SPECIES: Prosopodesmus jacobsoni Silvestri, by original des- 
ignation. 
Prosopodesmus jacobsoni Silvestri 1910, p. 362. 

Homodesmus parvus Chamberlin 1918a, p. 223; Loomis 1950, p. 166. 

TYPE: Lab. Zool., Portici ? (H) 

TYPE LOCALITY: Batavia, Java. 

OTHER CHECKLIST LOCALITIES: Barro Colorado Island, and Mad- 
den Forest Preserve, Canal Zone (Loomis 1964a); Chilibrillo Cave, near 
Chilibre, Panama Prov., Panama (Loomis 1964a). 


Genus RETTENMEYERIA Loomis 1959a, p. 6. 


TYPE SPECIES: Rettenmeyeria parvipes Loomis, by original desig- 


nation. 


a2 U.S. NATIONAL MUSEUM BULLETIN 266 


Rettenmeyeria parvipes Loomis 1959a, p. 6. 
TYPE: U.S. Nat. Mus., no. 2938. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 
Genus SAROLEPIS Loomis 1964a, p. 75. 


TYPE SPECIES: Sarolepis spiculatus Loomis, by original designation. 
Sarolepis spiculatus Loomis 1964a, p. 76. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro Prov., 


Panama. 
Genus STYRAXODESMUS Chamberlin 1918, p. 222. 
TYPE SPECIES: Styraxodesmus furcatus Chamberlin, by original des- 


ignation. 
Styraxodesmus chipinqueus Chamberlin 1943b, p. 68. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Chipinque, Nuevo Leén, Mexico. 
Undoubtedly Chipinque Mesa, near Monterrey. 


Genus TELAUXUS Chamberlin 1943b, p. 68. 
TYPE SPECIES: Telauxus fractus Chamberlin, by original designa- 


tion. 
Telauxus fractus Chamberlin 1943b, p. 68. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Fortin, Veracruz, Mexico. Probably Fortin de las 


Flores. 
Genus TRACHELOASPIS Loomis 1961, p. 98. 
TYPE SPECIES: Tracheloaspis tumida Loomis, by original designa- 


tion. 
Tracheloaspis tumida Loomis 1961, p. 99. 
TYPE: U.S. Nat. Mus., no. 2642. 
TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
PARATYPE LOCALITIES: Same, and Monte Lirio, Canal Zone; Alah- 


juela and Juan Dids, Panama Prov., Panama. 


Genus XENOPORUS Loomis 1961, p. 100. 


TYPE SPECIES: Xenoporus carinaceps Loomis, by original desig- 
nation. 
Xenoporus carinaceps Loomis 1961, p. 101. 

TYPE: U.S. Nat. Mus., no. 2643. 

TYPE AND PARATYPE LOCALITY: El Valle, Coclé Prov., Panama. 

OTHER LOCALITY: Almirante, Bocas del Toro Prov., Panama 
(Loomis 1964a). 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA a3 


Genus YUCODESMUS Chamberlin 1938a, p. 178. 
TYPE SPECIES: Yucodesmus viabilis Chamberlin, by original desig- 


nation. 
Yucodesmus alienus Chamberlin 1938a, p. 180. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Probably Hoctun Cave, Hoctun, 
Yucatan, Mexico. 
Yucodesmus dampfi Chamberlin 1947d, p. 101. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Hacienda Sayula, west of 
Veracruz, Veracruz, Mexico. 
Yucodesmus isidricus Chamberlin 1938a, p. 179. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: San Isidro Cave, Mérida, 
Yucatan, Mexico. 
Yucodesmus mumuztunicus Chamberlin 1938a, p. 182. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Muruztun Cave, Tizamin, 
Yucatan, Mexico. 
Yucodesmus viabilis Chamberlin 1938a, p. 179. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Luchil Cave, Tixcacal, Yucatan, Mexico. 

PARATYPE LOCALITIES: Same, and Spukil and Sazich Caves, 
Calcehtok; Gongora Cave, Oxkutzcab; San Isidro Cave, Mérida, Yucatan, 
Mexico. 


Family TRICHOPOLYDESMIDAE 
Genus CYCLOPSODESMUS Loomis 1964a, p. 80. 
TYPE SPECIES: Cyclopsodesmus scaurus Loomis, by original desig- 


nation. 
Cyclopsodesmus geniculatus Loomis 1964a, p. 82. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Cerro Punta, Chiriqui Prov., 
Panama. 
Cyclopsodesmus octosetosus Loomis 1964a, p. 83 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqui Prov., Panama. 
Cyclopsodesmus scaurus Loomis 1964a, p. 81. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Barca area, Finca Lerida, near Boquete, Chiriqui 
Prov., Panama. 

PARATYPE LOCALITIES: Same, and Cerro Punta, Chiriqui Prov., 


Panama. 


54 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus CYLINDROGONUS Loomis 1964a, p. 83. 
TYPE SPECIES: Cylindrogonus copiosus Loomis, by original desig- 


nation. 
Cylindrogonus copiosus Loomis 1964a, p. 84. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Casita Alta, Finca Lerida, near Boquete, Chiriqui 
Prov., Panama. 


PARATYPE LOCALITIES: Same and general region. 


Genus ENANTIOGONUS Loomis 1961, p. 103. 


TYPE SPECIES: Enantiogonus fragilis Loomis, by original desig- 
nation. 
Enantiogonus fragilis Loomis 1961, p. 104. 

TYPE: U.S. Nat. Mus., no. 2644. 

TYPE LOCALITY: Pina area, Canal Zone. 


Genus EUTYNELLUS Chamberlin 1940a, p. 9. 
TYPE SPECIES: Eutynellus flavior Chamberlin, by original desig- 


nation. 
Eutynellus flavior Chamberlin, 1940a, p. 10. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 


Genus HARPOGONOPUS Loomis 1960, p. 58. 


TYPE SPECIES: Harpogonopus confluentus Loomis, by original 
designation. 
Harpogonopus confluentus Loomis 1960, p. 59. 

TYPE: U.S. Nat. Mus., no. 2802. 

TYPE LOCALITY: Cottonwood Creek, 45 mi. east of San Diego, 
California. 

PARATYPE LOCALITIES: Same, and 14-20 mi. north of Ensenada, 


Baja California, Mexico. 


Genus IRAZUNUS Attems 1933, p. 260. 


TYPE SPECIES: J/razunus reimoseri Attems, by original designation. 
Irazunus chiriquensis Loomis 1964a, p. 87. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: West of Finca Palo Santo, near Nueva California, 
Chiriqui Prov., Panama. 


PARATYPE LOCALITIES: Same and general region. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 55 


Irazunus minusculus Attems 1933, p. 262. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE LOCALITY: Hamburg Farm, north of Puerto Limén, Costa Rica. 
Irazunus ovatus Loomis 1964a, p. 88. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Finca Lerida, near Boquete 
Chiriqui Prov., Panama. 
Irazunus penicillatus Loomis 1964a, p. 89. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Barca area, Finca Lerida, near Boquete, Chiriqui 
Prov., Panama. 

PARATYPE LOCALITIES: Same and general region. 
Irazunus reimoseri Attems 1933, p. 260. 

TYPE: Naturh. Mus. Wien. 

TYPE AND PARATYPE LOCALITY: Volcan Irazu, Costa Rica. 
Irazunus uncus Loomis 1964a, p. 90. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 
Prov., Panama. 
Irazunus velaripes Loomis 1964a, p. 91. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 

PARATYPE LOCALITIES: Same and general region. 


Genus IROGONUS Loomis 1964a, p. 92. 


TYPE SPECIES: J/rogonus reniformis Loomis, by original designation. 
Irogonus reniformis Loomis 1964a, p. 92. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqui Prov., Panama. 


Genus LEIOGONOPUS Loomis 1964a, p. 93. 
TYPE SPECIES: Leiogonopus echinus Loomis, by original desig- 


nation. 
Leiogonopus echinus Loomis 1964a, p. 94. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 
Prov., Panama. 
Leiogonopus bidentatus Loomis 1964a, p. 96. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: F1 Valle, Coclé Prov., Panama. 


56 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus PACHYGONOPUS Loomis 1964a, p. 96. 


TYPE SPECIES: Pachygonopus apiculatus Loomis, by original desig- 
nation. 
Pachygonopus apiculatus Loomis 1964a, p. 97. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 


Genus PHREATODESMUS Loomis 1960, p. 62. 
TYPE SPECIES: Phreatodesmus torreyanus Loomis, by original 


designation. 

Phreatodesmus torreyanus Loomis 1960, p. 63. 
TYPE: U.S. Nat. Mus., no. 2805. 
TYPE LOCALITY: Torrey Pines, above La Jolla, California. 
PARATYPE LOCALITIES: Same, and Temescal Canyon Road, near 


Corona, California; North of Ensenada, Baja California, Mexico. 


Genus SALVADORIA Kraus 1954b, p. 314. 


TYPE SPECIES: Salvadoria alata Kraus, by original designation. 
Salvadoria alata Kraus 1954b, p. 315. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Laguna Verde, Dept. Ahuachapan, El Salvador. 
Salvadoria alata propinqua Kraus 1954b, p. 315. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Laguna de las Ranas, Dept. 
Sonsonate, EF] Salvador. 
Salvadoria argentea Kraus 1954b, p. 316. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Hacienda les Planes, Metapan Mts., Dept. Santa 
Ana, El Salvador. 
Salvadoria fureata Kraus 1954b, p. 316. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Rainforest of Monte Cristo, Metapan Mts., Dept. 
Santa Ana, EF] Salvador. 
Salvadoria sagittalis Kraus 1954b, p. 316. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Laguna Verde, Dept. Ahuacha- 
pan, El Salvador. 


Genus TICHODESMUS Chamberlin 1940a, p. 13. 
TYPE SPECIES: Tichodesmus micrus Chamberlin, by original desig- 


nation. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 57 


Tichodesmus micrus Chamberlin 1940a, p. 13. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 


Family TRIDONTOMIDAE 
Genus AENIGMOPUS Loomis and Hoffman 1962, p. 151. 


TYPE SPECIES: Aenigmopus alatus Loomis and Hoffman, by original 
designation. 
Aenigmopus alatus Loomis and Hoffman 1962, p. 152. 

TYPE: U.S: Nat. Mus., no. 2824. 

TYPE AND PARATYPE LOCALITY: Scamay, about 3 mi. south of 


Senahu, Guatemala. 


Genus TRIDONTOMUS Loomis and Hoffman 1962, p. 146. 
TYPE SPECIES: Tridontomus procerus Loomis and Hoffman, by 


original designation. 
Tridontomus procerus Loomis and Hoffman 1962, p. 148. 
TYPE: U.S. Nat. Mus., no. 2823. 
TYPE AND PARATYPE LOCALITY: Sepacuite, Alta Verapaz, 


Guatemala. 


Family XYSTODESMIDAE 
Genus CRUZODESMUS Chamberlin 1943b, p. 48. 
TYPE SPECIES: Cruzodesmus ergus Chamberlin, by original desig- 


nation. 
Cruzodesmus browni Chamberlin 1943b, p. 49. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Near Fortin, Veracruz, Mexico; probably Fortin 
de las Flores. 
Cruzodesmus ergus Chamberlin 1943b, p. 48. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Near Fortin, Veracruz, Mexico; probably Fortin 
de las Flores. 
Cruzodesmus purojenus Chamberlin 1943b, p. 49. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Puroje Nuevo, Veracruz, Mexico. 


Genus RHYSODESMUS Cook 1895, p. 5. 


Damp faria Verhoeff 1932, p. 519; Attems 1938, p. 138. 
Aporiaria Chamberlin 1938b, p. 207; Chamberlin 1943c, p. 143 


58 U.S. NATIONAL MUSEUM BULLETIN 266 


Acentronus Chamberlin 1943b, p. 50. New synonymy, suggested by 
R. L. Hoffman. 

TYPE SPECIES: Polydesmus limax Saussure, by original desig- 
nation. 
Rhysodesmus acolhuus (Humbert and Saussure). 

Polydesmus acolhuus Humbert and Saussure 1869, p. 150. 

Polydesmus (Fontaria) acolhuus Saussure and Humbert 1872, p. 33; 
Pocock 1910, p. 203. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Valley of Moyapan and the 
Sierra de Agua, near Orizaba, Eastern Cordillera, Mexico. 
Rhysodesmus acuarius Attems 1931, p. 63. 

TYPE: Natur. Mus. Wien. (H) 

TYPE LOCALITY: Orizaba, Veracruz, Mexico. 
Rhysodesmus alpuyecus Chamberlin 1943b, p. 37. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Alpuyeca, Morelos, Mexico. 
Rhysodesmus amplinus Chamberlin 1952a, p. 571. 

TYPE: Chicago Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: El Fortin, Veracruz, Mexico. 
Probably Fortin de las Flores. 
Rhysodesmus angelus (Karsch). 

Polydesmus (Fontaria) angelus Karsch 1881, p. 39. 

Fontaria tepaneca Attems 1899, p. 259; Pocock 1910, p. 203. 

TYPE: Zool. Mus. Berlin (H). 

TYPE AND PARATYPE LOCALITY: Puebla, Mexico. 
Rhysodesmus arcuatus Pocock 1910, p. 201. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus attemsi Pocock 1909, p. 200. 

TYPE: British Mus. 

TYPE LOCALITY: Amula, Guerrero, Mexico. 

PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus bolivari Chamberlin 1943b, p. 38. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Garcia, Nuevo Leén, Mexico. 
Rhysodesmus bonus Chamberlin 1943b, p. 38. 

TYPE Collection of R. V. Chamberlin. 

TYPE LOCALITY: Fortin, Veracruz, Mexico. Probably Fortin de 
las Flores. 

PARATYPE LOCALITY: Orizaba, Mexico. 
Rhysodesmus brachus Chamberlin 1941c, p. 63. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, 


Nuevo Ledén, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 59 


Rhysodesmus championi Pocock 1909, p. 198. 

TYPE: British Mus. 

TYPE LOCALITY: Zapote, Pacific slope, Guatemala. 
Rhysodesmus consobrinus (Saussure). 

Polydesmus consobrinus Saussure 1859, p. 322. 

Polydesmus (Fontaria) consobrinus Saussure 1860, p. 317; Pocock 
1910, p. 204. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Anahuac, Mexico. 

PARATYPE LOCALITY: Volcdn Orizaba, Mexico. 
Rhysodesmus cuernavacae Chamberlin 1942c, p. 92. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Cuernavaca, Morelos, Mexico. 
Rhysodesmus cumbres Chamberlin 1943b, p. 39. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Cumbres, Veracruz, Mexico. 
Rhysodesmus dampfi (Verhoeff). New combination, by R. L. Hoffman. 

Damp faria damp fi Verhoeff 1932, p. 521. Hoffman states this species 
was mistakenly considered a synonym of R. attemsi Pocock, by Attems 
1938, p. 148. 

TYPE: Zool. Mus. Munchen (H). 

TYPE LOCALITY: Desierto de los Leones, Mexico. 
Rhysodesmus elestribus Chamberlin 1943b, p. 39. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Elestribo, Michoacan, Mexico. 

PARATYPE LOCALITY: Same and Tepoztlan, Morelos, Mexico. 
Rhysodesmus esperanzae Chamberlin 1943b, p. 40. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Esperanza, Guanajuato, Mexico 
Rhysodesmus eunis Chamberlin 1943b, p. 41. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Volcan Orizaba, Puebla, Mexico. 


Rhysodesmus eusculptus Chamberlin 194Ic, p. 64. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Tancitaro, Michoacan, Mexico. 


Rhysodesmus eutypus Chamberlin 1943b, p. 41. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Los Berros, Mexico. 

PARATYPE LOCALITY: Same and Desierto de los Leones, D. F., 
Mexico. 
Rhysodesmus flavocinctus Pocock 1909, p. 199. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Amula, Guerrero, Mexico. 


273-687 O- 68-5 


60 U.S. NATIONAL MUSEUM BULLETIN 266 


Rhysodesmus fraternus (Saussure). 

Polydesmus fraternus Saussure 1859, p. 323. 

Polydesmus (Fontaria) fratermus Saussure 1860, p. 310. 

? Fontaria fraterna Attems 1900, p. 260; Pocock 1910, p. 205. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Villages, eastern slope of 
Cordillera, Mexico. 
Rhysodesmus frionus Chamberlin 1943b, p. 42. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Rio Frio, D. F., Mexico. 

PARATYPE LOCALITIES: Same and San Rafael, D. F.; Rio Frio, 
Puebla, Mexico. 
Rhysodesmus garcianus Chamberlin 1943b, p. 42. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Garcia, Nuevo Leén, Mexico. 
Rhysodesmus godmani Pocock 1909, p. 199. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 

PARATYPE LOCALITY: Amula, Guerrero, Mexico. 
Rhysodesmus guardanus Chamberlin 1943b, p. 43. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: EI Guarda, D. F., Mexico 
Rhysodesmus intermedius Chamberlin 1943b, p. 43. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chilpancingo, Guerrero, Mexico. 
Rhysodesmus inustus Pocock 1910, p. 202. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus knighti Chamberlin 1941c, p. 65. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Horsetail Falls, Hacienda Vista Hermosa, Villa 
Santiago, Nuevo Leén, Mexico. 
Rhysodesmus leonensis Chamberlin 1941c, p. 65. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, Nuevo Leon, 
Mexico. 
Rhysodesmus limax (Saussure). 

Polydesmus limax Saussure 1859, p. 312. 

Polydesmus (Fontaria) limax Saussure 1860, p. 312; Cook 1895, p. 5. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Cérdoba, Veracruz, Mexico. 

PARATYPE LOCALITY: San Andréas Tuxtla, Veracruz, Mexico. 
Rhysodesmus malinche Chamberlin 1943b, p. 44. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: La Caffada, Malinche, Tlaxcala, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 61 


Rhysodesmus marcosus Chamberlin 1952a, p. 572. 

TYPE: Chicago Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: San Marcos, Guatemala. 
Rhysodesmus mayanus Chamberlin 1925, p. 44. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Yucatan, Mexico. 
Rhysodesmus minor (Chamberlin). New combination. 

Acentronus minor Chamberlin 1943b, p. 50; transfer suggested by 
R. L. Hoffman, after having examined the holotype. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chilpancingo, Guerrero, Mexico. 
Rhysodesmus morelus Chamberlin 1943b, p. 44. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tepoztlan, Morelos, Mexico. 
Rhysodesmus montezumae (Saussure). 

Polydesmus (Fontaria) montezumae Saussure 1859, p. 321; Pocock 
1909, p. 195. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Mirador, Cérdoba, and the 
plateau of Anahuac round Puebla and Chalchicomula, Mexico. 

OTHER LOCALITY: Peak of Orizaba, Mexico (Pocock 1909). 
Rhysodesmus mystecus (Humbert and Saussure). 

Polydesmus my stecus Humbert and Saussure 1869, p. 150. 

Polydesmus (Fontaria) mystecus Saussure and Humbert 1872, p. 32; 
Pocock 1910, p. 206. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Mid-region of Eastern Cordillera, 
Mexico. 
Rhysodesmus nahuus (Humbert and Saussure). 

Polydesmus nahuus Humbert and Saussure 1869, p. 150. 

Polydesmus (Fontaria) nahuus Saussure and Humbert 1872, p. 36; 
Pocock 1910, p. 206. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 
Rhysodesmus notostictus Pocock 1910, p. 202. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus otomitus (Saussure). 

Polydesmus otomitus Saussure 1859, p. 322. 


Polydesmus (Fontaria) otomitus Saussure 1860, p. 315; Pocock 1910, 
p- 207. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Plateau and temperate regions 
(Cérdoba), Mexico. 


62 U.S. NATIONAL MUSEUM BULLETIN 266 


Rhysodesmus pater Chamberlin 1943b, p. 45. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: El Fortin, Veracruz, Mexico. (Probably Fortin de 
las Flores.) 
Rhysodesmus perotenus Chamberlin 1943b, p. 45. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Eighteen mi. north of Perote, Veracruz, Mexico. 

PARATYPE LOCALITY: Perote, Veracruz, Mexico. 
Rhysodesmus potosianus Chamberlin 1942c, p. 91. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tamazunchale, San Luis Potosi, 
Mexico 
Rhysodesmus pusillus Pocock 1909, p. 197. 

TYPE: British Mus. 

TYPE LOCALITY: Mexico. 
Rhysodesmus rubrimarginis Chamberlin 1943b, p. 46. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tacubaya, D. F., Mexico. 
Rhysodesmus salvini Pocock 1909, p. 200. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus sandersi Causey 1954, p. 64. 

TYPE: Illinois St. Nat. Hist. Surv., Urbana. 

TYPE LOCALITY: Tamazunchale, San Luis Potosi, Mexico. 
Rhysodesmus seriatus Chamberlin 1947b, p. 31. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9954. 

TYPE LOCALITY: Cieneguilla, Nuevo Leén, Mexico. 

PARATYPE LOCALITIES: Pablillo, Nuevo Leén; Saltillo-Diamante, 
Coahuila, Mexico. 
Rhysodesmus simillimus (Humbert and Saussure). 

Polydesmus simillimus Humbert and Saussure 1869, p. 150. 

Polydesmus (Fontaria) simillimus Saussure and Humbert 1872, p. 31; 
Pocock 1910, p. 207. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Santa Cruz, near Orizaba, Veracruz, Mexico. 
Rhysodesmus smithi Pocock 1910, p. 201. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Rhysodesmus stolli Pocock 1909, p. 198. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Retalhuleu, Guatemala. 
Rhysodesmus tabascensis Pocock 1909, p. 198. 

TYPE: British Mus. 

TYPE LOCALITY: Teapa, Tabasco, Mexico. 

OTHER LOCALITY: Rancho Nuevo, Chiapas, Mexico (Causey 1954). 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 63 


Rhysodesmus tacubayae Chamberlin 1943b, p. 46. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Tacubaya, D. F., Mexico. 
Rhysodesmus tepanecus (Saussure). 

Polydesmus tepanecus Saussure 1859, p. 321. 

Polydesmus(Fontaria) tepanecus Saussure 1860, p. 319; Pocock 1910, 
p: 207. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Cérdoba, Mexico. 

PARATYPE LOCALITY: Moyoapan, SantaCruz, near Orizaba, Mexico. 
Rhysodesmus tepoztlanus Chamberlin 1943b, p. 47. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Tepoztlan, Morelos, Mexico. 
Rhysodesmus toltecus (Saussure). 

Polydesmus toltecus Saussure 1859, p. 322. 

Polydesmus mayus Saussure 1859, p. 322. 

Polydesmus granulosus Saussure 1859, p. 323; Pocock 1910, p. 208. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern slope of Cordillera, Mexico 

PARATYPE LOCALITY: Edge of plateau, Orizaba Mountain, Mexico. 
Rhysodesmus totanacus (Saussure). 

Polydesmus totanacus Saussure 1859, p. 321. 

Polydesmus (Fontaria) totanacus Saussure 1860; p. 322; Pocock 
1909, p. 196. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Plateau of Anahuac, Mexico. 

PARATYPE LOCALITY: Peak of Orizaba, Mexico. 
Rhysodesmus viabilis Chamberlin 1943b, p. 47. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Five mi. west of El Folorn, Tamaulipas, Mexico. 
(Correct locality probably Forlon.) 

PARATYPE LOCALITY: Same, and south of Ciudad Victoria, Tam- 
aulipas, Mexico. 

OTHER LOCALITIES: North of Mante, Mexico (Loomis 1944); Valles, 
San Luis Potost, Mexico (Causey 1954). 
Rhysodesmus vicinus (Saussure). 

Polydesmus vicinus Saussure 1859, p. 322. 

Polydesmus (Fontaria) vicinus Saussure 1860, p. 318; Pocock 1910, 
p. 209. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Plateau of Anahuac, Mexico. 

PARATYPE LOCALITY: Oaxaca, Mexico. 
Rhysodesmus violaceus (Brolemann). 

Fontaria violacea Brolemann 1900, p. 101; Pocock 1910, p. 208. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 


64 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE AND PARATYPE LOCALITY: Guatemala. 
OTHER LOCALITY: Hacienda des Palmes, Mt. Metapan, Dept. Santa 
Ana, El Salvador (Kraus 1954b). 


Rhysodesmus zapotecus (Saussure). 

Polydesmus (Fontaria) zapotecus Saussure 1860, p. 314; Pocock 1910, 
p- 209: 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: San Andreas Tuxtla, Veracruz, 
Mexico. 
Rhysodesmus zendalus (Humbert and Saussure). 

Polydesmus zendalus Humbert and Saussure 1869, p. 150. 

Polydesmus (Fontaria) zendalus Saussure and Humbert 1872, p. 34; 
Pocock 1910, p. 210. 

TYPE: Mus. Hist. Nat. Geneva (H). 

TYPE AND PARATYPE LOCALITY: Eastern Cordillera, Mexico. 


Genus STENODESMUS Saussure 1859, p. 327 


TYPE SPECIES: Stenodesmus mexicanus Saussure, by original des- 
ignation. 
Stenodesmus mexicanus Saussure 1859, p. 327. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Cérdoba, Mexico. 


Suborder PARADOXOSOMATIDEA 
Family PARADOXOSOMATIDAE 


This family has been known for years as the Strongylosomidae or 
Strongylosomatidae but the reasons for the above usage are presented 


by Jeekel 1963a. 


Genus MESTOSOMA Silvestri 1897b, p. 3. 
TYPE SPECIES: Strongylosoma salvadori Silvestri, by original des- 


ignation. An imposing list of synonyms of this genus, none of which 
includes any Mexican or Central American species, is given by Jeekel 
1963a. 
Mestosoma isthmianum Loomis 1961, p. 107. 

WYP ESTOS ANats Mus /2no., 2646: 

TYPE LOCALITY: Alahjuela, Panama Prov., Panama. 

PARATYPE LOCALITIES: Same, and Balboa, Paraiso, and Pedro 
Miguel, Canal Zone. 
Mestosoma pulvillatum (Attems). 

Strongylosoma pulvillatum Attems 1899, p. 293. 

Habrodesmus pulvillatus Attems 1914b, p. 214; Jeekel 1963a, p. 32. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 65 


TYPE: Naturh. Mus. Wien. 

TYPE LOCALITY: Paraguay. 

In Attems 1933, p. 257, this species is listed, without additional 
details, as found in Costa Rica. This fantastic extension of range is 
quite improbable and it seems most likely that the Costa Rican speci- 
men or specimens represent an undescribed species. Some additional 
remarks on this species in South America were made by Jeekel 1963a, 


p..32. 


Genus ORTHOMORPHA Bollman 1893, p. 159. 
TYPE SPECIES: Polydesmus coarctatus Saussure, by original des- 


ignation. 
Orthomorpha coarctata (Saussure). 
Polydesmus coarctatus Saussure 1860, p. 207. 
TYPE: Mus. Hist. Nat. Geneve (H). 
TYPE LOCALITY: Considered to be in the Malay Archipelago. 
OTHER LOCALITIES: Tropicopolitan. Has been or probably will be 
found in or near most large urban areas of Mexico and Central America. 
Extensive synonymy of this introduced genus and species is given by 
Chamberlin and Hoffman 1958, and Jeekel 1963a. 
Orthomorpha festae Silvestri 1896a, p. 4. 
TYPE: Mus. Zool. Royale, Torino (H). 
TYPE LOCALITY: Forest of the Rio Cianati, Darien Prov., Panama. 
Jeekel 1963a places this species in “‘species incertae sedis.” I 
believe it to be a euryurid, possibly of the genus Pseudamplinus. 


Genus OXIDUS Cook 19llc, p. 628. 


TYPESPECIES: Fontaria gracilis Koch, by original designation. 
Oxidus gracilis (Koch). 

Fontaria gracilis Koch 1847, p. 142; Cook 191lc, p. 631. 

TYPE: Location unknown (H). 

TYPE LOCALITY: Supposed to be East Indian but collected in a 
greenhouse in Austria. 

OTHER LOCALITIES: Cosmopolitan in greenhouses; in the open in 
much of Mexico and Central America. 

Full synonymy of this introduced genus and species is given in 


Chamberlin and Hoffman 1958. 


Order CHORDEUMIDA 
KEY TO THE SUBORDERS OF CHORDEUMIDA 


1. Body segments not more than 32 and lacking longitudinal crests and 
repugnatorial poresS...ccccccessessessvescece CHORDEUMIDEA 


66 U.S. NATIONAL MUSEUM BULLETIN 266 


Body segments 39 or more with pronounced longitudinal crests and 
conspicuous repugnatorial pores............ LYSIOPETALIDEA 


Suborder CHORDEUMIDEA 


KEY TO THE FAMILIES OF CHORDEUMIDEA 





1, Segments with a prominent lateral shoulder; gonopods united by a 
large, apically divided, anterior sternite ......... DY BASIIDAE 
Segments not prominently shouldered; gonopods not united by 

a large istiemiute Guin <c-rabbaie seekerteee tte th keep ciaictee es cele eee 2 

2. Gnathochilarium with mentum divided, having a distinct promentum; 
9th male legs with 4 or 5 joints beyond sternum. CLEIDOGONIDAE 
Gnathochilarium with mentum entire; 9th male legs 3-jointed beyond 


SHOUT ares crenata cae Cle soa Meee ce TRICHOPETALIDAE 





Family CLEIDOGONIDAE 
Genus ACAKANDRA Loomis 1964a, p. 99. 


TYPE SPECIES: Acakandra austrina Loomis, by original designation. 
Acakandra austrina Loomis 1964a, p. 99. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqui Prov., Panama. 

PARATYPE LOCALITIES: Same and near Nueva California, Chiriqui 


Prov., Panama. 


Genus CAVOTA Chamberlin 1942b, p. 8. 


TYPE SPECIES: Cavota crucis Chamberlin, by original designation. 
Cavota crucis Chamberlin 1942b, p. 8. 

TYPE: Escuela Nat. Mexico City. 

TYPE AND PARATYPE LOCALITY: Gruta de Atoyac, Veracruz, 


Mexico. 


Genus CLEIDOGONA Cook 1895, p. 3. 


Cryptotrichus Packard 1883, p. 189. 

Campodes Bollman 1893, p. 120 (not C. L. Koch). 

Cleidogona Cook and Collins 1895, p. 41. 

TYPE SPECIES: Spirostrephon caesioannulatus Wood, by monotypy. 

For nearly 70 years the genus Cleidogona has been attributed to Cook 
and Collins 1895, p. 41, overlooking the fact that on an earlier page 
Cook alone proposed the name to replace Cryptotrichus Packard, pre- 
occupied in Coleoptera, for specimens from Culmana, Alabama, and 
Ocean Springs, Mississippi, which Packard identified as Cryptotrichus 
(Spirostrephon) caesioannulatus (Wood). Although Packard undoubtedly 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 67 


was wrong in identifying his specimens as the same species as those 
from Alleghany County, Pennsylvania, on which Wood founded Spiro- 
strephon caesioannulatus, use of this name forthe genotype of Cleidogona 
is required by the current rules of nomenclature. 
Cleidogona atoyaca Chamberlin 1943b, p. 34. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Atoyac, Veracruz, Mexico. 
Cleidogona leona Chamberlin 1943b, p. 34. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Desierto de los Leones, D. F., 
Mexico. 
Cleidogona mandeli Chamberlin 1952b, p. 13. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Cleidogona mexicana (Humbert and Saussure). 

Cleidogona mexicanus Cook and Collins 1895, p. 52. 

Craspedosoma mexicanum Humbert and Saussure 1869, p. 153. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 
Cleidogona moderata Causey 1957, p. 115. 

TYPE: American Mus. Nat. Hist. 

TYPE LOCALITY: Fighteen mi. south of Tamazunchale, San Luis 
Potosi, Mexico. 
Cleidogona nueva Chamberlin 1941d, p. 250. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, Nuevo Leén, 
Mexico. 
Cleidogona nueva michoacana Chamberlin 1941d, p. 251. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tancitaro, Michoacan, Mexico. 
Cleidogona propia Causey 1957, p. 119. 

TYPE: American Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: Near Puerto Las Peras, on 
Highway 15, Michoacan, Mexico. 


Cleidogona punctifer Chamberlin 1952b, p. 13. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Chichivac, Guatemala. 
Cleidogona rafaela Chamberlin 1943b, p. 35. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: San Rafael, D. F., Mexico. 
Cleidogona secreta Causey 1957, p. 119. 

TYPE: American Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: Oaxaca, Oaxaca, Mexico. 


68 U.S. NATIONAL MUSEUM BULLETIN 266 


Cleidogona tajumulco Chamberlin 1952b, p. 15. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Cleidogona zempoala Chamberlin 1943b, p. 35. 
TYPE: Collection of R. VY. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Parque Nacionale de Zempoala, 
Morelos, Mexico. 
Cleidogona zimapaniensis Causey 1957, p. 117. 
TYPE: American Mus. Nat. Hist. 
TYPE AND PARATYPE LOCALITY: Pan-American Highway, 14 mi. 


north of Zimapan, Hidalgo, Mexico. 


Genus HIRSUTOGONA Kraus 1954b, p. 329. 


TYPE SPECIES: Hirsutogona minutissima Kraus, by original desig- 
nation. 
Hirsutogona ceibana (Chamberlin). 

Cleidogona ceibana Chamberlin 1922, p. 11; Kraus 1954, p. 329; placed 

with doubt. 

TYPE: U.S. Nat. Mus., no. 822. 

TYPE LOGALITY:~ La Gerba, Honduras! 
Hirsutogona godmani (Pocock). 

Cleidogona godmani Pocock 1903, p. 52; Kraus 1954b, p. 329. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Hirsutogona minutissima Kraus 1954b, p. 332. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: Laguna de las Ranas, Dept. Sonsonate, El 
Salvador. 
Hirsutogona mirabilis Kraus 1954b, p. 329. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Monte Cristo, Metapan Mts., 
Dept. Santa Ana, Fl Salvador. 
Hirsutogona stolli (Pocock). 

Cleidogona stolli Pocock 1903, p. 52; Kraus 1954b, p. 329. 

TYPE: British Mus. 

TYPE LOCALITY: Volcan de Agua, Guatemala. 


Genus MEXICEUMA Verhoeff 1926, p. 112. 
TYPE SPECIES: Mexiceuma maculata Verhoeff, by original desig- 


nation. 

Mexiceuma maculata Verhoeff 1926, p. 112. 
Py BE: Zool. Mus., Munchen (H). 
TYPE LOCALITY: D. F., Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 69 


Genus SOLAENOGONA Hoffman 1950a, p. 91. 


TYPE SPECIES: Solaenogona guatemalana Hoffman, by original des- 
ignation. 
Solaenogona guatemalana Hoffman 1950a, p. 91. 

TYPE: U.S. Nat. Mus., no. 1883. 

TYPE AND PARATYPE LOCALITY: Between Santa Cruz del Quiché 


and Totonicapan, Guatemala. 


Family DYBASIIDAE 
Genus DYBASIA Loomis 1964a, p. 100. 


TYPE SPECIES: Dybasia humerosa Loomis, by original designation. 
Dybasia humerosa Loomis 1964a, p. 101. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: ‘‘Casita Alta,’’ Finca Lerida, near Boquete, 
Chiriqui Prov., Panama. 

PARATYPE LOCALITIES: Same and general region; Cerro Punta, 


Chiriqui Prov., Panama. 


Genus OGKOMUS Loomis 1964a, p. 102. 


TYPE SPECIES: Ogkomus divergens Loomis, by original designation. 
Ogkomus divergens Loomis 1964a, p. 104. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 

PARATYPE LOCALITIES: Same and adjacent region. 


Genus SOLEMIA Loomis 1964a, p. 104. 


TYPE SPECIES: Solemia chiriquia Loomis, by original designation. 
Solemia chiriquia Loomis 1964a, p. 105. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Finca Lerida, near Boquete, Chiriqui Prov., 


Panama. 


PARATYPE LOCALITY: Cerro Punta, Chiriqut Prov., Panama. 


Family TRICHOPETALIDAE 
Genus MEXITERPES Causey 1963, p. 235. 


TYPE SPECIES: Mexiterpes sabinus Causey, by original designation. 
Mexiterpes sabinus Causey 1963, p. 235. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Big Room, 3,000 ft. from entrance, Sdétan6 del 
Arroyo, near Los Sabinos and 8 mi. north of Valles, San Luis Potosi, 


Mexico. 


70 U.S. NATIONAL MUSEUM BULLETIN 266 


Suborder LYSIOPETALIDEA 
Family LYSIOPETALIDAE 
Genus ABACION Rafinesque 1820, p. 9. 


Spirostrephon Brandt 1841, p. 105; Hoffman and Crabill 1953, p. 81. 

Platops Newport 1844, p. 267. 

TYPE SPECIES: A bacion tesselatum Rafinesque, by monotypy. 
Abacion texense (Loomis). 

Spirostrephon texensis Loomis 1937, p. 109; Chamberlin and Hoffman 

1958, p. 110. 
Spirostrephon jonesi Chamberlin 1942d, p. 17; Chamberlin and Hoffman 
1O58 cep: LO. 

TYPE: U.S. Nat. Mus., no. 1237: 

TYPE LOCALITY: Pierce, Wharton Co., Texas. 

PARATYPE LOCALITIES: Wharton, Wharton Co.; Smith, Dallas, and 
Tarrant Counties, Texas. 

OTHER LOCALITIES: ‘‘Great Plains from Ames, Iowa, and Lincoln, 
Nebraska, through Kansas, Oklahoma and western Missouri, south as 
far as Kerr, Bandera, and Wharton Counties, Texas, east through Arkan- 
sas and Louisiana to Rankin Co., Mississippi’’ (Chamberlin and Hoffman 
1958). Monterrey and Bustamente, Nuevo Ledén, Mexico. (Reported by 
letter from Dr. Nell B. Causey.) 


Genus COLACTIS Loomis 1937, p. 120. 


TYPE SPECIES: Colactis saxetana Loomis, by original designation. 
Colactis loomisi Hoffman 1954b, p. 1. 

TYPE: American Mus. Nat. Hist. 

TYPE LOCALITY: Providencia, Durango, Mexico. 
Colactis protenta Loomis 1937, p. 126. 

TYPE: U.S. Nat. Mus., no. 1246. 

TYPE AND PARATYPE LOCALITY: On road 15 mi. north of Ense- 
nada, Baja Califomia, Mexico. 
Colactis tiburona (Chamberlin). 

Lysiopetalum tiburonum Chamberlin 1923a, p. 402; Loomis 1937, 

ps 128: 
TYPE: Mus. California Acad. Sci., no. 1259. 
TYPE LOCALITY: South end of Isla Tiburén, Sonora, Mexico. 


Genus TYNOMA Loomis 1937, p. 117. 


TYPE SPECIES: Tynoma sedecimum Loomis, by original designation. 
Tynoma messicanum Chamberlin 1943b, p. 33. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chipinqui, Nuevo Leon, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 71 


Order STEMMIULIDA 
Family STEMMIULIDAE 
Genus PROSTEMMIULUS Silvestri 1916, p- 323. 


TYPE SPECIES: Prostemmiulus mexicanus Silvestri, by original 
designation. 
Prostemmiulus annulatus Loomis 1964a, p. 106. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 
Prov., Panama. 
Prostemmiulus cincinnatus Loomis 1%64a, p. 108. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 
Prov., Panama. 
Prostemmiulus cooki Chamberlin 1922, p. 11. 

TYPE: U.S. Nat. Mus., no. 821. 

TYPE AND PARATYPE LOCALITY: Joyabaj, Guatemala. 
Prostemmiulus leucus (Chamberlin). New combination. 

Stemmiulus leucus Chamberlin 1943b, p. 33. Above combination, with 
which I agree, was suggested by R. L. Hoffman because of the dif- 
ference of generic ranges. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Atoyac, Veracruz, Mexico. 

Prostemmiulus lombardiae Chamberlin 1922, p. 10. 

TYPE: U.S. Nat. Mus., no. 820. 

TYPE LOCALITY: Lombardia, Honduras. 

Prostemmiulus mexicanus Silvestri 1916, p. 323. 
TYPE: Location unknown. 
TYPE LOCALITY: Buena Ventura (Santa Rosa), Veracruz, Mexico. 
PARATYPE LOCALITIES: Jonala, Chiapas; Jalapa, Veracruz, 
Mexico. 
Prostemmiulus modicus Silvestri 1916,, p. 324. 

TYPE: Location unknown. 

TYPE AND PARATYPE LOCALITY: Jalapa, Veracruz, Mexico. 
Prostemmiulus modicus cordovana Silvestri 1961, p. 326. 

TYPE: Location unknown. 

TYPE AND PARATYPE LOCALITY: Cérdoba, Mexico. 
Prostemmiulus oculeus Loomis 1961, p. 109. 

TYPE: U.S. Nat. Mus., no. 2647. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 

PARATYPE LOCALITY: Juan Dias, Panama Prov., Panama. 

Prostemmiulus picadoi Silvestri 1916, p. 327. 

TYPE: Location unknown. 


72 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE LOCALITY: ‘‘La Estrella,’’ Costa Rica. 

PARATYPE LOCALITY: Same and ‘‘Planton,’’ Costa Rica. 
Prostemmiulus relictus Chamberlin 1922, p. 9. 

TYPE: U.S. Nat. Mus., no. 819. 

TYPE LOCALITY: La Ceiba, Honduras. 
Prostemmiulus teres Loomis 1964a, p. 109. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: West of Finca Palo Santo, near 
Nueva California, Chiriqui Prov., Panama. 
Prostemmiulus tristani Silvestri 1916, p. 226. 

TYPE: Location unknown. 

TYPE LOCALITY: ‘“‘La Estrella,’’ Costa Rica. 

PARATYPE LOCALITY: Same and Orosi, Costa Rica. 
Prostemmiulus vallaris Loomis 1964a, p. 109. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: El Valle, Coclé Prov., Panama. 


Genus STEMMIULUS Gervais 1844b, p. 28. 


TYPE SPECIES: Julus (Stemmiulus) bioculatus Gervais, probably by 
monotypy. 

Stemmiulus bioculatus (Gervais). 

lulus (Stemmiulus) bioculatus Gervais 1844, p. 28. 

TYPE: Location unknown. 

TYPE LOCALITY: Colombia. 

Although several authors have credited this species to Panama, pri- 
marily on Silvestri’s 1896a and 1897b identifications of specimens from 
Darien Province, it now seems quite unlikely that Silvestri’s specimens 
were of this species. 

Stemmiulus canalis Chamberlin 1940a, p. 14. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 

OTHER LOCALITIES: Same and Pifia area, Canal Zone (Loomis 
1961). 

Stemmiulus marginandus Loomis 1964a, p. 110. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 
Prov., Panama. 

Stemmiulus parallelus Loomis 1964a, p. 111. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: West of Finca Palo Santo, near 


° a ee 4 
Nueva California, Chiriqui Prov., Panama. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 73 


Order JULIDA 
KEY TO THE SUBORDERS OF JULIDA. 


1. Telopodite of anterior gonopods mostly absent or rudimentary, when 
present not moveable by muscles; gonopods sunk in a pocket and 
more or less concealed; grinding plate of mandibles with a pitted 
area and adjacent fluted band or file, or if this development is 
absent, the cutting teeth are also aborted or weak; labroclypeal 
PEGGESSE Swab SiON ty 6 oi rgakos <peyasis,a REE caecn mies cuba ecRe JULIDEA 

Telopodite of anterior gonopods strongly developed and always 
moveable by muscles; gonopods not sunk in a pocket, freely ex- 
posed; grinding plate of mandibles without pits and fluting; labro- 
eiypeal PrOCESSES PIESEME 2%... es oss ale site nae PARAIULIDEA 


Suborder JULIDEA 
Family JULIDAE 
Genus BRACHYIULUS Berlese 1884, p. 1. 


TYPE SPECIES: Julus pusillus Leach, by original designation? 
Brachyiulus lusitanus Verhoeff, 1898, p. 154, fig. 28. 

TYPE: Zool. Mus. Berlin. 

TYPE LOCALITY: ‘‘Herzegowina.”’ 

OTHER LOCALITIES: North America; Tacubaya, D. F.; Patzcuaro, 
Michoacan, Mexico (Jawlowski 1930). 


Genus DIPLOIULUS Berlese 1884, p. 2. 


TYPE SPECIES: Julus boleti Koch, by original designation? 

For synonymy and bibliography of this European genus and the fol- 
lowing introduced species see Chamberlin and Hoffman 1958. 
Diploiulus latistriatus (Curtis 1845), p. 29. 

TYPE: British Mus. 

TYPE LOCALITY: England. 

OTHER LOCALITIES: North America: reported as Cylindroiulus 
frisius Verhoeff, from Xochimilco, D. F., and Patzcuaro, Michoacan, 
Mexico (Jawlowski 1930). 


Suborder PARAIULIDEA 
KEY TO THE FAMILIES OF PARAIULIDEA 


1. Mandible with 4 pectinate lamellae; gnathochilarium of similar form 
in both sexes; neither the second nor the seventh pair of male 
legs modified; first legs of male with 1] to 6 joints. 

NEMASOMATIDAE 


74 U.S. NATIONAL MUSEUM BULLETIN 266 


Mandible with 7 or 8 pectinate lamellae; gnathochilarium different 
in the two sexes; first male legs with 6 joints and much thickened, 


the second and often the seventh legs also modified. PARAIULIDAE 


Family NEMASOMATIDAE 
Genus BLANIULUS Gervais 1836, p. 435. 


TYPE SPECIES: Julus guttulatus Bosc, by original designation? 
Blaniulus guttulatus (Bosc 1791), p. 10. 

TYPE: Not known to exist. 

TYPE LOGALIVY:” France. 

OTHER LOCALITIES: North America: Cultivated areas of Canada 
and the United States. Chapultepec park, and Mexico City, D. F., Mexico 
(Chamberlin 1943b). 

For synonymy and bibliography of this introduced European species 
see Chamberlin and Hoffman 1958. 


Genus NOPOIULUS Menge 185], p. 7. 


TYPE SPECIES: Nopoiulus punctulatus Menge, by original designation? 
Nopoiulus armatus Nem 1895, p. 2. 

TYPE: Present location? 

TYPE LOCALITY: Vicinity of Prague, Czechoslovakia. 

OTHER LOCALITY: North America: Patzcuaro, Michoacan, Mexico 
(Jawlowski 1930). 
Nopoiulus minutus (Brandt 1821), p. 105. 

TYPE: Not known to exist. 

TYPE LOCALITY: ‘‘Eastern shore of Virginia’ (Say), but introduced 
there from Europe. 

OTHER LOCALITIES: North America: United States and Canada. 
Chapultepec Park, and Salazar, D. F., Mexico (Chamberlin 1943b). 

For synonymy and bibliography of this introduced species see Cham- 
berlin and Hoffman 1958. 


Family PARAIULIDAE 
Genus PARAIULUS Humbert and Saussure 1869, p. 155. 


Mexicoiulus Verhoeff 1926, p. 66. New synonymy suggested by N. 
B. Causey. 

TYPE SPECIES: Paraiulus olmecus Humbert and Saussure, by mon- 
otypy. 
Paraiulus amulensis Pocock 1903, p. 55. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Amula, Guerrero, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 75 


Paraiulus aztecus Pocock 1903, p. 56. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Paraiulus dampfi (Verhoeff). New combination suggested by N.B. Causey. 

Mexicoiulus dampfi Verhoeff 1926, p. 67, figs. 4-11. 

TYPE: Zool. Mus. Munchen (H). 

TYPE LOCALITY: Desierto de los Leones, D. F., Mexico. 
Paraiulus filicornis (Saussure). 

Julus filicornis Saussure 1860, p. 377; Pocock 1907, p. 58. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Veracruz, Mexico. 

OTHER LOCALITY: Cuernavaca, Mexico (Saussure and Humbert 1872). 
Paraiulus olmecus Humbert and Saussure 1869, p. 155. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Moyoapan, Mexico. 
Paraiulus phloibius Chamberlin 1942a, p. 60. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Cerro Tancitaro, Michoacan, 
Mexico. 
Paraiulus pueblanus Chamberlin 1943b, p. 30. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Rio Frio, Puebla, Mexico. 
Paraiulus rasilis (Karsch). 

Julus rasilis Karsch 1881, p. 17; Pocock 1907, p. 58. 

TYPE: Zool. Mus. Berlin (H). 

TYPE AND PARATYPE LOCALITY: Puebla, Mexico. 
Paraiulus rosanus Chamberlin 1943b, p. 31. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Santa Rosa, D. F., Mexico. 
Paraiulus schachti (Chamberlin). 

Thriniulus schachti Chamberlin 1941b, p. 20; 1943b, p. 31. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Tancitaro, Michoacan, Mexico. 
Paraiulus schmidti Chamberlin 1952b, p. 18. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Paraiulus stylifer Pocock 1907, p. 57. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Guatemala. 
Paraiulus tarascus (Saussure). 

Julus tarascus Saussure 1860, p. 378; Saussure and Humbert 1872, 
p- 98. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Aguengueo District, Michoacan, Mexico. 


273-687 O - 68 - 6 


76 U.S. NATIONAL MUSEUM BULLETIN 266 


Paraiulus viganus Chamberlin 1952b, p. 19. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Las Vigas, Veracruz, Mexico. 
Paraiulus zempoalus Chamberlin 1943b, p. 31. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Parque Nacional de Zempoala, 


Morelos, Mexico. 


Genus PHENIULUS Chamberlin 1943b, p. 29. 
TYPE SPECIES: Pheniulus phenotypus Chamberlin, by original desig- 


nation. 
Pheniulus mimeticus Chamberlin 1943b, p. 30. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Ixtapan del Oro, D. F’., Mexico. 
Pheniulus phenotypus Chamberlin 1943b, p. 30. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Oaxtepec, Morelos, Mexico. 


Genus THRINIULUS Chamberlin 1940b, p. 21. 
TYPE SPECIES: Paraiulus leucoclius Chamberlin, by original desig- 


nation. 
Thriniulus leucoclius (Chamberlin). 
Paraiulus leucoclius Chamberlin 1922, p. 19; 1940b, p. 21. 
TYPE: U.S. Nat. Mus., no. 829. 
TYPE LOCALITY: Actele, Guatemala. (Probably Acteld.) 
PARATYPE LOCALITIES: Same and Tectic, Santa Rosa, Guatemala. 


Order SPIROBOLIDA 
KEY TO THE SUBORDERS OF SPIROBOLIDA 


1. Inner gonopods (phallopods or posterior gonopods) connected by 
membrane and a sclerite, or only by membrane; posterior plates 


(telopodites of anterior gonopods) contiguous or nearly so along 


inmerwbasialasidesi sees jen niece cee ethan RG ONMUTE TD RA 
Inner gonopods not directly connected; posterior plates well separated 
at base eeeeeeee @eeoeoeevseevoeveeevee eee eee @ @eeeeeeeee SPIROBOLIDEA 


Suborder SPIROBOLIDEA 
KEY TO THE FAMILIES OF SPIROBOLIDEA 


1. Prozonites of some segments usually with a pair of scobinae; an- 


terior sternal plate of gonopods with apex produced distally, nearly 





MILLIPEDS OF MEXICO AND CENTRAL AMERICA v7 


equalling or surpassing tips of anterior plates (coxae or coxal 
endites), which usually are mesally produced; distal joint of inner 
gonopods elongate, usually quite slender and outwardly bira- 
Tern Sige breed eet ld = abe wade 4% Pad abase See Sota ST fees RHINOCRICIDAE 
Prozonites without scobinae; anterior sternal plate of gonopods 
transverse, subtriangular, or rarely distally produced; inner gono- 

pods usually short and stout, not biramous near apex 

2. Inner gonopods with apical and basal elements forming a continuous 


IS TUVEATI CONG LT EAN as vscccy core das mia ess iaotie ocala aot ewe calc olga noe ea SPIROBOLIDAE 
Inner gonopods with apical and basalelements not forming a continuous 


SPT MENT U ES wan 08 Fei as in (o's Rye ids 3 os IS au eee ea! allo Ron arm 8 oe RIOTS Le IE 3 

3. Inner gonopods with outer joint in a nearly continuous straight line, 

with basal elements heavy................ ALLOPOCOCKIIDAE 

Inner gonopods with each terminal joint at an angle to the coxal 

Plenent; lowest jointyvslende€r. sits . 22.850 1 ces oe Ge. 4 

4. Anterior gonopods with mesal margins long and in contact much of 

length; segment 2 usually with a spirobolus-like production below 

lateral angle offsepment? Iygac.:cf) males... MESSICOBOLIDAE 

Anterior gonopods with mesal margins short or, if elongate, gen- 

erally well separated, rarely in contact for a short distance; seg- 

ment 2 seldom much produced below lateral angle of segment 1.. 5 

5. First segment with front and back margins each side converging 

sowuward fo an’ acute “angle... 5.2 cb aels ATOPETHOLIDAE 

First segment with front and back margins each side not strongly 
converging downward; lateral margin broadly rounded. 


SPIROBOLELLIDAE 


Family ALLOPOCOCKIIDAE 


In his treatment of this family in 1960, Keeton did not mention the 
genus Arolus Chamberlin 1922 although it was described as closely 
related to Allopocockia. He also appears to have overlookedthe paper by 
Kraus 1954 in which Arlus was placed as a synonym of that genus. 
He did consider that Chelogonobolus might be a synonym of Allopocockia 
but lack of material for study caused him to withhold judgement. It 
appears to me that Arolus purulanus offers distinct connection of these 
two genera but they are still kept apart until further study of Allopocockia 
tylopus can be made. 


Genus ALLOPOCOCKIA Brolemann 1913, p. 478. 


Arolus Chamberlin 1922, p. 34; Kraus 1954, p. 336. 
TYPE SPECIES: Spirobolellus tylopus Pocock, by original designa- 


tion. 


78 U.S. NATIONAL MUSEUM BULLETIN 266 


Allopocockia purulana (Chamberlin). 

Arolus purulanus Chamberlin 1922, p. 35; Kraus 1954, p. 336. 

TYPE: U.S. Nat. Mus., no. 842. 

TYPE AND PARATYPE LOCALITY: Purulha, Guatemala. 
Allopocockia tylopus (Pocock). 

Spirobolellus tylopus Pocock 1908, p. 87; Brolemann 1913, p. 478. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Tecpan, Guatemala. 


Genus CHELOGONOBOLUS Carl 1919, p. 399. 
TYPE SPECIES: Spirobolus nahuus Saussure and Humbert, by original 


designation. 
Chelogonobolus atriculus (Pocock). 
Spirobolellus articulus Pocock 1908, p. 88; Keeton 1960b, p. 137. 
Allopocockia articulus Kraus 1954, p. 336. 
TYPE: British Mus. 
TYPE LOCALITY: Volcan de Agua, Guatemala. 
Chelogonobolus nahuus (Humbert and Saussure). 
Spirobolus nahuus Humbert and Saussure 1869, p. 154. 
Spirobolellus nahuus Pocock 1908, p. 88; Carl 1919, p. 401. 
TYPE: Mus. Hist. Nat. Geneve. 
TYPE AND PARATYPE LOCALITY: Sierra de Moyoapan, Eastern 
Cordillera, Mexico. 
Chelogonobolus rotundatus (Kraus). New combination. 
Allopocockia rotundata Kraus 1954b, p. 337. 
TYPE: Senckenberg Mus., Frankfurt. 
TYPE AND PARATYPE LOCALITY: Monte Cristo, Metapan Mts., 
Dept. Santa Ana, FE] Salvador. 


Genus SCHMIDTOLUS Chamberlin 1953a, p. 149. 


TYPE SPECIES: Schmidtolus chichivacus Chamberlin, by original 
designation. 
Schmidtolus chichivacus Chamberlin 1953a, p. 149. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Chichivac, Guatemala. 

PARATYPE LOCALITIES: Same, and Sta. Elena, near Tecpan, Guat- 
emala. 
Schmidtolus parvior Chamberlin 1953a, p. 150. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 79 


Family ATOPETHOLIDAE 


Genus ATOPETHOLUS Chamberlin 1918b, p. 167. 


Orthichelus Chamberlin and Hoffman 1950, p. 7: Hoffman and Orcutt 
1960, p. 133. 
TYPE SPECIES: Atopetholus californicus Chamberlin, by original 
designation. 
Atopetholus nigrescens (Chamberlin). 
Onychelus nigrescens Chamberlin 1923a, p. 407; Hoffman and Orcutt 
1960, p. 139. 
TYPE: Mus. California Acad. Sci., no. 1265. 
TYPE LOCALITY: Isla Coronados, Gulf of California, Mexico. 


Genus CENTRELLUS Cook 191la, p. 154. 


Toltecolus Chamberlin 1943b, p. 27; Hoffman and Orcutt 1960, p- 123. 
TYPE SPECIES: Centrellus falcatus Cook, by original designation. 
Centrellus faleatus Cook 1911a, p. 156. 
TYPE: U.S. Nat. Mus., no. 800. 
TYPE AND PARATYPE LOCALITY: Guanajuato, Mexico. 
Centrellus kerrensis (Chamberlin and Mulaik). 
Eurellus kerrensis Chamberlin and Mulaik 1941, p. 61; Hoffman and 
Orcutt 1960, p. 124. 
Toltecolus garcianus Chamberlin 1943b, p. 27; Hoffman and Orcutt 
1960, p. 124. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Raven Ranch, Kerr Co., Texas. 
OTHER LOCALITIES: Garcia, near Monterrey, Nuevo Leén, Mexico 
(Chamberlin 1943b). Real, Presidio, and Val Verde Counties, Texas 
(Hoffman and Orcutt 1960). 


Genus COMANCHELUS Hoffman and Orcutt 1960, p. 128. 
TYPE SPECIES: Toltecolus chihuanus Chamberlin, by original desig- 


nation. 
Comanchelus chihuanus (Chamberlin). 

Toltecolus chihuanus Chamberlin 1947a, p. 10; Hoffman and Orcutt 
1960, p. 128. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chihuahua City, Chihuahua, Mexico. 


Genus CYCLOTHYROPHORUS Pocock 1908, p. 83. 


TYPE SPECIES: Cyclothyrophorus salvini Pocock, by original desig- 


nation. 


80 U.S. NATIONAL MUSEUM BULLETIN 266 


Cyclothyrophorus heteropygus (Humbert and Saussure). 
Spirobolus heteropygus Humbert and Saussure 1869, p. 154; Pocock 
1908, p. 85. 
TYPE: Mus. Hist. Nat. Geneve (H). 
TYPE LOCALITY: Cuernavaca, Mexico. 
Cyclothyrophorus salvini Pocock 1908, p. 84. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Amula, Guerrero, Mexico. 
OTHER LOCALITY: Colima, Mexico (Hoffman, unpublished). 
After examination of the Colima specimens, R. L. Hoffman assured 
me that the genus is an atopetholid. 
Cyclothyrophorus valvanus (Karsch). 
Spirobolus valvanus Karsch 1881, p. 55; Pocock 1908, p. 87. 
TYPE: Zool. Mus. Berlin (H). 
TYPE AND PARATYPE LOCALITY: Puebla, Mexico. 


Genus SAUSSUROBOLUS Carl 1919, p. 389. 


TYPE SPECIES: Julus nietanus Saussure, by original designation. 
Saussurobolus neglectus Carl 1919, p. 391. 

Spirebolus nietanus Saussure and Humbert 1872, p. 89, in part. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Cuernavaca, Morelos, Mexico. 
Saussurobolus nietanus (Saussure). 

Julus nietanus Saussure 1860, p. 565. 

Spirobolus nietanus Saussure and Humbert 1872, p. 89. 

Cyclothyrophorus nietanus Pocock 1908, p. 84; Carl 1919, p. 390. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Cuernavaca, Morelos, Mexico. 
Saussurobolus zacatecus (Chamberlin). 

Arinolus zacatecus Chamberlin 1947b, p. 51; Hoffman and Orcutt 

1960, p. 147. 
TYPE: Acad. Nat. Sci. Philadelphia, no. 9974. 
TYPE LOCALITY: Sierra Temperosa del Oro, Zacatecas, Mexico. 


Genus SCOBINOMUS Loomis 1953, p. 420. 


TYPE SPECIES: Scobinomus serratus Loomis, by original designa- 
tion. 
Scobinomus serratus Loomis 1953, p. 420. 

TYPE: U.S. Nat. Mus., no. 2091. 

TYPE LOCALITY: Fourteen mi. north of Ensenada, Baja California, 
Mexico. 

PARATYPE LOCALITIES: Same, and Ceregas Canyon, 8-10 mi. from 


Ensenada, Baja California, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 81 


Genus TARASCOLUS Chamberlin 1943b, p. 25. 
TYPE SPECIES: Tarascolus bolivari Chamberlin, by original designa- 


nation. 
Tarascolus bolivari Chamberlin 1943b, p. 26. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Zitacuaro, Michoacan, Mexico. 
Tarascolus clarus Chamberlin 1943b, p. 26. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Santa Rosa, D. F., Mexico. 

PARATYPE LOCALITIES: Same, and Pedregal de San Angel, Mexico 
wor .?). 


Genus WATICHELUS Chamberlin 1949, p. 169. 
TYPE SPECIES: Onychelus smithi Chamberlin, by original designa- 


tion. 
Watichelus cooki Loomis 1949, p. 243. 
TYPE: U.S. Nat. Mus., no. 2516. 
TYPE AND PARATYPE LOCALITY: Descanso, Baja California, 
Mexico. 
Watichelus emarginatus Loomis 1949, p. 243. 
TYPE: U.S. Nat. Mus., no. 2515. 
TYPE AND PARATYPE LOCALITY: Eight mi. south of Tijuana, 


Baja California, Mexico. 


Family MESSICOBOLIDAE New 


When Hoffman and Orcutt 1960 published their synopsis of the family 
Atopetholidae they excluded three genera that had been associated with 
it at one time or another. These genera are Anelus, Cyclothyrophorus, 
and Messicobolus, which they indicated belong in one or more unde- 
fined spirobolid families. Since then Hoffman has obtained fresh material 
of Cyclothyrophorus salvini, from Mexico, and has determined that the 
genus is a true atopetholid. While it is beyond the scope of this check- 
list to go into the study necessary to determine if the two remaining 
genera belong in only a single family, these waifs require some assign- 
ment for inclusion here and the new family Messicobolidae is proposed 
for them, with Oxobolus Chamberlin 1922 also included. 

This family is characterized as composed of intermediate to rather 
large size species with anterior plates of the gonopods more or less 
produced at the mesodistal corners; a short but broad sternum (ventral 
plate); and short, stout, generally quite simple inner gonopods. 


82 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus ANELUS Cook 191la, p. 160. 


TYPE SPECIES: Anelus reduncus Cook, by original designation. 
Anelus richardsoni (Pocock). 

Spirobolellus richardsoni Pocock 1908, p. 87; Cook 191la, p. 163. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Tampico, Tamaulipas, Mexico. 


Genus MESSICOBOLUS Brolemann 1914, p. 32. 


TYPE SPECIES: Spirobolus godmani Pocock, by original designation. 
Messicobolus collaris (Chamberlin). New combination suggested by 
W. T. Keeton and approved by R. L. Hoffman. 
Aztecolus collaris Chamberlin 1953a, p. 148. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Volcdn Tajumulco, Guatemala. 
Messicobolus eximius (Porat). New combination. 
Spirobolus eximius Porat 1889a, p. 248. 
TYPE: Location unknown but possibly Nat. Hist. Mus. Brussels (H). 
TYPE AND PARATYPE LOCALITY: Guatemala. 
Included in this genus on the basis of its association withM. mystecus 
(Saussure), Carl 1919. 
Messicobolus godmani (Pocock). 
Spirobolus godmani Pocock 1908, p. 78; Brolemann 1914, p. 32. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Messicobolus hoogstraali Chamberlin 1941d, p. 254; original spelling 
hoogstralli. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY: Ojo de Aguas, Sabinas Hidalgo, 
Nuevo Leon, Mexico. 
Messicobolus hoplomerus (Pocock). 
Spirobolus hoplomerus Pocock 1908, p. 76; Carl 1918, p. 460. 
TYPE: British Mus. 
TYPE LOCALITY: Costa Cuca, Pacific slope, Guatemala. 
OTHER LOCALITY: Guatemala (Carl 1918). 
Messicobolus monticola (Pocock). New combination. 
Spirobolus monticola Pocock 1908, p. 80. 
TYPE: British Mus. 
TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Above combination made on basis of Pocock’s association of the 
species with M. mystecus. 
Messicobolus mystecus (Saussure). 
Julus mystecus Saussure 1860, p. 569. 
Spirobolus mystecus Saussure and Humbert 1872, p. 177; Carl 1919, 
p. 386. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 83 


TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: Oaxaca, Mexico. 

OTHER LOCALITY: San Andreas Tuxtla, Mexico (Pocock 1908). 
Messicobolus pilsbryi Chamberlin 1947b, p. 53. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9976. 

TYPE LOCALITY: Pablillo, Nuevo Leén, Mexico. 
Messicobolus raui Chamberlin 1941b, p. 13. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Tamazunchale, San Luis Potosi, Mexico. 
Messicobolus stolli (Pocock). 

Spirobolus stolli Pocock 1908, p. 77; Carl 1918, p. 461. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Pachuta, Guatemala. 
Messicobolus tepanecus (Saussure). New combination. 

Julus tepanecus Saussure 1859, p. 332. 

Spirobolus tepanecus Saussure and Humbert 1872, p. 88. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Cérdoba, Mexico. 

Above combination made on basis of Saussure’s illustrations. 
Messicobolus totonacus Chamberlin 1943b, p. 25. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Cumbres, Veracruz, Mexico. 
Messicobolus tzendalus (Saussure). New combination. 

Julus tzendalus Saussure 1860, p. 570. 

Spirobolus tzendalus Saussure and Humbert 1872, p. 178. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Oaxaca, Mexico. 

Above combination made on basis of Pocock’s association of the spe- 
cies with mystecusas possibly an “‘old and large example of mystecus.”’ 
Messicobolus zonatus Carl 1918, p. 462. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Guatemala. 


Species of Uncertain Position. 


The following three species, all apparently founded on single females, 
cannot be assigned definitely to any genus but, for convenience and 
until topotype males are available, are tentatively placed in Messicobolus. 


Messicobolus amulensis (Pocock). New combination. 
Spirobolus amulensis Pocock 1908, p. 79. 
TYPE: British Mus. 
TYPE LOCALITY: Amula, Guerrero, Mexico. 
Messicobolus platyops (Pocock). New combination. 
Spirobolus platyops Pocock 1908, p. 76. 


84 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE: British Mus. 

TYPE LOCALITY: Mescala, Mexico. 
Messicobolus reptans (Porat). New combination. 

Spirobolus reptans Porat 1889a, p. 250. 

TYPE: Location unknown (H). 

TYPE LOCALITY: Guanajuato, Mexico. 


Genus OXOBOLUS Chamberlin 1922, p. 30. 
TYPE SPECIES: Oxobolus virilis Chamberlin, by original designa- 


tion. 
Oxobolus cinctus Chamberlin 1922, p. 32. 
TYPE: U.S. Nat. Mus., no. 839. 
TYPE LOCALITY: Trece Aguas, Guatemala. 
Oxobolus cratus Chamberlin 1922, p. 33. 
TYPE: U.S. Nat. Mus., no. 840. 
TYPE LOCALITY: ‘‘On road to Los Pifiales,’’ Guatemala. 
Oxobolus distinctus Kraus 1954b, p. 333. 
TYPE: Senckenberg Mus., Frankfurt. 
TYPE LOCALITY: Monte Cristo, Metapan Mts., Dept. Santa Ana, El 
Salvador. 
Oxobolus magnificus Causey 1954, p. 61. 
TYPE: Illinois St. Nat. Hist. Surv., Urbana. 
TYPE LOCALITY: Three mi. northwest of Rancho Monserrate, Chiapas, 
Mexico. 
Oxobolus mundus Chamberlin 1953b, p. 69. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: San Pedro Sula, Honduras. 
Oxobolus pictus Chamberlin 1922, p. 33. 
TYPE: U.S. Nat. Mus., no. 841. 
TYPE AND PARATYPE LOCALITY: Coban, Guatemala. 
Oxobolus santanus Chamberlin 1953b, p. 69. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE LOCALITY: Head of Rio Santa Ana, in San Pedro Sula, Hon- 
duras. 
Oxobolus tardus Causey 1954, p. 61. 


TYPE: Illinois St. Nat. Hist. Surv., Urbana. 
TYPE LOCALITY: Seventeen mi. southwest of Simojovel, Chiapas, 


Mexico. 
Oxobolus virilis Chamberlin 1922, p. 31. 
TYPE: U.S. Nat. Mus., no. 838. 
TYPE AND PARATYPE LOCALITY: Samac, Coban, Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 85 


Family RHINOCRICIDAE 
Genus EURHINOCRICUS Brolemann 1903, p. 131. 


TYPE SPECIES: Rhinocricus (Eurhinocricus) biolleyi Brolemann, by 
monotypy. 
Eurhinocricus barrios Chamberlin 1953a, p. 138. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Escobas, opposite Point Barrios, 
Guatemala. 
Eurhinocricus biolleyi (Brolemann). 

Rhinocricus (Eurhinocricus) biolleyi Brolemann 1903, p. 132; Hoffman 

1953d, p. 182. 

Rhinocricus biolleyi several authors. 

Rhinocricus wheeleri Chamberlin 1922, p. 21; Hoffman 1955, p. 33. 

Rhinocricus cocos Chamberlin 1947b, p. 38; Hoffman 1953d, p. 182. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE AND PARATYPE LOCALITY: Cocos Island, Costa Rica. 

OTHER LOCALITIES: San José, and Cachi, Costa Rica (Brolemann 
1905); Barro Colorado Island, Ft. Sherman, Pita area,Canal Zone (Loomis 
1961); Almirante, Bocas del Toro Prov., Panama (Loomis 1964). 
Eurhinocricus chichivacus Chamberlin 1953a, p. 139. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Chichivac, near Tecpan, Guatemala. 
Eurhinocricus cooki Loomis 1961, p. 110. 

TYPE: U.S. Nat. Mus., no. 2648. 

TYPE AND PARATYPE LOCALITIES: Piffa area, and Barro Colorado 
Island, Canal Zone. 
Eurhinocricus eutypus Chamberlin 1953a, p. 139. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Eurhinocricus fissus Verhoeff 1937, p. 97. 

TYPE: Zool. Mus. Munchen (H). 

TYPE LOCALITY: Sierra de las Victoria, between Todos Santos and 
Miraflores, Baja California, Mexico. 
Eurhinocricus omiltemae (Pocock). 

Rhinocricus omiltemae Pocock 1907, p. 67; Hoffman 1953d, p. 182. 

TYPE: British Mus. 

TYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Eurhinocricus parvissimus Hoffman 1953d, p. 182. 

TYPE: U.S. Nat. Mus., no. 2062. 

TYPE AND PARATYPE LOCALITY: Finca Guatemoc, on Volcan 
Tacana, above Cacahuatan, Chiapas, Mexico. 
Eurhinocricus pygmoides (Chamberlin). 

Rhinocricus pygmoides Chamberlin 1933, p. 22,; Hoffman 1955, p. 35. 


86 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Parismina, Costa Rica. 
Eurhinocricus storkani Verhoeff 1937, p. 98. 

TYPE: Zool. Mus. Munchen (H). 

TYPE LOCALITY: Manzanillo (Colima?), Mexico. 


Genus OXYPYGE Silvestri 1896a, p. 4. 


TYPE SPECIES: Oxypyge varicolor Silvestri, by monotypy. 


Oxypyge benedictus Chamberlin 1925, p. 40. 

TYPE: Mus. Comp. Zool. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 
Zone. 

OTHER LOCALITIES: Same and Pifia area, Canal Zone (Loomis 1961). 
Oxypyge confusa Chamberlin 1922, p. 28. 

TYPE: U.S. Nat. Mus., no. 835. 

TYPE LOCALITY: Cacao, Guatemala. This locality probably is about 
25 km. southeast of Puerto Barrios, near the Honduran border. 
Oxypyge curticaudus Chamberlin 1953a, p. 143. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Bobos River, Izabal, Guatemala. 
Oxypyge eqialis Chamberlin 1922, p. 29. 

TYPE: U.S. Nat. Mus., no. 837. 

TYPE LOCALITY: Trece Aguas, Guatemala. 
Oxypyge isolatus Chamberlin 1925, p. 40. 

TYPE: Mus. Comp. Zool. 

TYPE AND PARATYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 
Oxypyge socia Chamberlin 1922, p. 28. 

TYPE: U.S. Nat. Mus., no. 836. 

TYPE LOCALITY: Cacao, Guatemala. See note under Q. confusa. 
Oxypyge varicolor Silvestri 1896a, p. 4. 

TYPE: Zool. Mus. Univ. Torino (H). 

TYPE LOCALITY: Punta Sabana, near Lago de Pita, Darien Prov., 


Panama. 


OTHER LOCALITY: Taboga Island, Panama Prov., Panama (Chamber- 
lin 1947c). 


Genus OXYPYGIDES Chamberlin 1922, p. 24. 


TYPE SPECIES: Oxypygides mesites Chamberlin, by original desig- 
nation. 
Oxypygides lapidicina Chamberlin 1922, p. 26. 

TYPE: U.S. Nat. Mus., no. 833. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 87 


TYPE AND PARATYPE LOCALITY: Candalaria Rocks, Scamay Es- 
tuary, Guatemala. 
Oxypygides mesites Chamberlin 1922, p. 25. 

TYPE: U.S. Nat. Mus., no. 832. 

TYPE AND PARATYPE LOCALITY: Cacao, Guatemala. See note 


under O. confusa. 


Genus RHINOCRICUS Karsch 1881, p. 68. 
TYPE SPECIES: Spirobolus (Rhinocricus) parcus Karsch, by desig- 


nation of Pocock. 
Rhinocricus angusticollis (Karsch). 
Spirobolus (Rhinocricus) angusticollis Karsch 1881, p. 70; Pocock 
1907, p. 70. 
TYPE: Zool. Mus. Berlin (H). 
TYPE AND PARATYPE LOCALITY: Puebla, Mexico. 
Rhinocricus aposematus Pocock 1907, p. 63. 
TYPE: British Mus. 
TYPE LOCALITY: Santa Clara, Costa Rica. 
Rhinocricus approximatus Hoffman 1950b, p. 69. 
Rhinocricus simulans Chamberlin 1947b, p. 39, 
Rhinocricus simulatus Chamberlin 1950, p. 6. 
TYPE: Acad. Nat. Sci. Philadelphia, no. 9963. 
TYPE LOCALITY: Nicaragua. 
Rhinocricus atoyacus Pocock 1907, p. 65. 
TYPE: British Mus. 
TYPE LOCALITY: Atoyac, Veracruz, Mexico. 
Rhinocricus aurocinctus Pocock 1907, p. 62. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: Milpas, Durango, Mexico. 
Rhinocricus aztecus (Saussure). 
Julus aztecus Saussure 1859, p. 331; Pocock 1907, p. 68. 
TYPE: British Mus. 
TYPE AND PARATYPE LOCALITY: ‘‘Veracruz, Cordoba, Orizaba, 
etc,’ Mexico. 
Rhinocricus brevicollis (Voges). 
Spirobolus brevicollis Voges 1878, p. 191; Pocock 1907, p. 69. 
TYPE: Mus. Univ. Gottingen (H). 
TYPE AND PARATYPE LOCALITY: Original spelling ‘‘Misatlan,”’ 
?Mazatlan, Mexico. 
Rhinocricus centralis Chamberlin 1922, p. 21. 
TYPE: U.S. Nat. Mus., no. 830. 
TYPE LOCALITY: Costa Rica. 
Rhinocricus chichen Chamberlin 1953a, p. 145. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Chichen Itza, Yucatan, Mexico. 


88 U.S. NATIONAL MUSEUM BULLETIN 266 


Rhinocricus chichimecus (Saussure). 

Julus chichimecus Saussure 1859, p. 331. 

Spirobolus chichimecus Saussure and Humbert 1872, p. 82; Pocock 
1907, p. 69. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE AND PARATYPE LOCALITY: Mexico. 
Rhinocricus chitarianus Chamberlin 1933, p. 23. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chitaria, Costa Rica. 
Rhinocricus costaricensis Brolemann 1905, p. 375. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE LOCALITY: Cariblanco, Costa Rica. 
Rhinocricus dugesi (Bollman). 

Spirobolus (Rhinocricus) dugesi Bollman 1893, p. 194. 

TYPE: U.S. Nat. Mus.? or lost (H). 

TYPE LOCALITY: Guanajuato, Mexico. 
Rhinocricus edenus Chamberlin 1947b, p. 39. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9962. 

TYPE LOCALITY: Eden, Nicaragua. 
Rhinocricus ferrugineus (Daday). 

Spirobolus ferrugineus Daday 1889, p. 130; Pocock 1907, p. 70. 

TYPE: Hungarian Nat. Mus., Budapest? (H). 

TYPE LOCALITY: Panama. 
Rhinocricus hagedussi (Daday). 

Spirobolus hagedussii Daday 1889, p. 130; Pocock 1907, p. 70. 

TYPE: Hungarian Nat. Mus., Budapest? (H). 

TYPE AND PARATYPE LOCALITY: Panama. 
Rhinocricus ixtapanus Chamberlin 1943b, p. 18. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Ixtapan del Oro, Mexico. 
Rhinocricus lamprus Chamberlin 1943b, p. 19. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Xilitla, San Luis Potost!, Mexico. 
Rhinocricus malkini Chamberlin 1956, p. 157. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Musawas, Waspuc River, Nicaragua. 


Rhinocricus marci Pocock 1907, p. 66. 
TYPE: British Mus. 
TYPE LOCALITY: San Marcos, Nicaragua. 
Rhinocricus mayanus Chamberlin 1947b, p. 43. 
TYPE: Acad. Nat. Sci. Philadelphia, no. 9966. 
TYPE LOCALITY: Yucatan, Mexico. 
Rhinocricus mertensi Kraus 1954b, p. 334. 


TY PE: Senckenberg Mus., Frankfurt. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 89 


TYPE LOCALITY: At 43 km. beyond Sonsonate, Dept. Sonsonate, 
El] Salvador. 

PARATYPE LOCALITIES: Same and above Dorf Conchagua, Dept. 
Cuscatlan; Finca Los Angeles, Dept. La Libertad, F1 Salvador. 
Rhinocricus morelus Chamberlin 1943b, p. 20. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: ‘‘Chapultepec,’’ Cuernavaca, 
Morelos, Mexico. 

Rhinocricus motulensis Chamberlin 1938a, p. 172. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: San Bulha Cenote, Motul, Yucatan, Mexico. 
Rhinocricus mucronatus Brolemann 1911b, p. 120. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE LOCALITY: Cache, Costa Rica. 

Rhinocricus nicaraguanus Chamberlin 1922, p. 20. 

TYPE: Mus. Comp. Zool., cat. no. 5017. 

TYPE LOCALITY: Chontales Camoapa, Nicaragua. 
Rhinocricus nodosicollis Brolemann 1905, p. 372. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE AND PARATYPE LOCALITY: Cariblanco, Costa Rica. 
Rhinocricus obesus Brolemann 1900, p. 107. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE LOCALITY: Guatemala. 

Rhinocricus obesus rubicundus Brolemann 1905, p. 376. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE AND PARATYPE LOCALITY: Cariblanco, Costa Rica. 
Rhinocricus ocraceus Brolemann 1900, p. 124. 

Rhinocricus insulatus Chamberlin 1925, p. 39; Loomis 1964a, p. 114. 

Rhinocricus williamsi Chamberlin 1940a, p. 15. 

Eurhinocricus insulatus Hoffman 1955, p. 34. 

Eurhinocricus williamsi Hoffman 1955, p. 35. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE LOCALITY: Bas Obispo, near Gamboa, Canal Zone. 

OTHER LOCALITIES: Barro Colorado Island, Canal Zone (Chamberlin 
1925); many in Canal Zone and adjacent Panama Prov. Panama (Loomis 
1961). 

Rhinocricus olivaceus (Newport). 
Spirobolus olivaceus Newport 1844, p. 268. 
Julus olivaceus Gervais 1847, p. 184; Pocock 1907, p. 66. 


TYPE: British Mus. 

TYPE LOCALITY: Oaxaca, Mexico. 
Rhinocricus pedrocola Chamberlin 1947b, p. 41. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: Pedro Miguel, Canal Zone. 


90 U.S. NATIONAL MUSEUM BULLETIN 266 


Rhinocricus plesius Chamberlin 1914, p. 187. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: Rio Oro Valley, near Cachi, Costa Rica. 
Rhinocricus potosianus Chamberlin 1941d, p. 252. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Valles,7 mi. south of El Banito, San Luis Potosi, 
Mexico. 

OTHER LOCALITIES: Xilitla, San Luis Potost (Chamberlin 1943b); 
south of Mante, Tamaulipas (Loomis 1944); Tamazunchale, San Luis 
Potost (Chamberlin 1953a; Causey 1954); near towns of El Limon, El 
Forlon, and Victoria, Tamaulipas (Chamberlin 1943b). 

Rhinocricus rixi Pocock 1907, p. 64. 

TYPE: British Mus. 

TYPE LOCALITY: Chontales Copper Mine, Nicaragua. 
Rhinocricus rogersi Pocock 1907, p. 61. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Costa Rica. 
Rhinocricus salleanus Pocock 1907, p. 64. 

TYPE: British Mus. 

TYPE LOCALITY: Cérdoba, Mexico. 

Rhinocricus scobinatus Pocock 1907, p. 65. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Retalhuleu, Guatemala. 
Rhinocricus simulans Chamberlin 1922, p. 22. 

TYPE: U.S. Nat. Mus., no. 831. 

TYPE LOCALITY: Costa Rica. 

Rhinocricus smithi Pocock 1907, p. 62. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme and Amoguileca, 
Guerrero, Mexico. 

OTHER LOCALITY: El Soledad, Oaxaca, Mexico (Causey 1954). 
Rhinocricus stolli Pocock 1907, p. 62. 

TYPE: British Mus. 

TYPE LOCALITY: Cholhuitz, Guatemala. 

OTHER LOCALITY: Coban, Guatemala (Chamberlin 1922). 
Rhinocricus tejerianus Chamberlin 1953a, p. 147. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Tejeria, Veracruz, Mexico. 
Rhinocricus toltecus (Saussure). 

Julus toltecus Saussure 1859, p. 331. 

Spirobolus toltecus Saussure and Humbert 1872, p. 75; Pocock 1907, 
p. 68. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Cordoba, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 91 


Rhinocricus totonacus (Saussure). 

Julus totonacus Saussure 1860, p. 561; 

Spirobolus totonacus Saussure and Humbert 1872, p. 176; Pocock 1907, 
p- 69. 

TYPE: Mus. Hist. Nat. Geneve. 

TY PE LOCALITY: Orizaba, Mexico. 
Rhinocricus tristani Pocock 1907, p. 63. 

TYPE: British Mus. 

TYPE LOCALITY: Santa Clara, Costa Rica. 
Rhinocricus zapotecus (Saussure). 

Julus zapotecus Saussure 1860, p. 559. 

Spirobolus zapotecus Saussure and Humbert 1872, p. 176; Pocock 
1907, p. 69. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE AND PARATYPE LOCALITY: ‘‘The same country as the pre- 


ceding,” i.e., Mexico. 


Genus YUCATOBOLUS Chamberlin 1938a, p. 173. 
TYPE SPECIES: Yucatobolus spukilensis Chamberlin, by original 


designation. 


Yucatobolus spukilensis Chamberlin 1938a, p. 173. 
TYPE: Collection of R. V. Chamberlin. 
TYPE AND PARATYPE LOCALITY :Spukil Cave,Calcehtok, Yucatan, 


Mexico. 


Genus ZIPYGE Chamberlin 1925, p. 40. 


TYPE SPECIES: Oxypyge ferruginipes Chamberlin, by original desig- 
nation. 
Zipyge ferruginipes (Chamberlin). 

Oxypyge ferruginipes Chamberlin 1922, p. 27. 

TYPE: U.S. Nat. Mus., no. 834. 

TYPE AND PARATYPE LOCALITY: Cacao, Guatemala. Probably 


about 25 km. southeast of Puerto Barrios, near Honduran border. 


Family SPIROBOLELLIDAE 
Genus MICROSPIROBOLUS Silvestri 1898b, p. 75. 


Barrobolus Chamberlin 1925, p. 41; Loomis 1964a, p. 114. 

TYPE SPECIES: Microspirobolus pulchellus Silvestri, by monotypy. 
Microspirobolus grammicus (Chamberlin). 

Barrobolus grammicus Chamberlin 1925, p. 41; Loomis 1964a, p. 115. 

TYPE: Mus. Comp. Zool. 

TYPE AND TOPOTYPE LOCALITY: Barro Colorado Island, Canal 


Zone. 


273-687 O- 68-7 


92 U.S. NATIONAL MUSEUM BULLETIN 266 


Microspirobolus trifasciatus Loomis 1964a, p. 115. 
TYPE: Chicago Nat. Hist. Mus. 
TYPE AND PARATYPE LOCALITY: Almirante, Bocas del Toro 


Prov., Panama. 


Family SPIROBOLIDAE 
Genus AZTECOLUS Chamberlin 1943b, p. 28. 
TYPE SPECIES: Spirobolus nigrior Chamberlin, by original designa- 


tion. 
Aztecolus collaris Chamberlin 1953a, p. 148. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Aztecolus nigrior (Chamberlin). 

Spirobolus nigrior Chamberlin 1941d, p. 253; 1943b, p. 28. 

? Aztecolus nigrior Chamberlin 1947b, p. 47; Keeton 1960a, p. 35. 

TYPE: Collection of R. VY. Chamberlin. 

TYPE LOCALITY: Horsetail Falls, Hacienda Vista Hermosa, Villa 
Santiago, Nuevo Leén, Mexico. 

PARATYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, Nuevo Leon, 
Mexico. 

OTHER LOCALITIES: Linares Nuevo Leén (Chamberlin 1943b); 
Pablillo, Nuevo Leén (Chamberlin 1947b); Catton de las Anahuas, 
Linares, Nuevo Ledn; 13 mi. west of Milpillas, Zacatecas, Mexico 
(Keeton 1960a). 

Aztecolus pablillo Chamberlin 1947b, P. 47. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9969. 

TYPE LOCALITY: Pablillo, Nuevo Leén, Mexico. 

OTHER LOCALITY: Five mi. west of Gomez Farias, in Sierra Madre 
Oriental, Tamaulipas, Mexico (Keeton 1960a). 


Genus HILTONIUS Chamberlin 1918b, p. 166. 
TYPE SPECIES: Hiltonius pulchrus Chamberlin, by original desig- 


nation. 
Hiltonius callipus (Bollman). 

Spirobolus callipus Bollman 1893, p. 191; Keeton 1960a, p. 114. 

TYPE: Said to be in U.S. Nat. Mus. but not found recently. 

TYPE AND PARATYPE LOCALITY: Guanajuato, Mexico. 

Keeton lists this species under ‘‘nomen inquirendum’’ but states that 
Bollman’s description is sufficient to assign the species to Hiltonius. 
Hiltonius carpinus Chamberlin 1943b, p. 21. 

Hiltonius crassus Chamberlin 1943b, p. 21; Keeton 1960a, p- 106. 

Hiltonius tepoztlanus Chamberlin 1943b, p. 21; Keeton 1960a, p. 106. 

TYPE: Collection of R. V. Chamberlin. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 93 


TYPE LOCALITY: Limén, Tamaulipas, Mexico. 

PARATYPE LOCALITIES: Same, and Tepoztlan, Morelos, Mexico. 

OTHER LOCALITIES: Ixtapan del Oro, Mexico, Mexico (Chamberlin 
1943b). 

Hiltonius carpinus vulean (Chamberlin). 

Aztecolus vulcan Chamberlin 1953a, p. 148; Keeton 1960a, p. 107. 

Rhinocricus vulcan Chamberlin 1953a, p. 144; Keeton 1960a, p. 107. 

Aztecolus fratrellus Chamberlin 1953a, p. 148; Keeton 1960a, p. 107. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 

OTHER LOCALITY: Same locality for type of A. fratrellus. 

Hiltonius erythrotypus Chamberlin 1943b, p. 22. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Santa Rosa, D.F., Mexico. 

This species is listed as “‘nomen inquirendum”’ by Keeton 1960a, p.114. 
Hiltonius fossulifer (Pocock). 

Spirobolus fossulifer Pocock 1908, p. 78; Keeton 1960a, p. 102. 

TYPE: British Mus. (presumed lost, Keeton 1960a, p. 105). 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Hiltonius hebes (Bollman). 

Spirobolus hebes Bollman 1887, p. 31. 

Tylobolus hebes Cook 1904, p. 66; Loomis and Hoffman 1948, p. 51. 

Hiltonius balboanus Chamberlin 1941b, p. 10; Loomis and Hoffman 

1948, p. 51. 

TYPE: U.S. Nat. Mus. but not found recently. 

TYPE AND PARATYPE LOCALITY: San Diego, California. 

OTHER LOCALITIES: Several in San Diego Co., California (Loomis 
and Hoffman 1948); Ceregus Canyon, Ensenada, Baja California, Mexico 
(Keeton 1960a). 

Hiltonius mexicanus (Saussure). 

Julus mexicanus Saussure 1859, p. 332. 

Spirobolus mexicanus Saussure and Humbert 1872, p. 177. 

Eurelus tancitarus Chamberlin 1941d, p. 255; Keeton 1960a, p. 100. 

Hiltonius federalis Chamberlin 1943b, p. 22; Keeton 1960a, p. 100. 

Hiltonius michoacanus Chamberlin 1943b, p. 23; Keeton 1960a, p. 100. 

Hiltonius veracruzanus Chamberlin 1943b, p. 24; Keeton 1960a, p. 100. 

TYPE: Mus. Hist. Nat. Geneve. 

TYPE LOCALITY: Mexico. 

OTHER LOCALITIES: D.F., and states of Michoacan, Morelos, and 


Veracruz, Mexico (see Keeton 1960a for exact localities). 


Genus TYLOBOLUS Cook 1904, p. 65. 


Californibolus Verhoeff 1944, p. 55; Keeton 1960a, p. 114. 
Auxobolus Chamberlin 1949, p. 163; Keeton 1960a, p. 114. 
TYPE SPECIES: Tylobolus deses Cook, by original designation. 


94 U.S. NATIONAL MUSEUM BULLETIN 266 


Tylobolus claremontus Chamberlin 1918b, p. 165. 
? Spirobolus californicus Bollman 1887, p. 50; Keeton 1960a, p. 121. 
Tylobolus stebbinsi Chamberlin 1944, p. 113; Keeton 1960a, p. 121. 
Auxobolus claremontus Chamberlin 1949, p. 163. 
Auxobolus stebbinsi Chamberlin 1949, p. 165; Keeton 1960a, p. 122. 
Auxobolus simulatus Chamberlin 1949, p. 165; Keeton 1960a, p. 122. 
TYPE: Mus. Comp. Zool. 
TYPE LOCALITY: Claremont, California. 
OTHER LOCALITIES: St. Vincent, Baja California, Mexico (Bollman 


1887). All other localities of above synonymous species in California. 


Suborder TRIGONIULIDEA 
Family TRIGONIULIDAE 


Genus SPIROSTROPHUS Saussure and Zehntner 1902, p. 150. 


TYPE SPECIES: Spirobolus naresii Pocock. 

For full synonymy and bibliography of this genus and the following 
introduced species see Loomis and Hoffman 1948, p. 52. 
Spirostrophus naresi (Pocock). 

TYPE: British Mus. 

TYPE LOCALITY: Mahé Island, Seychelles. 

LOCALITIES IN CENTRAL AMERICA: Costa Rica (Chamberlin 1922; 
Cook 191la; Loomis and Hoffman 1948). 


Order SPIROSTREPTIDA 
KEY TO THE SUBORDERS OF SPIROSTREPTIDA 


1. Both anterior and posterior gonopods present and functional, the an- 
terior pair usually with long flagellae; laminae linguales completely 
separated by the mentumauw. au destreee terse eres eC AMD AIDE A 

Posterior pair of gonopods rudimentary or completely absent, an- 


terior pair elaborate; laminae linguales not separated by the 


TING MLE MSs eee aM crete, REARS wiatobaitcl ate meet anen ne SPIROSTREPTIDEA 


Suborder CAMBALIDEA 


KEY TO THE FAMILIES OF CAMBALIDEA 


1. Mentum divided, the promentum clearly separated ... CAMBALIDAE 
Mentum entire, no separate promentum .........6-. LEIODERIDAE 


Family. CAMBALIDAE 
Genus CAMBALA Gray 1832, pl. 135. 


Eclytus Chamberlin 1952c, p. 10; preoccupied. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 95 


Eclomus Chamberlin 1952d, p. 71; Causey 1964a, p. 237. 

TYPE SPECIES: Julus annulatus Say, by monotypy. 
Cambala speobia (Chamberlin). 

Eclytus (not Holmgren 1855) speobius Chamberlin 1952c, p. 11 

Eclomus speobius Chamberlin 1952d, p- 71; Causey 1964a, p. 243. 

Cambala caeca Loomis 1953, p. 417; Causey 1964a, p- 243. 

Cambala captiosa Causey 1959, p. 69; Causey 1964a, p. 243. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Wyatt Cave, Edwards Co., Texas. 

PARATYPE LOCALITIES: Same and Felton Cave, Edwards Co., Texas. 

OTHER LOCALITIES: Found in numerous caves of central and south- 
west Texas. In letter of Nov. 18, 1964, Dr. Causey wrote that she had 
received specimens of a new subspecies of speobia from Cueva de los 
Lagos, Coahuila, Viexico, and suggested that I include the record here. 


Genus MEXICAMBALA Causey 1964a, p. 244, 


TYPE SPECIES: Mexicambala russelli Causey, by original designation. 
Mexicambala russelli Causey 1964a, p. 244. 

TYPE: American Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: Cueva de la Parra, 5 km. north 


of Xilitla, San Luis Potosi, Wexico. 


Family LEIODERIDAE 
Genus ERGENE Chamberlin 1943a, p. 4. 


TYPE SPECIES: Ergene setosus Chamberlin, by original designation. 
Ergene setosus Chamberlin 1943a, p. 5. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: ‘‘Highway No. 1 and Tropic of 


Cancer,’’ 19 mi. south of Ciudad Victoria, Tamaulipas, Mexico. 


Genus LEIODERE Loomis 1938, p. 60. 


TYPE SPECIES: L eiodere torreyana Loomis, by original designation. 
Leiodere torreyana Loomis 1938, p. 61. 

TYPE: U.S. Nat. Vus., no. 1313. 

TYPE LOCALITY: Torrey Pines, La Jolla, California. 

PARATYPE LOCALITY: Same and 20 mi. below Tijuana, Baja Cali- 


fornia, Mexico. 


Genus MORELENE Chamberlin 1943b, p. 10. 


TYPE SPECIES: Morelene mundus Chamberlin, by original designation. 
Morelene mundus Chamberlin 1943b, p. 11. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Oaxtepec, Morelos, Mexico. 


96 U.S. NATIONAL MUSEUM BULLETIN 266 


Suborder SPIROSTREPTIDEA 
KEY TO THE FAMILIES OF SPIROSTREPTIDEA 


1. Size small to intermediate, slender; gonopods relatively simple, 
lacking long, twisted, anterior branches... EPINANNOLENIDAE 

Size large, heavy bodied; gonopods quite complicated, each with a 
long, contorted, anterior branch ........ SPIROSTREPTIDAE 


Family EPINANNOLENIDAE 
Genus EPINANNOLENE Brolemann 1903, p. 135. 


TYPE SPECIES: Epinannolene pittieri Brolemann, by monotypy. 
Epinannolene affinis Loomis 1961, p. 114. 

TYPE: U.S. Nat. Mus., no. 2649. 

TYPE LOCALITY: Piiva area, Canal Zone. 

PARATYPE LOCALITIES: Same, and El Valle, Coclé Prov., Panama. 


Epinannolene bicornis Brolemann 1905, p. 356. 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE LOCALITY: Cariblanco, Costa Rica. 
Epinannolene caeca Loomis 1964a, p. 120. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
Epinannolene pittieri Brolemann 1903, p. 136 

TYPE: Mus. Nat. Hist. Natur., Paris. 

TYPE AND PARATYPE LOCALITY: Cocos Island, Costa Rica. 
Epinannolene plana Loomis 1961, p. 116. 

TYPE: U.S. Nat. Mus., no. 2651. 

TYPE AND PARATYPE LOCALITY: El Valle, Cocle Prov., Panama. 
Epinannolene robusta Loomis 1961, p. 115. 

TYPE: U.S. Nat. Mus., no. 2650. 

TYPE AND PARATYPE LOCALITY: Cerro Campana, Panama Prov., 
Panama. 
Epinannolene suleata Loomis 1964a, p. 120. 

TY PE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: West of Finca Palo Santo, near Nueva California, 


ey 
Chiriqui Prov., Panama. 


Family SPIROSTREPTIDAE 
Genus GYMNOSTREPTUS Brolemann 1901, p. 153. 


TYPE SPECIES: Spirostreptus (Gymnostreptus) perfidus Brolemann 
1901, by present designation. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA Oy 


Gymnostreptus laetus Chamberlin 1922, p. 13. 

TYPE: U.S. Nat. Mus., no. 823. 

TYPE AND PARATYPE LOCALITY: Joyabaj, Guatemala. 
Gymnostreptus vagans Chamberlin 1922, p. 13. Return to original 
combination. 

Epistreptus (Microtrulius) vagans Attems 1950, ps 220). 

Gymnostreptus pacificus Chamberlin 1922, p. 14; Attems 1950, p. 220. 

Epistreptus (Microtrulius) pacificus Attems 1950, p. 220. 

TYPE: U.S. Nat. Mus., no. 824. 

TYPE LOCALITY: Candalaria Rocks, Scamay Estuary, Guatemala. 

PARATYPE LOCALITY: Trece Aguas, Guatemala. 

OTHER LOCALITIES: Santo Domingo de San Mateo, Costa Rica 
(Chamberlin 1922 for G. pacificus); various localities in nine departments 
of El Salvador (Kraus 1954b). 

On the basis of the structure of the gonopods it seems necessary to 
return this species to its original designation. 


Genus ISOPOROSTREPTUS Silvestri 1898b, p. 73. 


Epistreptus Carl 1914, p. 865 (not Silvestri). 

Pemptoporus Attems 191]4a, p. 121; Hoffman 1953a, p. 150. 

Scaphiostreptus Attems 1950, pp. 230, 236, in part (not Brolemann); 

Hoffman 1953a, p. 150. 

TYPE SPECIES: /soporostreptus villicola Silvestri, by monotypy., 
Isoporostreptus pittieri Hoffman 1953a, p. 151. 

TYPE: U.S. Nat. Mus., no. 2018. 

TYPE LOCALITY: Los Siguas, Chiriqui Prov., Panama. 


Genus ORTHOPORUS Silvestri 1897a, p. 7. 


Diaporus Silvestri 1897a, p. 8; Pocock 1909, p- 93. 

Scaphiostreptus Brolemann 1901, p. 150; Pocock 1909, p. 93. 

TYPE SPECIES: Orthoporus diaporoides Silvestri, by monotypy. 
Orthoporus absconsus Chamberlin 1922, p. 15. Return to original designa- 
tion. 

Scaphiostreptus absconsus Attems 1950, p. 231. 

TYPE: U.S. Nat. Mus., no. 826. 

TYPE LOCALITY: Santo Domingo de San Mateo, Pacific side, Costa 
Rica. 

OTHER LOCALITIES: San José, Costa Rica(Chamberlin 1933); seven 
departments. E] Salvador (Kraus 1954b); Zamorano, Tegucigalpa, 
Honduras (Loomis 1959b). 

Orthoporus acanthethus Chamberlin 1947b, p. 57. 

TYPE: Acad. Nat. Sci. Philadelphia, no. 9980, 

TYPE LOCALITY: Piedras Negras, Guatemala. 


98 U.S. NATIONAL MUSEUM BULLETIN 266 


Orthoporus ampussis (Karsch). 
Spirostreptus ampussis Karsch 1881, p. 43; Pocock 1909, p. 102. 
TYPE: Zool. Mus. Berlin (H). 
TYPE LOCALITY: Puebla, Mexico. 
Orthoporus asper (Attems). New combination. 
Scaphiostreptus asper Attems 1950, p. 234. 
TYPE: Naturh. Mus. Wien. (H). 
TYPE LOCALITY: Mexico. 
Orthoporus bisuleatus Chamberlin 1952b, p. 21. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Tejeria, Veracruz, Mexico. 
Orthoporus bobos Chamberlin 1952b, p. 23. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Bobos River, I[zabal, Guatemala. 
Orthoporus canalis Chamberlin 1925, p. 38. 
TYPE: Mus. Comp. Zool. 
TYPE LOCALITY: Barro Colorado Island, Canal Zone. 


OTHER LOCALITIES: Same, Monte Lirio, and Pina area, Canal Zone 
(Loomis 1961). 
Orthoporus capucinus (Attems). New combination. 

Scaphiostreptus capucinus Attems 1950, p. 234. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE AND PARATYPE LOCALITY: Mexico 
Orthoporus chihuanus Chamberlin 1947a, p. 11. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Chihuahua, Chihuahua, Mexico. 

Suspected by Loomis 1963, p. 125, of being a synonym of punctilliger, 
itself a synonym of ornatus. 
Orthoporus chiriquensis Pocock 1909, p. 97. 

Diaporus chiriquensis Chamberlin 1922, p. 18; Loomis 1964a, p. 118. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Volcan de Chiriqut, Chiriqui, 
Panama. 

The species redescribed by Attems 1950, p. 244, as Scaphiostreptus 
(Diaporus) chiriquensis is not that species. 
Orthoporus cienegonus Chamberlin 1952b, p. 23. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Cienega de Flores, Nuevo Leon, 
Mexico. 
Orthoporus cobanus Chamberlin 1922, p. 16. 

TYPE: U.S. Nat. Mus., no. 828. 

TYPE LOCALITY: Cobdan-Caché, Guatemala. 

PARATYPE LOCALITIES: Sepacuite; Secanquim; Joyabaj; Trece 


Aguas, Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 99 


Orthoporus comminutus (Attems). New combination. 

Scaphiostreptus comminutus Attems 1950, p. 239. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE AND PARATYPE LOCALITY: Mexico. 
Orthoporus confragosus (Karsch). 

Spirostreptus confragosus Karsch 1881, p. 44. 

Spirostreptus (Scaphiostreptus) confragosus Brolemann 1905, p. 367; 

Pocock 1909, p. 101. 

TYPE: Zool. Mus. Berlin (H). 

TYPE LOCALITY: San José, Costa Rica. Additional specimens 
Brolemann 1905. 
Orthoporus conifer (Attems). New combination. 

Scaphiostreptus (Diaporus) conifer Attems 1950, p. 243. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE LOCALITY: Pons, Costa Rica. 
Orthoporus cordovanus Pocock 1909, p. 98. 

?Spirostreptus otomitus Saussure 1859, p. 330. 

TYPE: British Vus. 

TYPE LOCALITY: Cordoba, Mexico. 
Orthoporus discriminans Chamberlin 1922, p. 16. Return to original 
designation. 

Scaphiostreptus (S.) discriminans Causey 1960, p. 278. 

TYPE: U.S. Nat. Mus., no. 827. 

TYPE LOCALITY: Pancajché, Guatemala. 

OTHER LOCALITY: Cueva Lanquin, Alta Verapaz, Guatemala (Causey 
1960). 
Orthoporus dybasi (Chamberlin). 

Scaphiostreptus dybasi Chamberlin 1952b, p. 28; Causey 1964b, p. 175. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: El Fortin, Veracruz, Mexico. 

OTHER LOCALITIES: Veracruz-Jalapa Highway, near Cunejos; El 
Fortin, Pasada Loma, Veracruz, Mexico (Causey 1964b). 
Orthoporus esperanzae Chamberlin 1943b, p. 13. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Esperanza, Guanajuato, Mexico. 
Orthoporus euthus Chamberlin 1952b, p. 25. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Esquintla, Tiquisate, Guatemala. 
Orthoporus festae (Silvestri). 

Plusioporus festae Silvestri 1896a, p. 3; Pocock 1909, p. 102. 

Diaporus culebrae Chamberlin 1922, p. 18; Loomis 1961, p. 114. 

Diaporus barroensis Chamberlin 1925, p. 39; Loomis 1961, p. 114. 

TYPE: Mus. Zool. Univ. Torino (H). 


100 U.S. NATIONAL MUSEUM BULLETIN 266 


TYPE LOCALITY: Punta Sabana, Darien, Panama. 

OTHER LOCALITIES: Canal Zone: Culebra (Chamberlin 1922); 
Barro Colorado Island (Chamberlin 1925); Barro Colorado Island, Frijoles, 
Monte Lirio, Pina area (Loomis 1961). Panama Prov.: Juan Dfas and 
Taboga Island (Loomis 1961). Chiriquf Prov.: Boquete, Panama (Loomis 
1961). 

Orthoporus fortinus Chamberlin 1943b, p. 13. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: El Fortin, Veracruz, Mexico. 
Orthoporus fraternus (Saussure). 

Julus fratermus Saussure 1860, p. 374. 

?Spirostreptus fraternus Saussure and Humbert 1872, p. 70; Pocock 
1909, p. 101. 

Orthoporus amulensis (?Spirostreptus fraternus) Pocock 1909, p. 98; 

Causey 1964b, p. 177. 

Orthoporus morelus Chamberlin 1943b, p. 16; Causey 1964b, p. 177. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Yautepec, Morelos, Mexico. 

OTHER LOCALITIES: Amula, Guerrero (Pocock 1909); Tepoztlan, 
Morelos (Chamberlin 1943b); 8mi. east of Yautepec, and near Amacuzac, 
Morelos; between Jalapa and Banderilla, Veracruz; Oaxaca, Oaxaca; 
near Tehuitzingo, Puebla (Causey 1964b), Mexico. 

Orthoporus gracilior Chamberlin 1952b, p. 25. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Orthoporus guerreronus (Chamberlin). 

Gymnostreptus guerreronus Chamberlin 1942b, p. 8; 1943b, p. 13. 

TYPE: Escuela Nat. Mexico City. 

TYPE AND PARATYPE LOCALITY: Cueva de Jutxlahuaca, Guerrero, 
Mexico. 

Orthoporus hoctunicolens Chamberlin 1938a, p. 170. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Hoctun Cave, Hoctun, Yucatan, Mexico. 
Orthoporus kiemi Loomis 1962, p. 48. 

TYPE: U.S. Nat. Mus., no. 2778. 

TYPE LOCALITY: Cave on Hacienda San Bernardo, 5 mi. from San 
Bernardo Sta., railroad between Merida and Maxcanu, Yucatan, Mexico. 
Orthoporus leius Chamberlin 1943b, p. 14. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Esperanza, Guanajuato, Mexico. 

Orthoporus lenonus Chamberlin 1943b, p. 14. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Two mi. north of Lenon, Tamaulipas, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 101 


Orthoporus leonicus Chamberlin 1941d, p. 252. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Ojo de Agua, Sabinas Hidalgo, 
Nuevo Leon, Mexico. 

OTHER LOCALITIES: Garcia, Nuevo Leén; Xilitla, San Luis Potosi, 
Mexico (Chamberlin 1943b). 

Thought possibly a synonym of O. punctilliger (itself a synonym of O. 
oratus) by Loomis 1963, p. 124. 
Orthoporus linares Chamberlin 1943b, p. 15. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Linares, Nuevo Leon, Mexico. 
Orthoporus luchilicolens Chamberlin 1938a, p. 169. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Luchil Cave, Tixcacal, Yucatan, 
Mexico. 
Orthoporus margarites (Chamberlin). 

Diaporus margarites Chamberlin 1946, p. 146, Loomis 1964a, p. 131. 

TYPE: American Mus. Nat. Hist. 

TYPE AND PARATYPE LOCALITY: San José Island, Pearl Islands, 
Panama Prov., Panama. 
Orthoporus mimus Chamberlin 1943b, p. 15. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Nineteen mi. north of El Limon, Tamaulipas, 
Mexico. 
Orthoporus montezumae (Saussure). 

Julus montezumae Saussure 1859, p. 330. 

? Spirostreptus montezumae Saussure and Humbert 1872, p. 69; Pocock 

1909, p. 99. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Orizaba, Mexico. 

OTHER LOCALITY: Veracruz, Mexico (Saussure and Humbert 1872). 
Orthoporus mundus Chamberlin 1942a, p. 60. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Apatzingan, La Majada, Michoacan, Mexico. 
Orthoporus nesiotes Chamberlin 1923a, p. 405. 

TYPE: Mus. California Acad. Sci., no. 1261. 

TYPE AND PARATYPE LOCALITY: South end of Tiburon Island, 
Gulf of California, Sonora, Mexico. 

Suspected by Loomis 1963, p. 125, of being asynonym of O. punctil- 
liger, itself a synonym of O. omatus. 
Orthoporus nicaraguanus Chamberlin 1925, p. 38. 

TYPE: Mus. Comp. Zool. 

TYPE LOCALITY: Nicaragua. 


102 U.S. NATIONAL MUSEUM BULLETIN 266 


Orthoporus omalopyge (Brolemann). Return to original combination. 
Spirostreptus (Scaphiostreptus) omalopyge Brolemann 1905, p. 365; 
Pocock 1909, p. 103. 


Diaporus omalopyge Chamberlin 1922, p. 19. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE LOCALITY: La Palma, Costa Rica. 

PARATYPE LOCALITIES: Carillo and Cache, Costa Rica. 

OTHER LOCALITY: E1 Gallito, Costa Rica (Chamberlin 1933). 
Orthoporus ornatus (Girard). 

Julus ornatus Girard 1853, p. 274. 

Spirobolus ornatus Wood 1865, p. 208. 

Spirostreptus montezumae Bollman 1888, p. 343. 

Orthoporus punctilliger Chamberlin 1923a, p. 405. 

Orthoporus wichitanus Chamberlin 1931, p. 99; Causey 1964b, p. 177. 

Orthoporus vallicolens Chamberlin 1943c, p. 149; Loomis 1963, p.124. 

Orthoporus producens Chamberlin 1947a, p. 11; Loomis 1963, p. 124. 

Orthoporus crotonus Chamberlin 1952b, p. 24; Loomis 1963, p. 124. 

Orthoporus arizonicus Loomis 1953, p. 418; Loomis 1963, p. 124. 

TYPE: Probably no longer existing. 

TYPE LOCALITY: Determined as probably in Randal Co., Texas, by 
Causey, Tulane Stud. Zool., vol. 2, pp. 67-8, 1954. 

OTHER LOCALITIES: A very widely distributed species; in the 
United States known from Arizona, New Mexico, Oklahoma, and Texas; 
in Mexico from Chihuahua, Nuevo Ledn, San Luis Potosi, Sonora, and 
Tamaulipas. 

Orthoporus otomitus (Saussure). 
Julus otomitus Saussure 1859, p. 330. 
Spirostreptus otomitus Saussure and Humbert 1872, p. 69; Pocock 1909, 
p- 100. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Cérdoba, Veracruz, Mexico (first locality named). 

PARATYPE LOCALITIES: Veracruz and San Andrea Tuxtla, Vera- 
cruz, Mexico. 

Orthoporus palmensis (Brolemann). 
Spirostreptus (Scaphiostreptus) typotopyge palmensis Brolemann 1905, 
p- 362; Pocock 1909, p. 96. 

TYPE AND PARATYPE LOCALITY: La Palma, Costa Rica. 
Orthoporus paxillicauda Loomis 1962, p. 50. 

TYPE: U.S. Nat. Mus., no. 2779. 

TYPE AND PARATYPE LOCALITY: Zapotilla, Lake Izabal, Guate- 
mala. 

Orthoporus rodriguezi (Brolemann). 
Spirostreptus rodriguezi Brolemann 1900, p. 104; Pocock 1909, p. 103. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 103 


TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE AND PARATYPE LOCALITY: Guatemala. 
Orthoporus rodriguezi coriaceus (Brolemann). 

Spirostreptus rodriguezi coriaceus Brolemann 1900, p. 106; Pocock 

1909, p. 103. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE LOCALITY: Guatemala. 
Orthoporus rugiceps (Attems). New combination. 

Scaphiostreptus rugiceps Attems 1950, p. 240. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE AND PARATYPE LOCALITY: Mexico. 
Orthoporus rugiceps reimoseri (Attems). New combination. 

Scaphiostreptus rugiceps reimoseri Attems 1950, p. 241. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE LOCALITY: Bebedero, Costa Rica. 
Orthoporus salvadoricus (Kraus). New combination. 

Scaphiostreptus salvadoricus Kraus 1954, p. 337. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE LOCALITY: La Rabida, San Salvador, Dept. San Salvador, FE] 
Salvador. 

PARATYPE LOCALITIES: Departments of Santa Ana, Cuscatlan, 
San Vincente, and Usulutan, EF] Salvador. 
Orthoporus solicolens Chamberlin 1938a, p. 170. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Hoctun Cave, Hoctun, Yucatan, Mexico. 
Orthoporus striatulus Pocock 1909, p. 99. 

Orthoporus chiapasus Chamberlin 1943b, p. 12; Causey 1964b, p. 179. 

TYPE: British Mus. Now lost (H). 

TYPE LOCALITY: Tuxtla, Mexico. Causey, loc. cit., states that 
‘‘The records indicate that this is surely Tuxtla Gutiérrez,’’ Chiapas. 

OTHER LOCALITIES: Huixtla, Chiapas (Chamberlin 1943b); Tuxtla 
Gutiérrez and vicinity, also San Jerénimo, near Tapachula, Chiapas; Pan- 
American Highway near Chiapas-Oaxaca boundary, Tehuantepec, Oaxaca; 
west bank of Rio de Sierra, Jalapa, Tabasco (Causey 1964b). 
Orthoporus teapensis Pocock 1909, p. 97. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Teapa, Tabasco, Mexico. 

OTHER LOCALITIES: Tuxtla Gutiérrez and vicinity, probably also at 
San Jeronimo, near Tapachula, Chiapas; Pan-American highway near 


Oaxaca-Chiapas boundary, and Tehuantepec, Oaxaca, Mexico (Causey 
1964b). 


104 U.S. NATIONAL MUSEUM BULLETIN 266 


Orthoporus tehuacanus Chamberlin 1952b, p. 26. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Tehuacan, Puebla, Mexico. 
Orthoporus texicolens Chamberlin 1938b, p. 207. 
Scaphiostreptus texicolens Chamberlin 1952b, p. 28; Causey 1964b, 
p- 180. 

Orthoporus victorianus Chamberlin 1943b, p. 18; Causey 1964b, p. 180. 

TYPE LOCALITY: Edinburg, Hidalgo Co., Texas. 

OTHER RECORDS: Helotes, Bexar Co., Texas; 3 mi. South of Vic- 
toria, and 5 mi. west of El Forlén, Tamaulipas, Mexico (Chamberlin 
1943b); Brownsville, Cameron Co., Texas (Chamberlin 1953); Hidalgo, 
Hidalgo Co., Texas (Loomis 1963). One or two localities in each of the 
following Texas counties—Cameron, Jim Wells, Karnes, San Patricio, 
Uvalde, Victoria, and Wilson (Causey 1964b). 

Orthoporus tizamensis Chamberlin 1938a, p. 168. 

TYPE: Collection of R. V. Chamberlin 

TYPE LOCALITY: Muruztum Cave, Tizamin, Yucatan, Mexico. 
Orthoporus torreonus Chamberlin 1943b, p. 16. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Twenty mi. southwest of Torreon, Coahuila, 
Mexico. 

PARATYPE LOCALITIES: Same and 15 mi. southwest of Torreon, 
also 15 mi. west of Torreon, Coahuila, Mexico. 

Orthoporus trisuleatus (Daday). New combination. 

Spirostreptus trisulcatus Daday 1889, p. 127. 

TYPE: Hungarian Nat. Mus., Budapest? (H) 

TYPE LOCALITY: Panama. 

Orthoporus typotopyge (Brolemann). New combination. 
Spirostreptus (Scaphiostreptus) typotopyge Brolemann 1905, p. 359; 
Pocock 1909, p. 95. 

Diaporus typotopyge Chamberlin 1922, p. 19. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE LOCALITY: Probably Surubres, near San Mateo, Costa Rica. 

PARATYPE LOCALITIES: Cache, Cariblanco, La Palma, Costa Rica. 
Orthoporus ugmalanus Chamberlin 1943b, p. 17. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Ugmala, Guerrero, Mexico. 

PARATYPE: Without locality. 

Orthoporus unciferens Chamberlin 1952b, p. 27. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: San Marcos, El Porvenir, Guatemala. 
Orthoporus unicus Causey 1954, p. 55. 
TYPE: Illinois St. Nat. Hist. Surv., Urbana. 
TYPE LOCALITY: Two mi. northwest of Rancho Nuevo, Chiapas, 


Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 105 


PARATYPE LOCALITIES: Same and Trinitaria, Chiapas, Mexico. 
Orthoporus zizicolens (Chamberlin). New combination. 

Gymnostreptus zizicolens Chamberlin 1938a, p. 168. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Ziz Cave, Oxkutzcab, Yucatan, 


Mexico. 
Superorder COLOBOGNATHA 


The two most recent revisionary papers on the superorder Colobognatha 
are Verhoeff 194] and Attems 1951. The first proposes numerous nomen- 
clatorial changes, principally in genera of the family Siphonophoridae, 
and adds no newspecies from the area encompassed by this list. Attems’ 
paper is a revision of the superorder but it omitted mention of so many 
previously described genera, and contained such obvious errors, as well 
as departures from the rules of nomenclature, that it is a most unsatis- 
factory treatment. To produce a workable and acceptable classification, 
both papers needathorough study by someone with a large reference col- 
lection of species at hand. The proposals in these papers have been 
limited here to inclusion of the two species described by Attems from 
Costa Rica. 


KEY TO THE ORDERS OF COLOBOGNATHA 
1, Gnathochilarium having the usual parts represented in millipeds; seg- 


ments with a distinctly impressed median sulcus. 


PLATYDESMIDA 

Gnathochilarium represented by a triangular plate, or the usual di- 
visions indistinctly indicated at most; segments without an im- 
pressed median SulcuS...sceceeceeccccscccccces POLY ZONIIDA 


Order PLATYDESMIDA 
KEY TO THE FAMILIES OF PLATYDESMIDA 


1. Sterna narrow, the coxae close together separated only by a prominent 


BLETH Al APFOGESS \ateicve cllclinia ele 's € dias a,clalsle als sicle.\e ANDROGNATHIDAE 
Sterna very broad and lacking median processes; coxae very widely 
separated ereeeeee eeoeoeeeeeeeeeeeeoeeeeoseeese PL ATYDESMIDAE 


Family ANDROGNATHIDAE 
Genus BRACHYCYBE Wood 1864, p. 187. 


TYPE SPECIES: Brachycybe lecontei Wood, by monotypy. 
Brachycybe producta Loomis 1936, p. 367. 

TYPE AND PARATYPE LOCALITY: Baja California, Mexico. 

OTHER LOCALITY: Vernal Falls, Yosemite National Park, Mariposa 
Co., California, (Causey 1955). 


106 U.S. NATIONAL MUSEUM BULLETIN 266 


Family PLATYDESMIDAE 


Genus DESMETHUS Chamberlin 1922, p. 6. 

TYPE SPECIES: Desmethus ‘setifer Chamberlin, by original designa- 
nation. 
Desmethus chiriquensis Loomis 1964a, p. 126. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqui Prov., Panama. 
Desmethus setifer Chamberlin 1922, p. 6. 

TYPE: U.S. Nat. Mus., no. 814. 

TYPE AND PARATYPE LOCALITY: San Rafael, Guatemala 


Genus PLATYDESMUS Lucas 1843, p. 51. 


Piestodesmus Lucas 1849, p. 598; Pocock 1903, p. 42. 

TYPE SPECIES: Platydesmus polydesmoides Lucas, by monotypy. 
Platydesmus analis Pocock 1903, p. 46. 

TYPE: British Mus. 

TYPE LOCALITY: ?Guatemala City, Guatemala. 
Platydesmus calus Chamberlin 1952b, p. 30. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Veracruz Penuela, Veracruz, Mexico. 

PARATYPE LOCALITIES: Same, and El Fortin, Veracruz, Mexico. 
Platydesmus cerrobius Chamberlin 1942a, p. 57. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Cerro Tancitaro, Michoacan, 
Mexico. 
Platydesmus corzoi Chamberlin 1943b, p. 10. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Rio Frio, D. F., Mexico. 

PARATYPE LOCALITIES: Same, and Desierto de los Leones, D.F.; 
Rio Frio, Puebla, Mexico. 
Platydesmus crucis Chamberlin 1952b, p. 31. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Las Vigas, Veracruz, Mexico. 
Platydesmus excisus Chamberlin 1952b, p. 31. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Sierra Santa Elena, Guatemala. 

PARATYPE LOCALITIES: Same and Tecpan, Guatemala. 
Platydesmus guatemalensis Brolemann 1900, p. 13. 

Platydesmus guatemalae Chamberlin 1922, p. 6. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE LOCALITY: Guatemala. 
Platydesmus hirudo Pocock 1903, p. 46. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 107 


Platydesmus interruptus Chamberlin 1922, p. 5. 

TYPE: U.S. Nat. Mus, no. 812. 

TYPE LOCALITY: San Juan Pueblo, Honduras. 

PARATYPE LOCALITY: Same and La Ceiba, Honduras. 
Platydemus interruptus simplex Chamberlin 1922, p. 5. 

TYPE: U.S. Nat. Mus., no. 813. 

TYPE LOCALITY: La Ceiba, Honduras. 

PARATYPE LOCALITIES: Same and San Juan Pueblo, Honduras. 
Platydesmus lankesteri Brolemann 1905, p. 354. 

TYPE: Mus. Nat. Hist. Natur., Paris (H). 

TYPE AND PARATYPE LOCALITY: EI Reventado, Costa Rica. 

OTHER LOCALITIES: Hacienda Buena Vista, Volcan Izalco, Dept. 
Sonsonate; Finca Alicante, Cumbre de Santa Tecla, Dept. La Libertad, 
El Salvador, (Kraus 1954b). 
Platydesmus lineatus Pocock 1903, p. 46. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Volcan de Orizaba, Mexico. 
Platydesmus marmoreus Pocock 1903, p. 47. 

TYPE: British Mus. 

TYPE LOCALITY: Cholhuitz, Guatemala. 

OTHER LOCALITY: Monte Cristo, Matapan Mts., Dept. Santa Ana, F1 
Salvador (Kraus 1954b). 
Platydesmus mesomelas Pocock 1903, p. 46. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Omilteme, Guerrero, Mexico. 
Platydesmus mexicanus Humbert and Saussure 1869, p. 156. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 

PARATYPE LOCALITIES: Same and Sierra de Agua, Mexico. 
Platydesmus moreleti (Lucas). 

Piestodesmus moreletii Lucas 1849, p. 599; Pocock 1903, p. 48. 

TYPE: Location unknown (H). 

TYPE AND PARATYPE LOCALITY: Tabasco, Mexico. 
Platydesmus nicaraguae Chamberlin 1956, p. 158. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Musawas, Waspuc River, 


Nicaragua. 


Platydesmus nicaraguanus Chamberlin 1925, p. 37. 
TYPE: Mus. Comp. Zool. 
TYPE LOCALITY: Nicaragua. 

Platydesmus perditus Chamberlin 1952b, p. 33. 
TYPE: Collection of R. V. Chamberlin. 
TYPE LOCALITY: Guatemala. 


273-687 O - 68 - 8 


108 U.S. NATIONAL MUSEUM BULLETIN 266 


Platydesmus perpictus Pocock 1903, p. 47. 

TYPE: British Mus. 

TYPE LOCALITY: Senahu, Guatemala. 

PARATYPE LOCALITIES: Cholhuitz, Guatemala, and Belize, British 
Honduras. 

OTHER LOCALITY: Chichivac, near Tecpan, Guatemala (Chamberlin 
1952b). 
Platydesmus polydesmoides Lucas 1843, p. 52. 

TYPE: Location unknown (H). 

TYPE LOCALITY: Guatemala. 
Platydesmus subovatus Loomis 1964a, p. 127. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Almirante, Bocas del Toro Prov., Panama. 
Platydesmus taureus Loomis 1964a, p. 128. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Negra Creek, Almirante, Bocas del Toro Prov., 
Panama. 
Platydesmus triangulifer Pocock 1903, p. 45. 

TYPE: British Mus. 

TYPE AND PARATYPE LOCALITY: Volcan de Acatenango, 
Guatemala. 


OTHER LOCALITY: Izabal, Escobas, Guatemala (Chamberlin 1952b). 


Genus RHAPHONOTUM Attems 1951, p. 198. 

TYPE SPECIES: Rhaphonotum tuberculosum Attems, by present 
designation. 
Rhaphonotum reimoseri Attems 1951, p. 198. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE AND PARATYPE LOCALITY: Costa Rica. 
Rhaphonotum tuberculosum Attems 1951, p. 198. 

TYPE: Naturh. Mus. Wien. (H) 

TYPE AND PARATYPE LOCALITY: Hambourg Farm, Costa Rica. 


Order POLYZONIIDA 
KEY TO THE FAMILIES OF POLYZONIIDA 


1. Eyes absent; head narrowed forward or prolonged into a beak; pro- 
zonites decidedly smaller than the metazonites producing constric- 
tions between the metazoniteS.....ccceseee SIPHONOPHORIDAE 

One or more ocelli each side; head not strongly narrowed forward or 
produced into a beak; prozonites not forming constrictions between 


the*metazonittes! ses a swe « tele G.cielecccone: sheret ool ieieione POLY ZONIIDAE 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 109 


Family POLYZONIIDAE 
Genus SIPHONOTUS Brandt 1837, p. 179. 


TYPE SPECIES: Siphonotus brasiliensis Brandt, by monotypy. 
Siphonotus angulifer Chamberlin 1940a, p. 16. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

OTHER LOCALITIES: Summit (Chamberlin 1947c); Barro Colorado 
Island, Las Cascadas, Monte Lirio, Pifia area, Canal Zone (Loomis 
1961). 

Siphonotus centralis Chamberlin 1940a, p. 16. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 
Siphonotus panamanus Loomis 1964a, p. 122. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Chilibrillo Cave, near Chilibre, Panama Prov., 
Panama. 

PARATYPE LOCALITIES: Same, and Madden Forest Preserve, Canal 


Zone. 


Family SIPHONOPHORIDAE 
Genus SIPHONACME Cook and Loomis 1928, p. 7. 
TYPE SPECIES: Siphonacme lyttoni Cook and Loomis, by original 


designation. 
Siphonacme pseustes (Chamberlin). 
Siphonophora pseustes Chamberlin 1923a, p. 400; Cook and Loomis 
1928, p. 10. 
TYPE: Mus. California Acad. Sci., no. 1258. 
TYPE LOCALITY: San Pedro Bay, Sonora, Mexico. 


Genus SIPHONOCYBE Pocock 1903, p. 50. 


TYPE SPECIES: Siphonophora harti Pocock, by original designation. 
Siphonocybe laticollis Loomis 1%4a, p. 122. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Barca area, Finca Lerida, near Boquete, Chiriqut 
Prov., Panama. 

PARATYPE LOCALITIES: Finca Lerida, and Cerro Punta, Chiriqui 
Prov., Panama. 
Siphonocybe pilosa Loomis 1961, p. 118. 

TYPE: U.S. Nat. Mus., no. 2652. 

TYPE LOCALITY: Ft. Sherman, Canal Zone. 

PARATYPE LOCALITIES: Same, and Piffa area, Canal Zone. 


110 U.S. NATIONAL MUSEUM BULLETIN 266 


Genus SIPHONOPHORA Brandt 1837, p. 179. 


TYPE SPECIES: Siphonophora portoricensis Brandt, by subsequent 
designation of Silvestri 1896b. 
Siphonophora aviceps Loomis 1961, p. 119. 

TYPE: U.S. Nat. Mus., no. 2653. 

TYPE LOCALITY: Barro Colorado Island, Canal Zone. 

PARATYPE LOCALITIES: Same and Pifia area, Canal Zone; Juan 
Dias, Panama Prov., Panama. 
Siphonophora barberi Chamberlin 1922, p. 7. 

TYPE: U.S. Nat. Mus., no. 815. 

TYPE AND PARATYPE LOCALITY: Jocalo, Guatemala. 
Siphonophora brevicornis Pocock 1903, p. 50. 

TYPE: British Mus. 

TYPE LOCALITY: Volcan de Agua, Guatemala. 

PARATYPE LOCALITIES: Same and doubtfully Omilteme, Guerrero, 
Mexico. 
Siphonophora coclensis Loomis 1964a, p. 124. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Trail to Las Minas, FE] Valle, Coclé Prov., Panama. 
Siphonophora conicornis Chamberlin 1952b, p. 34. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Izabal, Escobas, Guatemala. 
Siphonophora cornuta Pocock 1903, p. 50. 

TYPE: British Mus. 

TYPE LOCALITY: Volcan de Acetenango, Guatemala. 

PARATYPE LOCALITIES: Same and Volcan de Agua, Guatemala. 
Siphonophora costaricae Chamberlin 1914, p. 186. 

TYPE: Acad. Nat. Sci. Philadelphia. 

TYPE LOCALITY: Juan Vinas, Costa Rica. 
Siphonophora fallens Chamberlin 1922, p. 8. 

TYPE: U.S. Nat. Mus., no. 817. 

TYPE AND PARATYPE LOCALITY: Joyabaj-Chiche, Guatemala. 

OTHER LOCALITY: Chamberlin 1940a reported specimens from Barro 
Colorado Island, Canal Zone, as this species but it is more likely they 
were S. aviceps Loomis 1961, p. 121. 
Siphonophora fallens annecta Chamberlin 1952b, p. 34. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Volcan Tajumulco, Guatemala. 
Siphonophora globiceps Pocock 1903, p. 51. 

TYPE: British Mus. 

TYPE LOCALITY: Purulha, Guatemala. 

OTHER LOCALITIES: Santa Rosa (Chamberlin 1922); Chichivac 
(Chamberlin 1952b), Guatemala. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 111 


Siphonophora mexicana Humbert and Saussure 1869, p. 155. 

TYPE: Mus. Hist. Nat. Geneve (H). 

TYPE LOCALITY: Eastern Cordillera, Mexico. 

PARATYPE LOCALITY: Sierra de Agua, near Orizaba, Mexico. 
Siphonophora montana Loomis 1964a, p. 125. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Cerro Punta, Chiriqui Prov., Panama. 

PARATYPE LOCALITY: Finca Lerida, near Boquete, Chiriquf Prov., 
Panama. 

Siphonophora panamensis Loomis 1961, p. 121. 

TYPE: U.S. Nat. Mus., no. 2654. 

TYPE LOCALITY: Pifia area, Canal Zone. 

PARATYPE LOCALITIES: Barro Colorado Island, Frijoles, Monte 
Lirio, Ft. Sherman, Canal Zone. Alahjuela and Juan Dias, Panama Prov.; 
El Valle, Coclé Prov., Panama. 

Siphonophora parvula Loomis 1964a, p. 125. 

TYPE: Chicago Nat. Hist. Mus. 

TYPE LOCALITY: Barca area, Finca Lerida, near Boquete, Chiriqut 
Prov., Panama. 

PARATYPE LOCALITIES: Same and adjacent areas; Finca Palo 
Santo, near Nueva California, Chiriqui Prov., Panama. 

Siphonophora progressor Chamberlin 1922, p. 9. 

TYPE: U.S. Nat. Mus., no. 818. 

TYPE AND PARATYPE LOCALITY: Progreso, Honduras. 
Siphonophora sabachana Chamberlin 1938a, p. 166. 

TYPE: Collection of R. V. Chamberlin. 

TYPE LOCALITY: Sabacha Cave, Tekax, Yucatan, Mexico. 
Siphonophora telana Chamberlin 1922, p. 7. 

TYPE: U.S. Nat. Mus., no. 816. 

TYPE LOCALITY: Tela, Honduras. 

Siphonophora trifini Kraus 1954b, p. 343. 

TYPE: Senckenberg Mus., Frankfurt. 

TYPE AND PARATYPE LOCALITY: Monte Cristo, Metapan Mts., 
Dept. Santa Ana, EF] Salvador. 

Siphonophora valeri Chamberlin 1933, p. 24. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Pavas, Costa Rica. 
Siphonophora vera Chamberlin 1952b, p. 34. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Las Vigas, Veracruz, Mexico. 


Genus BONETESMUS Chamberlin 1942b, p. 11 (see page 28) 


TYPE SPECIES: Bonetesmus verus Chamberlin, by original designation. 
Bonetesmus verus Chamberlin 1942b, p. 11. 

TYPE: Collection of R. V. Chamberlin. 

TYPE AND PARATYPE LOCALITY: Gruta de Atoyac, Vera Cruz, 


Mexico. 


112 U.S. NATIONAL MUSEUM BULLETIN 266 


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U.S. NATIONAL MUSEUM BULLETIN 266 


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Soc. Washington, vol. 68, pp. 87-94, illustr. 

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CHAMBERLIN, R. V. 


1914. 


1918a. 


1918b. 


1922. 


1923a. 


1923b. 


1923c. 


1924. 


1925. 


1931. 


1933. 


1938a. 


1938b. 


1940a. 


On a collection of Myriapoda from Costa Rica. Trans. Amer. Ent. 
Soc., vol. 40, pp. 185-194, illustr. 

The Chilopoda and Diplopoda of the West Indies. Bull. Mus. Comp. 
Zool., vol. 62, no. 5, pp. 151-262. 

New spirobolid diplopods. Proc. Biol. Soc. Washington, vol. 31, 
pp- 165-170. 

The millipeds of Central America. Proc. U.S. Nat. Mus., vol. 60, 
no. 8, pp. 1-75, illust. 

On chilopods and diplopods from islands in the Gulf of Califomia. 
Proc. California Acad. Sci., vol. 12, no. 18, pp. 389-407, illustr. 

On four termitophilous millipeds from British Guiana. Zool., New 
York Zool. Soc., vol. 3, no. 21, pp. 411-421, illustr. 

A new cryptodesmoid milliped from Santo Domingo. Proc. Biol. 
Soc. Washington, vol. 36, pp. 189-190. 

A new leptodesmoid milliped from Nicaragua. Ent. News., vol. 35, 
p- 174, illustr. 

Notes on chilopods and diplopods from Barro Colorado Id., and 
other parts of the Canal Zone, with diagnoses of new species. 
Proc. Biol. Soc. Washington, vol. 38, pp. 35-44. 

On a collection of Chilopoda and Diplopoda from Oklahoma. Ent. 
News, vol. 42, no. 4, pp. 97-104, illustr. 

On a collection of centipeds and millipeds from Costa Rica. Pan- 
Pacific Ent., vol. 9, no. 1, pp. 11-24, illustr. 

Diplopoda from Yucatan. Carnegie Inst. Washington, no. 491, 
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New diplopods. Proc. Biol. Soc. Washington, vol. 51, pp. 205-208. 

On a diplopod collection from Barro Colorado Island, Panama. Bull. 
Univ. Utah, vol. 30, no. 9, pp. 3-16. 


1940b. 


1941a. 


1941b. 


194]1c. 


1941d. 


194le. 


1942a. 


1942b. 


1942c. 


1942d. 


1943a. 


1943b. 


1943c. 


1944. 


1946. 


1947a. 


1947b. 


1947c. 


1947d. 


1949, 


1950. 


1951. 


1952a. 


1952b. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 115 


New genera and species of North American Paraiulidae. Bull. 
Univ. Utah, vol. 30, no. 11, pp. 3-39, illustr. 

A new oniscodesmoid diplopod from Barro Colorado Island. Pomona 
Coll. Journ. Ent. Zool., vol. 33, pp. 57-58, illustr. 

New American millipeds. Bull. Univ. Utah, vol. 31, no. 11, pp. 3-39, 
illustr. 

On five new polydesmid millipeds from Mexico. Proc. Biol. Soc. 
Washington, vol. 54, pp. 63-66, illustr. 

Seven new millipeds from Mexico (Chilopoda). Ent. News, vol. 52, 
pp- 250-57, illustr. 

On a collection of millipedes and centipedes from northeastern 
Peru. Bull. Amer. Mus. Nat. Hist., vol. 78, no. 7, pp. 473-535, 
illustr. 

New millipeds from Michoacan. Proc. Biol. Soc. Washington, vol. 
55, pp- 57-62, illustr. 

On centipeds and millipeds from Mexican caves. Bull. Univ. Utah, 
vol. 33, no. 4, pp. 3-19, illustr. 

Two millipeds of the genus Rhysodesmus from Mexico, Canadian 
Ent., vol. 74, no. 5, pp. 81-92, illustr. 

On a collection of myriopods from Iowa. Canadian Ent., vol. 74, 
pp- 15-17, illustr. 

On some genera and species of American millipeds. Bull. Univ. 
Utah, vol. 34, no. 6, pp. 3-20, illustr. 

On Mexican millipeds. Bull. Univ. Utah, vol. 34, no. 7, pp. 3-103, 
illustr. 

Some records and descriptions of American diplopods. Proc. Biol. 
Soc. Washington, vol. 56, pp. 143-152, illustr. 

Two millipeds from Southern California. Proc. Biol. Soc., Washing- 
ton, vol. 57, pp. 113-116, illustr. 

Anewcentiped and two newmillipeds from the Pearl Islands, Colom- 
bia. Pan-Pacific Ent., vol. 22, no. 4, pp. 145-147, illustr. 

Seven new American millipeds. Proc. Biol. Soc. Washington, vol. 
60, pp. 9-16, illustr. 

Some records and descriptions of diplopods chiefly in the collec- 
tion of the Academy. Proc. Acad. Nat. Sci. Philadelphia, vol. 99, 
pp- 21-58, illustr. 

Two new species of Trichomorpha from Panama. Proc. Biol. Soc. 
Washington, vol. 60, pp. 63-66, illustr. 

Anew myrmecophilous milliped from Mexico. Pan-Pacific Ent., vol. 
23, no. 3, pp. 101-102, illustr. 

On some Western millipeds of the order Spirobolida. Journ. Wash- 
ington Acad. Sci., vol. 39, no. 5, pp. 163-169, illustr. 

Three new genera and eight new species of Western millipeds. Nat. 
Hist. Misc., Chicago Acad. Sci., no. 68, pp. 1-6, illustr. 

On Diplopoda of north-east Angola. Public. Cult. Comp. Diam. 
Angola (separ. 10), pp. 63-94, illustr. 

Some American polydesmid millipeds in the collection of the 
Chicago Museum of Natural History. Ann. Ent. Soc. America, 
vol. 45, no. 4, pp. 553-584, illustr. 

Further records and descriptions of American millipeds. Great 
Basin Nat., vol. 12, nos. 1-4, pp. 13-34, illustr. 


116 U.S. NATIONAL MUSEUM BULLETIN 266 


1952c. Three cave-dwelling millipeds. Ent. News, vol. 63, pp. 10-12. 

1952d. Eclomus nom. nov. Diplopoda. Ent. News, Vol. 63, p. 71. 

1953a. Some American millipeds of the order Spirobolida. Amer. Midland 
Nat., vol. 50, no. 1, pp. 138-151, illustr. 

1953b. Six new American millipeds, with notes on several cave-dwelling 
species. Proc. Biol. Soc. Washington, vol. 66, pp. 67-71 illustr. 


1956. Three newNicaraguan millipeds. Ent. News, vol. 67, no. 6, pp. 157- 
159, illustr. 


CHAMBERLIN, R. V., and HOFFMAN, R. L. 
1950. On some genera and families of North American diplopods. Chicago 
Acad. Sci. Nat. Hist. Misc., no. 71, pp. 1-7. 


1958. Checklist of the millipeds of North America. U.S. Nat. Mus. Bull. 
212, pp. 1-236. 


CHAMBERLIN, R. V., and MULAIK, S. 


1941. On a collection of millipeds from Texas and New Mexico. Journ. 
New York Ent. Soc., vol. 49, no. 1, pp. 57-64. 


CONDE, B., and TERVER, D. 
1964. Penicillates du Surinam et du Guatemala (Diplopoda, Lophoprocti- 
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COOK, O. F. 


1895. Introductory note on the families of Diplopoda, in Cook and Collins, 
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1896. Brandtia, A series of occasional papers. Pp. 1-75. 

1898. American oniscoid Diplopoda of the order Merocheta. Proc. U.S. 
Nat. Mus., vol. 21, pp. 451-468, illustr. 

1901. Duoporus,a new diplopod from Mexico. Proc. Ent. Soc. Washington, 
vol. 4, no. 4, pp. 402404. 

1904. Myriapoda of northwestern North America, in Harriman, Alaska Ex- 
pedition, vol. 8 (Insects, pt. 1), pp. 47-82, illustr. 

191la. Notes onthe distribution of millipeds in southern Texas, with de- 
scriptions of new genera and species from Texas, Arizona, 
Mexico and Costa Rica. Proc. U.S. Nat. Mus., vol. 40, no. 1810, 
pp- 147-167. 

1911b. New tropical millipeds of the order Merocheta, with an example of 
kinetic evolution. Proc. U.S. Nat. Mus., vol. 40, no. 1831, pp. 451- 
473, illustr. 

19llc. The hothouse milliped as a new genus. Proc. U.S. Nat.Mus,, vol. 
41, no. 1842, pp. 625-631. 


COOK, O. F., and COLLINS, G. N. 


1895, The Craspedosomatidae of North America. Ann. New York Acad. 
Sci., vol. 9, pp. 1-100, illustr. 


COOK, O. F., and COOK, A. C. 


1894. A monograph of Scytonotus. Ann. New York Acad. Sci., vol. 8, 
pp- 233-248, illustr. 





MILLIPEDS OF MEXICO AND CENTRAL AMERICA 117 


COOK, O. F., and LOOMIS, H. F. 
1928. Millipeds of the order Colobognatha, with descriptions of six new 
genera and type species, from Arizona and California. Proc. U.S. 
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DADAY, E. 


1889. Myriapods extranea mussei nationalis Hungarici. Term. Fuzetek, 
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GERVAIS, P. 
1836. Nouvelles especes de myriapodes. L’ Institut, sect. 1, vol. 4, no. 190, 
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1844. Etudes sur les myriapodes. Ann. Sci. Nat. Zool., vol. 2, ser. 3, 
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1847. Myriapodes, in Walckenaer and Gervais, Histoire naturelle des in- 
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GERVAIS P., and GOUDOT, J. 


1844. Description des Myriapodes recueillis par J. Goudot en Colombie. 
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GILL, DR. (See Kenyon, F. C.) 


GIRARD, C. 


1853. Myriapods, in Marcy, Report on exploration of the Red River of 
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GOLDFUSS, GEORGE AUGUST 


1820. Handbuch der Zoologie, theil 3, in Schubert, Handbuch der 
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GRAY, J. E. 


1832. Myriapods, in Griffith, The animal kingdom arranged in conformity 
with its organization by the Baron Cuvier, vol. 15 (Class In- 
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HOFFMAN, R. L. 


1950a. A preliminary list of the cleidogonid millipeds, with descriptions 
of a new genus from Guatemala and a new species from Virginia. 
Journ. Washington Acad. Sci., vol. 41, no. 3, pp. 87-92, illustr. 

1950b. Systematic notes on some Central American millipeds. Proc. Biol. 
Soc. Washington, vol. 63, pp. 69-72. 

1951, A new genus of Central American milliped (Family Euryuridae) 
with notes on American genera. Proc. U.S. Nat. Mus., vol. 102, 
no. 3300, pp. 235-243, illustr. 

1953a. Studies on spirostreptid millipeds. I. The genus /soporostreptus 
Silvestri. Lloydia, vol. 16, pp. 149-156, illustr. 

1953b. Psammodesmus, a _ neglected milliped genus (Polydesmida- 
Platyrhacidae). Journ. Washington Acad. Sci., vol. 43, no. 9, 
pp- 299-304, illustr. 


1953c. A new Central American milliped of the genus Platyrhacus. Proc. 
Ent. Soc. Washington, vol. 55, pp. 251-258, illustr. 

1953d. Studies on spirobolid millipeds. I. The genus Eurhinocricus Brole- 
mann. Proc. Biol. Soc. Washington, vol. 66, pp. 179-183. 

1954a. Further studies on American millipeds of the family Euryuridae 
(Polydesmida). Journ. Washington Acad. Sci., vol. 44, no. 2, 
pp. 49-58, illustr. 


118 U.S. NATIONAL MUSEUM BULLETIN 266 


1954b. A new milliped of the genus Colactis from Mexico (Chordeumida- 
Lysiopetalidae). Amer. Mus. Nat. Hist., no. 1673, pp. 144, illustr. 

1955. Studies on Spirobolid millipeds. II]. A second paper on the genus 
Eurhinocricus. Proc. Biol. Soc. Washington, vol. 68, pp. 31-36. 

1959, A newdiplopod genus from Guatemala,closely related to Desmonus, 
and its effect on the validity of the families Desmonidae and 
Cyclodesmidae (Polydesmida, Sphaeriodesmidae). Proc. Biol. 
Soc. Washington, vol. 72, pp. 173-182, illustr. 

1960. A fourth contribution to the knowledge of neotropical platyrhacid 
millipeds (Diplopoda; polydesmida). Proc. U.S. Nat. Mus., vol. 
111, no. 3423, pp. 17-31, illustr. 

1962. A new species of the milliped genus Polylepiscus from Mexico, 
with some remarks on the status of the genus (Polydesmida- 
Euryuridae). Proc. Ent. Soc. Washington, vol. 64, no. 2, pp. 135- 
140, illustr. 

1964. <A third smooth species of Polylepiscus (Polydesmida-Euryuridae). 
Proc. Biol. Soc. Washington, vol. 77, pp. 171-174. 


HOFFMAN, R. L., and CRABILL, R. E. 
1953. C.S. Rafinesque as the real father of American Myriapodology: an 
analysis of his hitherto unrecognized species. Florida Ent., vol. 
36, no. 2, pp. 73-82. 


HOFFMAN, R. L., and ORCUTT, B. S. 
1960. A synopsis of the Atopetholidae, a family of spirobolid millipeds. 
Proc. U.S. Nat. Mus., vol. 111, no. 3426, pp. 95-166, illustr. 


HUMBERT, A., and SAUSSURE, H. 
1869. Myriapoda nova Americana. Rev. Mag. Zool., ser. 2, vol. 21, pp. 149= 
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1870. Myriapoda nova Americana: Description du divers Myriapodes nou- 
veaux du musee de Vienne. Rev. Mag. Zool., vol. 22, 
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JAWLOWSKI, H. 
1930. On European Diplopoda introduced into America. Fragmenta Mus. 
Zool. Polon, T. 1, no. 7, pp. 181-185, illustr. 


JEEKEL, C. A. W. 
1963a. Diplopoda of Guiana (1-5). Nat. Stud. Suriname en de Nederlandse 
Antillen, vol. 4, no. 27, pp. 1-157, illustr. 


1963b. Alloproctinus nom. nov. (Diplopoda, Pollyxenida), Ent. Bericht., 
vol. 23, p. 156. 


KARSCH, F. 
1881. Neue Juliden des Berlinen-Museums, als prodromus einer juliden 
Monographie. Zeitschr. Naturw. (ser. 3, vol. 6), vol. 54, pp. 1-79. 


KEETON, W. T. 
1960a. A taxonomic study of the milliped family Spirobolidae (Diplopoda; 
Spirobolida). Mem. Amer. Ent. Soc., Philadelphia, no. 17, pp. 1-146. 
illustr. 
1960b. A new family of millipeds of the order Spirobolida, with notes on 
an established family. Proc. Biol. Soc. Washington, vol. 73, pp. 
131-140, illustr. 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 119 


KENYON, F. C. 
1899, Anew Mexican diplopod, Decaporodesmus motzoranginis, type of a 
new family Decaporodesmidae. Proc. Ent. Soc. Washington, vol. 
4, pp. 299-300. 


KOCH, C. L. 
1847. System der myriapoden, in Herrich-Schaffer, Kritische Revision der 


insectfaune Deutschlands, vol. 3, pp. 1496, illustr. 


KRAUS, O. 
1954a. Myriapoden aus Peru, II. Senck. Biol., vol. 35 (1/2), pp. 17-55, 
illustr. 
1954b. Myriapoden aus El] Salvador. Senck. Biol., vol. 35 (5/6), pp. 293- 
349, illustr. 


LATREILLE, P. A. 
1802. Histoire Naturelle...de crustaces et des Insects. vol. 3, pp. 1-467. 


LATZEL, ROBERT 
1884. Die myriapoden der Osterreichisch-Ungarischen Monarchie..., 
vol. 2, Die Symphylen, Pauropoden und Diplopoden. pp. 1-414, 
illustr. 


LOOMIS, H. F. 

1934. Millipeds of the West Indies and Guiana collected by the Allison 
V. Armour Expedition in 1932. Smithsonian Misc. Coll., vol. 89, 
no. 3244, pp. 1-69, illustr. 

1936. Three new millipeds of the order Colobognatha from Tennessee, 
Texas and Lower California, with records of previously known 
species. Proc. U.S. Nat. Mus., vol. 83, no. 2989, pp. 361-368, 
illustr. 

1937. Crested millipeds of the family Lysiopetalidae in North America, 
with descriptions of new genera and species. Proc, U.S. Nat. 
Mus., vol. 84, no. 3006, pp. 97-135, illustr. 

1938. The cambaloid millipeds of the United States, including a family 
new to the fauna and new genera and species, Proc. U.S. Nat. 
Mus., vol. 86, no. 3043, pp. 27-66, illustr. 


1944. Millipeds principally collected by Professor F. E. Shelford in the 
eastern and southeastern states. Psyche, vol. 51, pp. 166-177, 
illustr. 

1949. New millipeds of the spirobolid genus Watichelus from the Pacific 
Coast. Journ. Washington Acad. Sci., vol. 39, no. 7, pp. 241- 
244, illustr. 

1950. Synonymy of some native American andintroducedmillipeds. Journ. 
Washington Acad. Sci., vol. 40, no. 5, pp. 164-166, illustr. 

1953. New millipeds of the western states and Lower California. Journ. 
Washington Acad. Sci., vol. 43, no. 12, pp. 417-422, illustr. 

1958. A newmilliped family on Barro Colorado Island, Canal Zone. Journ. 
Washington Acad. Sci., vol. 48, no. 1, pp. 235-237, illustr. 

1959a. New myrmecophilous millipeds from Barro Colorado Island, Canal 
Zone, and Mexico. Journ. Kansas Ent. Soc., vol. 32, pp. 1-7, 
illustr. 

1959b. A new milliped from Honduras. Ceiba, Escuela Agric. Panameri- 
cana, vol. 8, no. 2, pp. 1-3, illustr. 


120 U.S. NATIONAL MUSEUM BULLETIN 266 


1960. Millipeds of the order Polydesmida from the western states and 
Baja California. Journ. Kansas Ent. Soc., vol. 33, pp. 57-68, 
illustr. 

1961. New and previously known millipeds of Panama. Proc. U.S. Nat. 
Mus., vol. 113, no. 3454, pp. 77-123, illustr. 

1962. Twounusual Central American spirostreptid milliped species. Proc. 
Biol. Soc. Washington, vol. 75, pp. 47-51, illustr. 

1963. Millipeds from states immediately north and south of the Mexican 
boundary. Journ. Kansas Ent. Soc., vol. 36, no. 2, pp. 118-126, 
illustr. 

1964a. The millipeds of Panama (Diplopoda). Fieldiana: Zoology, Chicago 
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1964b. Three new polydesmoid millipeds from Central America. Proc. Biol. 
Soc. Washington, vol. 77, pp. 183-188, illustr. 


LOOMIS, H. F., and HOFFMAN, R. L. 

1948. Synonymy of various diplopods. Proc. Biol. Soc. Washington, vol. 
61, pp. 51-54. 

1953. Psammodemus schmitti Loomis and Hoffman, in Hoffman, P sammodes- 
mus, aneglected milliped genus (Polydesmida: Platyrhacidae), 
Journ. Washington Acad. Sci., vol. 43, no. 9, pp. 299-304, illustr. 

1962. A remarkable new family of spined polydesmoid Diplopoda, includ- 
ing a species lacking gonopods in the male sex. Proc. Biol. Soc. 
Washington, vol. 75, pp. 145-158, illustr. 


LUCAS, H. 
1840. Histoirenaturelle des crustacés, des arachnides, etdes myriapodes. 
Paris, pp. 511-551. 
1843. Observations sur un nouveau genre de la classe des myriapodes, 
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MENGE, F. A. 
1851. Myriapoden der Umgegend von Danzig. Neu. Schrift. Naturf. Ges. 
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NEWPORT, G. 
1844. A list of the species of myriapods in the collection of the British 
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NICHOLAS, Bro. G. 
1962. Checklist of troglobitic organisms of Middle America. Amer. Midl. 
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PACKARD, A. S. 
1883. A revision of the Lysiopetalidae, a family of chilognath Myriapoda, 
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PETERS, W. C. H. 

1864. Ubersicht der im Konigl, zoologischen museum befindlich Myria- 
poden aus der familie der polydesmi, so wie beschreibungen 
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MILLIPEDS OF MEXICO AND CENTRAL AMERICA 121 


POCOCK, R. I. 

1893. Upon the identity of some of the types of Diplopoda contained in 
the collection of the British Museum, together with descriptions 
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1894. Chilopoda, Symphyla, and Diplopoda from the Malay Archipelago, 
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1903- Chilopoda and Diplopoda, Biologia Centrali-Americana, pp. 41-217, 

1910. illustr. 


PORAT, C. O. 
1889a. Uber einige exotischen Juliden des Briisseler-Museums. Ann. Soc. 
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1889b. Om nagra exotiska myriapoder. Bihang K. Svenska Vet. Akad. Handl., 
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RAFINESQUE, C.S. 
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SAUSSURE, H. 

1859. Note sur la famille des polydesmides principalement au point de 
vue des especes Americaines. Diagnoses de divers myriapods 
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1860. Essai d’une faune des myriapodes du Mexique, avec la description 
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SAUSSURE, H., and HUMBERT, A. 
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SAUSSURE, H., and ZEHNTER, L. 
1902. Myriapodes de Madagascar, in Grandidier, Histoire physique, 
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SAY, T. 
1821. Description of the Myriapodae of the United States. Journ. Acad. 
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SCHUBART, O. 

1946. Contribuicao ao conhecimento do genero Leptodesmus (Familia 
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1952. Um Diplopoda (Calymmodesmus sodalis n. sp.), companheiro de 
formigas em migracao (Proterospermophora, Stylodesmidae). Dos 
Anais da Acad. Brasileira de Ciencias, vol. 24,no. 4, pp. 437- 
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1955. Uma Idiurodesminae de Goias: (Diplopoda, Proterospermophora, 
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122 


SILVESTRI, F. 
1896a. Chilopodi e Diplopodi reccolti dal Dott. E. Festa a La Guayra, 
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1897a. Descrizione di una famiglia di Diplopodi del Messico. Boll. Mus. 


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1897c. Viaggio del Dr. Enrico Festa nell’ Ecuador e regioni vicine. Boll. 
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Contributo alla conoscenze dei mirmecofili de Mexico. Boll. Lab. 
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Descrizione di un nuovo genere di Polydesmidae (Diplopoda) 
termitofilo di Costa Rica. Boll Lab. Zool. Portici, vol. 17, pp. 
172-175, illustr. 


meridionale. Boll. Lab. Ent. Agric., Portici, vol. 8, pp. 1-32, 
illustr. 


1948b. Tavola sinottica dei generi dei Diplopoda Penicillata. Boll. Lab. 


1949. 


Ent. Agric., Portici, vol. 8, pp. 214-220. 
Descrizioni di nuovi diplopodi Penicellati. Boll. Lab. Ent. Agric., 
Portici.,vol. 9, pp. 1-7, illustr. 


VERHOEFF, K. W. 


1926. 


1932. 


1937. 


1941. 


1944. 


Chilognathen-Beitrage. Zool. Anz., Leipzig, vol. 68, pp. 109-127, 
illustr. 

Diplopoden-Beitrage. Zool. Jahrb. Jena (Syst.), vol. 62, Heft. 5-6, 
pp. 469-524, illustr. 

Zur kenntnis der Rhinocricidae. Zool. Anz., Leipzig, vol. 118, 
pp- 90-102, illustr. 

Versuch eines  Siphonophoriden-systems und _ geographische- 
phylogenetische beurteilung der gonopoden. Zool. Anz., Leipzig, 
vol. 134, pp. 212-224, illustr. 

Some Californian Chilognatha. Bull. Southern California Acad. Sci., 
vol. 43, pt. 2, pp. 53-70, illustr. 














MILLIPEDS OF MEXICO AND CENTRAL AMERICA 123 


VOGES, E. 
1878. Beitrage zur kenntnis der iuliden. Zeitscher. Wissen Zool., vol. 31, 
pp. 127-194. 


WOOD; H.C. 
1864. Descriptions of new genera and species of North American Myria- 
poda. Proc. Acad. Nat. Sci. Philadelphia, vol. 16, pp. 186-187. 
1865. The Myriapoda of North America. Trans. Amer. Philos. Soc., n. Se, 
vol. 13, (1869), pt. 2, art. 7, pp. 137-248, illustr. 


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Abacion, 70 
aberrans, Tiphallus, 42 
absconsus, Orthoporus, 97 
Scaphiostreptus, 97 
Acakandra, 66 
acanthethus, Orthoporus, 97 
acanthurus, Chondrodesmus, 12 
Polydesmus, 12 
Acentronus, 58 
Aceratophallus, 35 
acolhuus, Polydesmus, 58 
Rhysodesmus, 58 
actaeon, Polylepiscus, 22 
acuarius, Rhysodesmus, 58 
Acutangulus, 37 
acuticollis, Chondrodesmus, 12 
adunca, Rhachidomorpha, 39 
aduncus, Polydesmus, 39 
Aenigmopus, 57 
affinis, Epinannolene, 96 
Airocaulon, 46 
alata propinqua, Salvadoria, 56 
alata, Salvadoria, 56 
alatus, Aenigmopus, 57 
alboalatus, Nyssodesmus, 32 
albomarginis, Nyssodesmus, 35 
alidens, Chondrodesmus, 12 
alienus, Yucodesmus, 53 
Allarithmus, 15 
allenae, Chondrodesmus, 12 
Allopocockia, 77 
Allopocockiidae, 77 
Alloproctinus, 5 
Alloproctus, 5: 
Alocodesmus, 11 
Alpuyecus, Rhysodesmus, 58 
amblus, Pararhachistes, 39 
Amplinus, 21 
amplinus, Rhysodesmus, 58 
ampussis, Orthoporus, 98 
Spirostreptus, 98 
amulensis, Messicobolus, 83 
Orthoporus, 100 
Paraiulus, 74 
Spirobolus, 83 
analis, Platydesmus, 106 
andinus, Calymmodesmus, 47 
Andrognathidae, 105 
Anelus, 81, 82 
angelus, Polydesmus, 58 
Rhysodesmus, 58 
angulatus, Oncodesmoides, 19 
angulifer, Siphonotus, 109 
angustatus, Alcodesmus, 11] 


INDEX 


angusticollis, Rhinocricus, 87 
Spirobolus, 87 

angustus, Eusphaeriodesmus, 43 
Polydesmus, 35 
Sphaeriodesmus, 43 

anisorhabdus, Alloproctinus, 5 

annulatus, Julus, 95 
Prostemmiulus, 71 

antius, Nyssodesmus, 32 
Platyrhacus, 32 

Aphelidesmus, 21 

apiculatus, Pachygonopus, 56 

Apomus, 29 

Aporiaria, 57 

aposematus, Rhinocricus, 87 

approximatus, Rhinocricus, 87 

Apsyma, 46 

archimedes, Cyrtodesmus, 19 
Peltedo, 19 

arcuatus, Rhysodesmus, 58 

areatus, Amplinus, 23 
Orthomorpha, 23 
Pseudamplinus, 23 

argentea, Salvadoria, 56 

arizonicus, Orthoporus, 102 

armatus, Amplinus, 23 
Chondrodesmus, 12 
Nopoiulus, 74 
Orthomorpha, 23 
Pseudamplinus, 23 

Arolus, 77 

asper, Orthoporus, 98 
Scaphiostreptus, 98 

atopa, Apsyma, 46 

Atopetholidae, 77, 79 

Atopetholus, 79 

atopophallus, Schistides, 15 

atoyaca, Cleidogona, 67 

atoyacus, Rhinocricus, 87 

atriculus, Allopocockia, 78 
Chelogonobolus, 78 
Spirobolellus, 78 

atrophus, Chondrodesmus, 12 

attemsi, Nyssodesmus, 32 
Rhysodesmus, 58 

Atylophor, 11 

Augesmus, 46 

aurantium, Rhachidomorpha, 40 

aurocinctus, Rhinocricus, 87 

austrina, Acakandra, 66 

Auxobolus, 93 

aviceps, Siphonophora, 110 

Aymaresmus, 33 

Aztecolus, 92 


125 


126 U.S. NATIONAL MUSEUM BULLETIN 266 


aztecus, Cyclodesmus, 42 
Julus, 87 
Leptodesmus, 37 
Neoleptodesmus, 37 
Paraiulus, 75 
Polydesmus, 37 
Rhinocricus, 87 


balboanus, Hiltonius, 93 
barberi, Siphonophora, 110 
barretti, Duoporus, 27 
barricolens, Glomeridesmus, 7 
barrios, Eurhinocricus, 85 
Barrobolus, 91 
Barrodesmus, 28 
barroensis, Diaporus, 99 
Dorsoporus, 20 
Barroxenus, 6 
Barydesmus, 33 
benedictus, Oxypyge, 86 
bicolor, Cyrtodesmus, 19 
Glomeridesmus, 7 
bicornis, Epinannolene, 96 
bidentatus, Leiogonopus, 55 
biensifer, Calymmodesmus, 47 
bifidus, Peridysodesmus, 44 
bifurcus, Cyrtodesmus, 19 
bilineatus, Platyrhacus, 33 
Polydesmus, 33 
bioculatus, Iulus, 72 
Stemmiulus, 72 
biolleyi, Colobodesmus, 42 
Eurhinocricus, 85 
Platyrhacus, 35 
Rhinocricus, 85 
Tirodesmus, 35 
biramosus, Botrydesmus, 47 
bisulcatus, Orthoporus, 98 
bivirgatus, Nyssodesmus, 32 
Platyrrhacus, 32 
Blaniulus, 74 
bobos, Orthoporus, 98 
boleti, Julus, 73 
Bolivaresmus, 46 
bolivari, Rhysodesmus, 58 
Tarascolus, 81 
Bonetesmus, 28, 111 
boneti, Glomeris, 7 
bonus, Rhysodesmus, 58 
Botrydesmus, 46 
brachus, Rhysodesmus, 58 
Brachycybe, 105 
Brachyiulus, 73 
brasiliensis, Siphonotus, 109 
brevicollis, Rhinocricus, 87 
Spirobolus, 87 
brevicornis, Siphonophora, 110 
browni, Cruzodesmus, 57 
bukowinus, Sphaeriodesmus, 44 
burgeri, Polylepiscus, 22 


caeca, Cambala, 95 
Epinannolene, 96 

caecus, Glomeroides, 7 

caeruleus, Zeuctodesmus, 28 

caesioannulatus, Spirostrephon, 66 

calcehtokanus, Aceratophallus, 35 

Californibolus, 93 

californicus, Atopetholus, 79 
Spirobolus, 94 

callipus, Hiltonius, 92 
Spirobolus, 92 

calus, Platydesmus, 106 

calverti, Aphelidesmus, 22 

Calymmodesmus, 47 

Cambala, 94 

Cambalidae, 94 

Cambalidea, 94 

campanulae, Polylepiscus, 22 

Campodes, 66 

canalis, Orthoporus, 98 
Stemmiulus, 72 

capillata, Trichomorpha, 16 

captiosa, Cambala, 95 

capucinus, Orthoporus, 98 
Scaphiostreptus, 98 

carinaceps, Xenoporus, 52 

carpinus, Hiltonius, 92 

carpinus vulcan, Hiltonius, 93 

Catapleuradesmus, 48 

Cavota, 66 

ceibana, Cleidogona, 68 
Hirsutogona, 68 

celatus, Cynedesmus, 50 
Lophodesmus, 50 

centralis, Glomeridesmus, 8 
Glomeroides, 7 
Rhinocricus, 87 
Siphonotus, 109 

Centrellus, 79 

Ceratesmus, 48 

cerroazulensis, Amplinus, 21 
Seminellogon, 21 

cerrobius, Platydesmus, 106 

cervantes, Ethophallus, 14 

Ceuthauxus, 40 

chamberlini, Chondrodesmus, 13 

championi, Rhysodesmus, 59 

chapini, Platyrhacus, 34 

chapultepecus, Polydesmus, 35 

Chelodesmidae, 11 

Chelogonobolus, 77, 78 

chiapasus, Orthoporus, 103 

chichen, Rhinocricus, 87 

chichimecus, Julus, 88 
Rhinocricus, 88 
Spirobolus, 88 

chichivacus, Eurhinocricus, 85 
Schmidtolus, 78 

chihuanus, Comanchelus, 79 
Orthoporus, 98 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 127 


Toltecolus, 79 
Chilognatha, 5, 6 
chipinqueus, Styraxodesmus, 52 
chiriquensis, Desmethus, 106 

Diaporus, 98 

Irazunus, 54 

Orthoporus, 98 

Scaphiostreptus, 98 
chiriquia, Solemia, 69 
chitarianus, Amplinus, 21 

Rhinocricus, 88 

Seminellogon, 21 
Chondrodesmus, 12 
Chordeumida, 8, 65 
Chordeumidea, 65, 66 
Chytodesmidae, 10, 17 
cienegonus, Orthoporus, 98 
cincinnatus, Prostemmiulus, 71 
cinctus, Oxobolus, 84 
circularis, Glomeridesmus, 8 
clara, Orthyma, 50 
claremontus, Auxobolus, 94 

Tylobolus, 94 
clarus, Ceratesmus, 48 

Tarascolus, 81 
Cleidogona, 66 
Cleidogonidae, 66 
coarctata, Orthomorpha, 65 
coarctatus, Polydesmus, 65 
cobanus, Colobodesmus, 42 

Orthoporus, 98 

Sphaeriodesmus, 42 
coccineus, Acutangulus, 37 

Polydesmus, 37 
coclensis, Siphonophora, 110 
cocos, Rhinocricus, 85 
Colactis, 70 
collaris, Aztecolus, 82, 92 

Messicobolus, 82 
Colobodesmus, 42 
Colobognatha, 6, 105 
colotlipa, Ilyma, 49 
Comanchelus, 79 
comminutus, Orthoporus, 99 

Scaphiostreptus, 99 
Comodesmidae, 10, 17 
complanatus, Julus, 35 
concolor, Nyssodesmus, 32 
conflyentus, Cyrtodesmus, 19 

Harpogonopus, 54 
conformans, Sphaeriodesmus, 44 
confragosus, Orthoporus, 99 

Spirostreptus, 99 
confusa, Oxypyge, 86 
conicornis, Siphonophora, 110 
conifer, Botrydesmus, 47 

Orthoporus, 99 

Scaphiostreptus, 99 


consobrinus, Polydesmus, 59 
Rhysodesmus, 59 

constrictus, Amplinus, 23 
Cylionus, 42 
Pseudamplinus, 23 

contortus, Cyclorhabdus, 15 
Leptodesmus, 15 
Solaenorhabdus, 15 

convexus, Amplinus, 23 
Orthomorpha, 23 
Pachyurus, 23 
Pseudamplinus, 23 

cooki, Eurhinocricus, 85 
Prostemmiulus, 71 
Watichelus, 81 


copiosus, Cylindrogonus, 54 
cordobana, Hexodontia, 29 
cordovanus, Orthoporus, 99 
coriaceus, Sphaeriodesmus, 44 
cornuta, Siphonophora, 110 
coronatus, Botrydesmus, 47 
Cyrtodesmus, 19 
corzoi, Platydesmus, 106 
cos, Psammodesmus, 34 
costaricae, Siphonophora, 110 
costaricensis, Rhinocricus, 88 
crassus, Hiltonius, 92 
cratus, Oxobolus, 84 
crenus, Amplinus, 24 
Pseudamplinus, 24 
crotonus, Orthoporus, 102 
crucis, Cavota, 66 
Platydesmus, 106 
cruzanus, Ceuthauxus, 40 
Strongylodesmus, 40 
Cruzodesmus, 57 
Cryptodesmidae, 10, 18 
Cryptotrichus, 66 
Crypturodesmus, 18 
cryptus, Botrydesmus, 47 
Cryptyma, 48 
cuernavacae, Rhysodesmus, 59 
culebrae, Diaporus, 99 
cumbres, Rhysodesmus, 59 
Curodesmus, 37 
curticaudus, Oxypyge, 86 
cyaneus, Strongylodesmus, 40, 41 
Cyclodesmus, 42 
Cyclopsodesmus, 53 
Cyclorhabdus, 15 
Cyclothyrophorus, 79, 81 
Cylindrogonus, 54 
Cyliocyrtus, 19 
Cylionus, 42 
Cyphodesmus, 43 
Cyrtodesmidae, 9, 18 
Cyrtodesmus, 19 


128 U.S. NATIONAL MUSEUM BULLETIN 266 


Dampfaria, 57 
dampfi, Dampfaria, 59 


Mexicoiulus, 75 
Paraiulus, 75 
Rhysodesmus, 59 
Yucodesmus, 53 
Decaporodesmus, 48 
delus, Nabotomus, 50 
dentatus, Cyrtodesmus, 19 
depilis, Cyrtodesmus, 20 
deses, Tylobolus, 93 
Desmacrides, 16 
Desmethus, 106 
diadematus, Catapleuradesmus, 48 
Diaphanacme, 18 
diaporoides, Orthoporus, 97 
Diaporus, 97 
dichrus, Desmacrides, 16 
digitata, Poratia, 51 
digitatus, Scytonotus, 51 
Sphaeriodesmus, 44 
Diploiulus, 73 
Diplopoda, 5 
Dirhabdophallus, 12 
discriminans, Orthoporus, 99 
Scaphiostreptus, 99 
dispersus, Neoleptodesmus, 38 
distinctus, Oxobolus, 84 
divergens, Ogkomus, 69 
diversunguis, Lophoproctinus, 5 
Docodesmiella, 17 
Dominicodesmus, 51 
Dorsoporidae, 9, 20 
Dorsoporus, 20 
dromeus, Alocodesmus, 11 
dubius, Gasatomus, 49 
dugesi, Rhinocricus, 88 
Spirobolus, 88 
Duoporus, 27 
dux, Aceratophallus, 36 
dybasi, Euphallus, 37 
Orthoporus, 99 
Scaphiostreptus, 99 
Dybasia, 69 
Dybasiidae, 66, 69 
Dynesmus, 33 


echinus, Leiogonopus, 55 

Eclomus, 95 

Eclytus, 94 

edenus, Rhinocricus, 88 

Eirenyma, 49 

Elcarmenia, 27 

electa, Hexodontia, 30 

electus, Peridontodesmus, 30 
Rhexiphloeus, 30 
Trachyphloeus, 30 

elegans, Trichomorpha, 16 

elestribus, Rhysodesmus, 59 


elevatus, Pararhachistes, 39 
emarginatus, Watichelus, 81 
emersoni, Gasatomus, 49 
Enantiogonus, 54 
engelhardi, Elcarmenia, 27 
ensiger, Chondrodesmus, 12 
Dirhabdophallus, 12 
Epinannolene, 96 
Epinannolenidae, 96 
Epistreptus, 97 
equalis, Oxypyge, 86 
Ergene, 95 
ergus, Cruzodesmus, 57 
erichsoni, Amplinus, 24 
Orthomorpha, 24 
Polydesmus, 24 
Pseudamplinus, 24 
Ermostyx, 34 
erosus, Hypsiloporus, 18 
erythrotypus, Hiltonius, 93 
esperanzae, Orthoporus, 99 
Rhysodesmus, 59 
Ethophallus, 14 
euliotus, Chondrodesmus, ]2 
eunis, Rhysodesmus, 59 
Euphallus, 37 
Euplinus, 22 
Eurhinocricus, 85 
Euryuridae, 10, 21 
eusculptus, Rhysodesmus, 59 
Eusphaeriodesmus, 43 
euthus, Orthoporus, 99 
eutropis, Kalesmus, 30 
Eutynellus, 54 
Eutyporachis, 14 
eutypus, Amplinus, 24 
Eurhinocricus, 85 
Lignydesmus, 29 
Mesesmus, 29 
Oniscodesmus, 29 
Pseudamplinus, 24 
Rhysodesmus, 59 
evidens, Trichomorpha, 16 
Typophallus, 16 
excisus, Platydesmus, 106 
eximius, Messicobolus, 82 
Spirobolus, 82 
extrema, Trichomorpha, 16 


falcatus, Calymmodesmus, 47 
Centrellus, 79 
falciphallus, Chondrodesmus, 12 
fallens, Siphonophora, 110 
fallens annecta, Siphonophora, 110 
federalis, Hiltonius, 93 
ferrugineus, Rhinocricus, 88 
Spirobolus, 88 
Zeuctodesmus, 28 
ferruginipes, Oxypyge, 91 
Zipyge, 91 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 129 


festae—Continued 

festae, Orthomorpha, 65, 99 
Orthoporus, 99 
Plusioporus, 99 

filicomis, Julus, 75 
Paraiulus, 75 

fimbriatus, Platyrrhacus, 35 
Polydesmus, 35 
Tirodesmus, 35 

fissus, Eurhinocricus, 85 

flagellatus, Peridontodesmus, 31] 

flavicornis, Amplinus, 24 
Orthomorpha, 24 
Pseudamplinus, 24 

flavior, Eutynellus, 54 

flavocinctus, Rhysodesmus, 59 

foetidus, Proletus, 15 

folium, Camptomorpha, 16 
Leptodesmus, 16 
Trichomorpha, 16 

forficula, Isidrona, 15 

formicarius, Myrmecodesmus, 50 

formicatus, Calymmodesmus, 47 

fortinus, Orthoporus, 100 

fossata, Poratia, 51 

fossulifer, Hiltonius, 93 
Spirobolus, 93 

fractus, Telauxus, 52 

fragilis, Enantiogonus, 54 

fraterna, Fontaria, 60 

fraternus, Julus, 100 
Nyssodesmus, 32 
Orthoporus, 100 
Platyrhacus, 32 
Platyrrhacus, 32 
Polydesmus, 60 
Rhysodesmus, 60 
Spirostreptus, 100 

fratrellus, Aztecolus, 93 
Trichomorpha, 16 

frauenfeldianus, Chondrodesmus, 12 
Leptodesmus, 12 
Polydesmus, 12 

frionus, Rhysodesmus, 60 

fulgens, Botrydesmus, 47 
Prionodesmus, 47 

furcata, Salvadoria, 56 

furcatus, Styraxodesmus, 52 

furcifer, Polylepiscus, 22 


galeanae, Pararhachistes, 39 

garcianus, Rhysodesmus, 60 
Toltecolus, 79 

Gasatomus, 49 

geddesi, Strongylodesmus, 41 

geniculatus, Cyclopsodesmus, 53 
Neoleptodesmus, 38 

geophilus, Dominicodesmus, 51 

glaphyros, Aphelidesmus, 22 
Euryurus, 22 


globiceps, Siphonophora, 110 
globulosus, Inodesmus, 18 
Glomerida, 6, 7 
Glomeridae, 7 
Glomeridesmida, 6, 7 
Glomeridesmidae, 7 
Glomeridesmus, 7, 44 
Glomeris, 7 
Glomeroides, 7 
godmani, Cleidogona, 68 
Hirsutogona, 68 
Messicobolus, 82 
Rhysodesmus, 60 
Spirobolus, 82 
egracilior, Orthoporus, 100 
gracilipes, Platyrhacus, 34 
gracilis, Calymmodesmus, 47 
Cylionus, 43 
Fontaria, 65 
Oxidus, 65 
Sphaeriodesmus, 42, 43 
grammicus, Barrobolus, 91 
Microspirobolus, 91 
eranosus, Chondrodesmus, 13 
Cyrtodesmus, 20 
Dirhabdophallus, 13 
Leptodesmus, 13 
Oncodesmus, 20 
Polydesmus, 20 
granulifer, Phorositus, 39 
granulofrons, Poratia, 51 
Psochodesmus, 51 
Treseolobus, 51 
granulosus, Polydesmus, 63 
griseus, Sphaeriodesmus, 44 
guardanus, Rhysodesmus, 60 
guatemalae, Aceratophallus, 36 
Platydesmus, 106 
guatemalana, Solaenogona, 69 
guatemalensis, Curodesmus, 38 
Neoleptodesmus, 38 
Platydesmus, 106 
guerreronus, Gymnostreptus, 100 
Orthoporus, 100 
guineensis, Alloproctus, 5 
guttulatus, Blaniulus, 74 
Julus, 74 
Gymnostreptus, 96 


Habrodesmus, 15 
hagedussii, Rhinocricus, 88 
Spirobolus, 88 
Harpogonopus, 54 
harti, Siphonophora, 109 
hebes, Hiltonius, 93 
Spirobolus, 93 
Tylobolus, 93 
Helminthomorpha, 6. 8 
hermaphroditus, Aphelidesmus, 22 
heteropygus, Cyclothyrophorus, 80 
Spirobolus, 80 


130 U.S. NATIONAL MUSEUM BULLETIN 266 


heterosculptus, Pachyurus, 23 
Polylepiscus, 23 
heterosculptus pococki, 
Polylepiscus, 23 
heterotuberculata, Poratia, 5] 
Hexodontia, 29 
hidalgonus, Cyphodesmus, 43 
Sierresmus, 32 
Hiltonius, 92 
Hirsutogona, 68 
hirsutus, Peridontodesmus, 31 
hirudo, Platydesmus, 106 
hispidulosus, Cyrtodesmus, 20 
hoctunanus, Aceratophallus, 36 
hoctuni-slens, Orthoporus, 100 
hoffmanni, Chondrodesmus, 13 
Dirhabdophallus, 13 
Rhacophorus, 13 
Holistophallidae, 10, 27 
Holistophallus, 27 
Homodesmus, 51 
hondurasanus, Sphaeriodesmus, 44 
hoogstraali, Maderesmus, 31] 
Messicobolus, 82 
Peridontodesmus, 31 
hoplomerus, Messicobolus, 82 
Spirobolus, 82 
Huanucodesmus, 29 
humerosa, Dybasia, 69 
humerosum, Airocaulon, 46 
Hybocestus, 43 
Hypsiloporus, 17 


Idiurodesmus, 49 

Ilyma, 49 

inferus, Lophoproctinus, 5 

Inodesmus, 18 

inquinatus, Calymmodesmus, 47 

insulanus, Lignydesmus, 29 
Mesesmus, 29 

insularis, Docodesmiella, 17 

insulatus, Eurhinocricus, 89 
Rhinocricus, 89 

intermedius, Aphelidesmus, 22 
Polydesmus, 38 
Rhysodesmus, 60 

intermittens, Amplinus, 24 
Pseudamplinus, 24 

interruptus, Platydesmus, 107 

interruptus Simplex, Platydesmus, 107 

inustus, Rhysodesmus, 60 

Irazunus, 54 

Irogonus, 55 

isidricus, Yucodesmus, 53 

Isidrona, 15 

isolatus, Barrodesmus, 28 
Oxypyge, 86 
Sphaeriodesmus, 44 

Isoporostreptus, 97 

isthmianum, Mestosoma, 64 

ixtapanus, Rhinocricus, 88 


jacobsoni, Homodesmus, 51 
Prosopodesmus, 5] 

jamaicensis, Inodesmus, 18 

jonesi, Spirostrephon, 70 

Julida, 8, 73 

Julidae, 73 

Julidea, 73 


Kalesmus, 30 
kalonotus, Pachyurus, 2] 
Kapyrodesmus, 49 
Katantodesmus, 18 
kauanus, Cylionus, 43 
kerrensis, Centrellus, 79 
Eurellus, 79 
kiemi, Orthoporus, 100 
klugi, Amplinus, 24, 26 
Pachyrus, 24 
Polydesmus, 24 
Pseudamplinus, 24 
knighti, Rhysodesmus, 60 
kraussi, Trichomorpha, 16 


laetus, Gymnostreptus, 97 

lagura, Scolopendra, 6 

lamellifer, Aceratophallus, 36 

laminatus, Cynedesmus, 50 
Lophodesmus, 50 

laminiger, Tunodesmus, 28 

lamprus, Rhinocricus, 88 

lankesteri, Platydesmus, 107 

lapidicina, Oxypygides, 86 

Lasiodesmus, 18 

laticollis, Siphonocybe, 109 

latistriatus, Diploiulus, 73 

latus, Diaphanacme, 18 
Glomeridesmus, 8 

latzeli, Amplinus, 21 
Pycnotropis, 21 

lecontei, Brachycybe, 105 

Leiodere, 95 

Leioderidae, 94, 95 

Leiogonopus, 55 

leius, Orthoporus, 100 

lenonus, Orthoporus, 100 

leon, Amplinus, 25 
Pseudamplinus, 25 

leona, Cleidogona, 67 

leonensis, Rhysodesmus, 60 

leonicus, Orthoporus, 101 

leucoclius, Paraiulus, 76 
Thriniulus, 76 

leucus, Prostemmiulus, 71 
Stemmiulus, 71 

Ligiodesmus, 28 

Lignydesmus, 29 

limax, Polydesmus, 58, 60 
Rhysodesmus, 60 

limonensis, Nyssodesmus, 33 
Platyrrhacus, 33 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 131 


linares, Orthoporus, 101 
lineatus, Platydesmus, 107 
lobata, Cryptyma, 48 
lombardiae, Prostemmiulus, 7 1 
longiramus, Sphaeriodesmus, 44 
longitubus, Sphaeriodesmus, 44 
loomisi, Colactis, 70 
Lophodesmus, 50 
Lophoproctidae, 5 
Lophoproctinus, 5 
luchilicolens, Orthoporus, 101 
lusitanus, Brachyiulus, 73 
lutosus, Botrydesmus, 46 
lyrapes, Cyrtodesmus, 20 
Lysiopetalidae, 70 
Lysiopetalidea, 66, 70 

lyttoni, Siphonacme, 109 


macrosetosus, Cyrtodesmus, 20 
maculata, Mexiceuma, 68 
Maderesmus, 30 
magna, Docodesmiella, 17 
magnificus, Oxobolus, 84 
magnus, Amplinus, 21 
Seminellogon, 21 
major, Aphelidesmus, 23 
Polylepiscus, 23 
malinche, Rhysodesmus, 60 
malkini, Rhinocricus, 88 
mandeli, Cleidogona, 67 
manni, Amplinus, 25 
Pseudamplinus, 25 
marci, Rhinocricus, 88 
marcosus, Rhysodesmus, 61 
margarites, Diaporus, 101 
Orthoporus, 101 
marginandus, Stemmiulus, 72 
marginatus, Oniscus, 7 
marmoreus, Platydesmus, 107 
maya, Aceratophallus, 36 
mayanus, Rhinocricus, 88 
Rhysodesmus, 61 
mayus, Polydesmus, 63 
mediator, Ceuthauxus, 41 
Strongylodesmus, 41 
medius, Peridontodesmus, 3] 
Sphaeriodesmus, 44 
mertensi, Rhinocricus, 88 
Mesesmus, 29 
mesites, Oxypygides, 86, 87 
mesomelas, Platydesmus, 107 
messicanum, Tynoma, 70 
Messicobolidae, 77, 81 
Messicobolus, 81, 82 
Mestosoma, 64 
Metaphallus, 37 
Mexicambala, 95 
mexicana, Cleidogona, 67 
Siphonophora, 111 


mexicanum, Craspedosoma, 67 
mexicanus, Calymmodesmus, 47 
Cleidogona, 67 
Cyphodesmus, 43 
Glomeridesmus, 44, 45 
Hiltonius, 93 
Julus, 93 
Lophoproctinus, 5 
Oniscodesmus, 43 
Platydesmus, 107 
Platyrhacus, 33, 34 
Polydesmus, 34 
Prostemmiulus, 71 
Sphaeriodesmus, 44, 45 
Spirobolus, 93 
Stenodesmus, 64 
Mexiceuma, 68 
Mexicoiulus, 74 
Mexiterpes, 69 
michoacanus, Hiltonius, 93 
Sphaeriodesmus, 45 
Tancitares, 4] 
Microrhachis, 39 
Microspirobolus, 91 
micrus, Tichodesmus, 56, 57 
mimeticus, Pheniulus, 76 
mimus, Amplinus, 25 
Metaphallus, 37 
Nyssodesmus, 33 
Orthoporus, 101 
Pseudamplinus, 25 
minor, Acentronus, 6] 
Rhysodesmus, 61 
minusculus, Irazunus, 55 
minutissima, Hirsutogona, 68 
minutus, Nopoiulus, 74 
mirabilis, Hirsutogona, 68 
moderata, Cleidogona, 67 
modestus, Myrmecodesmus, 50 
modicus, Prostemmiulus, 71 
modicus cordovana, 
Prostemmiulus, 71 
montana, Siphonophora, 111 
montanus, Calymmodesmus, 48 
Chondrodesmus, 13 
Dirhabdophallus, 13 
Hypsiloporus, 18 
montezumae, Julus, 101] 
Orthoporus, 101 
Polydesmus, 61 
Rhysodesmus, 61 
Spirostreptus, 101, 102 
monticola, Messicobolus, 82 
Spirobolus, 82 
monticolens, Pinesmus, 31] 
montivagus, Nyssodesmus, 33 
Platyrhacus, 33 
morela, Hexodontia, 30 
Ilyma, 49 
Morelene, 95 


s2 U.S. NATIONAL MUSEUM BULLETIN 266 


moreleti, Piestodesmus, 107 
Platydesmus, 107 

morelus, Ceuthauxus, 40, 41 
Orthoporus, 100 
Peridontodesmus, 30 
Rhinocricus, 89 
Rhysodesmus, 61 
Strongylodesmus, 41 

motulensis, Rhinocricus, 89 

motzoronginis, Decaporodesmus, 48 

mucronatus, Rhinocricus, 89 

mulegensis, Kapyrodesmus, 49,50 
Xerodesmus, 49, 50 

multiloba, Trichodesmiella, 17 

munda, Kirenyma, 49 

mundus, Cyrtodesmus, 20 
Morelene, 95 
Orthoporus, 101 
Oxobolus, 84 

murphyi, Chondrodesmus, 13 

muruztunicus, Yucodesmus, 53 

Myrmecodesmus, 50 

mystecus, Julus, 82 
Messicobolus, 82 
Polydesmus, 61 
Rhysodesmus, 61 
Spirobolus, 82 


Nabotomus, 50 

nahuus, Chelogonobolus, 78 
Polydesmus, 61 
Rhysodesmus, 61 
Spirobolellus, 78 
Spirobolus, 78 

nannus, Chondrodesmus, 13 

naresi, Spirobolus, 94 
Spirostrophus, 94 

nee ects Acutangulus, 37 

aussurobolus, 80 

Sphaeriodesmus, 45 

Nemasomatidae, 73, 74 

Neoleptodesmus, 37 

nesiotes, Orthoporus, 101 

nicaraguae, Chondrodesmus, 14 
Platydesmus, 107 

nicaraguanus, Nyssodesmus, 33 
Orthoporus, 101 
Platydesmus, 107 
Rhinocricus, 89 

nidicola, Trichomorpha, 16 

nietanus, Cyclothyrophorus, 80 
Julus, 80 
Saussurobolus, 80 
Spirobolus, 80 

nigrescens, Atopetholus, 79 
Onychelus, 79 

nigricaudus, Nyssodesmus, 33 

nigrior, Aztecolus, 92 
Spirobolus, 92 


niteus, Amplinus, 25 
Pseudamplinus, 25 
nitidus, Amplinus, 25 
Orthomorpha, 25 
Pachyurus, 25 
Platyrrhacus, 25 
Pseudamplinus, 25 
nodosicollis, Rhinocricus, 89 
nodulosus, Sphaeriodesmus, 45 
Nopoiulus, 74 
notandus, Lophoproctinus, 6 
notostictus, Rhysodesmus, 61 
nueva, Cleidogona, 67 
nueva michoacana, Cleidogona, 67 
nuevus, Ceuthauxus, 41 
Pararhachistes, 4] 
Strongylodesmus, 41 
Nyssodesmus, 32 


obesus, Rhinocricus, 89 

obesus rubicundus, Rhinocricus, 89 

ocraceus, Rhinocricus, 89 

octonodus, Hybocestus, 43, 44 

octosetosus, Cyclopsodesmus, 53 

oculeus, Prostemmiulus, 71 

Ogkomus, 69 

Oligodesmus, 48 

olivaceus, Julus, 89 
Rhinocricus, 89 
Spirobolus, 89 

olmecus, Paraiulus, 74, 75 

oltramarei, Eutyporachis, 14 
Leptodesmus, 14 

omalopyge, Diaporus, 102 
Orthoporus, 102 
Spirostreptus, 102 

omiltemae, Eurhinocricus, 85 
Rhinocricus, 85 

Oncodesmoides, 19 

Oncodesmus, 19 : 

Oniscodesmidae, 9, 28 

oniscus, Sphaeriodesmus, 45 

orientalis, Chondrodesmus, 13 

orizaba, Ilyma, 49 

orizabae, Leptodesmus, 38 
Neoleptodesmus, 38 
Polydesmus, 38 

ornatus, Julus, 102 
Orthoporus, 98, 101, 102 
Pammicrophallus, 27 
Spirobolus, 102 

orphnius, Amplinus, 23, 25 
Pseudamplinus, 25 

Orthichelus, 79 

orthogonus, Tunodesmus, 28 

Orthomorpha, 65 

Orthoporus, 97 

Orthyma, 50 

otomitus, Julus, 102 
Orthoporus, 102 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 133 


otomitus--Continued 
Polydesmus, 61 
Rhysodesmus, 61 
Spirostreptus, 99, 102 

ovatus, Irazunus, 55 

Oxidus, 65 

oxkutzcabus, Aceratophallus, 36 

Oxobolus, 81, 84 

Oxypyge, 86 

Oxypygides, 86 


pablillo, Aztecolus, 92 
Pachygonopus, 56 
pacificus, Epistreptus, 97 
Gymnostreptus, 97 
palicaudatus, Amplinus, 25 
Orthomorpha, 25 
Pachyurus, 25 
Pseudamplinus, 25 
palmensis, Orthoporus, 102 
Spirostreptus, 102 
palmitonus, Ceuthauxus, 41 
Strongylodesmus, 41 
Pammicrophallus, 27 
Panamadesmus, 29 
panamanus, Barroxenus, 6 
Lignydesmus, 29 
Siphonotus, 109 
panamensis, Siphonophora, 111 
panamenus, Chondrodesmus, 13 
panamica, Trichomorpha, 16 
panamicus, Amplinus, 21 
Aphelidesmus, 21 
Dominicodesmus, 51 
Seminellogon, 21 
Paradoxosomatidae, 64 
Paradoxosomatidea, 9, 64 
Paraiulidae, 74 
Paraiulidea, 73 
Paraiulus, 74 
parallelus, Stemmiulus, 72 
Parapomus, 29 
Pararhachistes, 39 
parcus, Spirobolus, 87 
parvior, Glomeridesmus, 8 
Schmidtolus, 78 
parvipes, Rettenmeyeria, 51, 52 
parvissimus, Eurhinocricus, 85 
parvula, Siphonophora, 111] 
parvulus, Allarithmus, 15 
Phylactophallus, 15 
parvus, Homodesmus, 51 
Peridontodesmus, 31 
pasaquinae, Pammicrophallus, 27 
pater, Rhysodesmus, 62 
paxillicauda, Orthoporus, 102 
pedrocola, Rhinocricus, 89 
Peltedo, 19 
Pemptoporus, 97 
penicillatus, Irazunus, 55 


Pentazonia, 6 
perditus, Platydesmus, 107 
peregrinus, Holistophallus, 27 
perfidus, Spirostreptus, 96 
Peridontodesmidae, 10, 29 
Peridontodesmus, 3] 
Peridysodesmus, 44 
perotenus, Rhysodesmus, 62 
perparvus, Cynedesmus, 50 
Lophodesmus, 50 
perpictus, Platydesmus, 108 
phanus, Kalesmus, 30 
Pheniulus, 76 
phenotypus, Pheniulus, 76 
phloibius, Paraiulus, 75 
Phorositus, 39 
Phreatodesmus, 56 
Phylactophallus, 15 
picadoi, Prostemmiulus, 71 
picteti, Polydesmus, 24 
pictus, Oxobolus, 84 
Pammicrophallus, 28 
Piestodesmus, 106 
pilosa, Siphonocybe, 109 
pilsbryi, Messicobolus, 83 
Pinesmus, 3] 
pittieri, Chondrodesmus, 13 
Epinannolene, 96 
Isoporostreptus, 97 
plagiodon, Hybocestus, 44 
plana, Epinannolene, 96 
Platops, 70 
Platydesmida, 105 
Platydesmidae, 105, 106 
Platydesmus, 106 
platyops, Messicobolus, 83 
Spirobolus, 83 
Platyrhacidae, 10, 32 
Platyrhacus, 33 
plesius, Rhinocricus, 90 
pococki, Nyssodesmus, 34 
Orthomorpha, 26 
Platyrhacus, 34 
Pseudamplinus, 26 
poecilus, Polyxenus, 6 
Polydesmida, 8, 9 
Polydesmidae, 10, 35 
Polydesmidea, 9 
polydesmoides, Platydesmus, 
106, 108 
Polydesmus, 35, 40 
Polylepiscus, 22 
Polyxenida, 5 
Polyxenidae, 5, 6 
Polyxenus, 6 
Polyzoniida, 105, 108 
Polyzoniidae, 108, 109 
Poratia, 51 
porcellus, Glomeridesmus, 7 
portoricensis, Siphonophora, 110 


134 U.S. NATIONAL MUSEUM BULLETIN 266 


potosianus, Ceuthauxus, 40, 41 
Rhinocricus, 90 
Rhysodesmus, 62 
Strongylodesmus, 41 

potosina, Ilyma, 49 

potosinus, Pararhachistes, 39 

prehensor, Eusphaeriodesmus, 43 
Sphaeriodesmus, 43 

Prionodesmus, 46 

procerus, Tridontomus, 57 

proclivis, Hypsiloporus, 17, 18 

producens, Orthoporus, 102 

producta, Brachycybe, 105 

progressor, Siphonophora, 111 

projectus, Lignydesmus, 29 

Proletus, 15 

promiscus, Glomeroides, 7 

propia, Cleidogona, 67 

propinquus, Nyssodesmus, 34 
Platyrhacus, 34 

Prosopodesmus, 51 

Prostemmiulus, 71 

protenta, Colactis, 70 

Psammodesmus, 34 

Pselaphognatha, 5 

Pseudamplinus, 23 

pseustes, Siphonacme, 109 
Siphonophora, 109 

Ptyxodesmus, 17 

pueblanus, Paraiulus, 75 

pulchellus, Microspirobolus, 91 

pulchrus, Hiltonius, 92 

pulvillatum, Mestosoma, 64 
Strongylosoma, 64 

pulvillatus, Habrodesmus, 64 

punctifer, Cleidogona, 67 

punctilliger, Orthoporus, 

98, 101, 102 

punctulatus, Nopoiulus, 74 

purojenus, Cruzodesmus, 57 

purulana, Allopocockia, 78 

purulanus, Arolus, 77, 78 

purulicus, Periodontodesmus, 31 

pusillus, Julus, 73 
Ligiodesmus, 28 
Lophodesmus, 50 
Rhysodesmus, 62 

Pycnotropis, 21 

pygmoides, Eurhinocricus, 85 
Rhinocricus, 85 

python, Polydesmus, 32 


quadratus, Aceratophallus, 36 


rafaela, Cleidogona, 67 

rafaelanus, Atylophor, 11 

rasilis, Julus, 75 
Paraiulus, 75 

raui, Messicobolus, 83 


recta, Trichomorpha, 16 

reduncus, Anelus, 82 

reimosieri, Irazunus, 54, 55 
Rhaphonotum, 108 

relictus, Prostemmiulus, 72 

remotus, Psammodesmus, 34 

reniformis, Irogonus, 55 

reptans, Messicobolus, 84 
Spirobolus, 84 

rettenmeyeri, Calymmodesmus, 48 

Rettenmeyeria, 51 

Rhachidomorpha, 39 

Rhachis, 40 

Rhachodesmidae, 11, 35 

Rhachodesmus, 40 

Rhaphonotum 108 

Rhexiphloeus, 29 

Rhinocricidae, 77, 85 

Rhinocricus, 87 

Rhysodesmus, 57 

richardsoni, Anelus, 82 
Spirobolellus, 82 

riparius, Nyssodesmus, 34 
Platyrhacus, 34 

rixi, Rhinocricus, 90 

robusta, Epinannolene, 96 

robustus, Eusphaeriodesmus, 43 
Sphaeriodesmus, 43 

rodriguezi, Chondrodesmus, 13 
Dirhabdophallus, 14 
Leptodesmus, 13 
Orthoporus, 102 
Spirostreptus, 102 

rodriguezi coriaceus, 

Orthoporus, 103 

Spirostreptus, 103 

rogersi, Rhinocricus, 90 

rosanus, Paraiulus, 75 

rotundata, Allopocockia, 78 

rotundatus, Chelogonobolus, 78 
Glomeridesmus, 8 

rubriceps, Oniscodesmus, 29 

rubrimarginis, Rhysodesmus, 62 

rugiceps, Orthoporus, 103 
Scaphiostreptus, 103 

rugiceps reimoseri, Orthoporus, 103 
Scaphiostreptus, 103 

russelli, Mexicambala, 95 


sabachana, Siphonophora, 111] 

sabachanus, Chondrodesmus, 14 

sabinus, Bolivaresmus, 46 
Mexiterpes, 69 

sagittalis, Salvadoria, 56 

Sakophallus, 40 

salleanus, Rhinocricus, 90 

salvadorensis, Neoleptodesmus, 38 

salvadori, Strongylosoma, 64 

Salvadoria, 56 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 135 


salvadoricus, Orthoporus, 103 
Sacphiostreptus, 103 

salvini, Cyclothyrophorus, 79, 80, 81 
Rhysodesmus, 62 

sandersi, Rhysodesmus, 62 

santanus, Oxobolus, 84 

Sarolepis, 52 

saussurei, Sphaeriodesmus, 45 

Saussurobolus, 80 

saxetana, Colactis, 70 

scaber, Polydesmus, 33 

Scaphiostreptus, 97 

scaurus, Cyclopsodesmus, 53 

schachti, Paraiulus, 75 
Thriniulus, 75 

Schistides, 15 

schmidti, Amplinus, 26 
Paraiulus, 75 
Pseudamplinus, 26 

Schmidtolus, 78 

schmitti, Psammodesmus, 34 

scobinatus, Rhinocricus, 90 

Scobinomus, 80 

Scolopendra, 6 

sculptilis, Lignydesmus, 29 
Panamadesmus, 29 

secreta, Cleidogona, 67 

sedecimum, Tynoma, 70 

Seminellogon, 21 

seriatus, Rhysodesmus, 62 

serratus, Amplinus, 26 
Hypsiloporus, 18 
Pseudamplinus, 26 
Scobinomus, 80 
Seminellogon, 21 
Sigmogonotropis, 21 

setifer, Desmethus, 106 

setosus, Ergene, 95 
Pinesmus, 31 

Sierresmus, 32 

Sigmogonotropis, 21 

silvestri, Alocodesmus, 1] 
Leptodesmus, 1] 

simillimus, Polydesmus, 62 
Rhysodesmus, 62 

simplex, Sakophallus, 40 

simulans, Rhinocricus, 87, 90 
Synthodesmus, 28 

simulatus, Auxobolus, 94 
Rhinocricus, 87 

singularis, Chondrodesmus, 14 

sinuatus, Neoleptodesmus, 38 

Siphonacme, 109 

Siphonocybe, 109 

Siphonophora, 110 

Siphonophoridae, 108, 109 

Siphonotus, 109 

smithi, Onychelus, 81 
Rhinocricus, 90 
Rhysodesmus, 62 


socia, Oxypyge, 86 

sodalis, Calymmodesmus, 48 
Solaenogona, 69 
Solaenorhabdus, 15 
Solemia, 69 

solicolens, Orthoporus, 103 


spathulata, Docodesmiella, 17 
spatulatus, Chondrodesmus, 14 
Dirhabdophallus, 14 
speobia, Cambala, 95 
speobius, Eclomus, 95 
Eclytus, 95 
Sphaeriodesmidae, 9, 42 
Sphaeriodesmus, 44 
spiculatus, Ptyxodesmus, 17 
Sarolepis, 52 
Spirobolellidae, 77, 91 
Spirobolida, 8, 76 
Spirobolidae, 77, 92 
Spirobolidea, 76 
Spirostrephon, 70 
Spirostreptida, 9, 94 
Spirostreptidae, 96 
Spirostreptidea, 94, 96 
Spirostrophus, 94 
spukilensis, Yucatobolus, 9] 
stebbinsi, Auxobolus, 94 
Tylobolus, 94 
Stemmiulida, 71 
Stemmiulidae, 71 
Stemmiulus, 72 
Stenodesmus, 64 
stenomerus, Phylactophallus, 15 
stenopterus, Nyssodesmus, 33 
Platyrhacus, 33, 34 
stolli, Cleidogona, 68 
Hirsutogona, 68 
Messicobolus, 83 
Polylepiscus, 22, 23 
Rhinocricus, 90 
Rhysodesmus, 62 
Spirobolus, 83 
storkani, Eurhinocricus, 86 
striatulus, Orthoporus, 103 
Strongylodesmus, 40 
stylifer, Eusphaeriodesmus, 43 
Paraiulus, 75 
Sphaeriodesmus, 43 
Stylodesmidae, 10, 46 
Styraxodesmus, 52 
subovatus, Platydesmus, 108 
sulcata, Epinannolene, 96 
sumichrasti, Neoleptodesmus, 38 
Polydesmus, 37 
sumucus, Aceratophallus, 36 
Synthodesmus, 28 


tabascensis, Rhysodesmus, 62 
tacarcuna, Trichomorpha, 16 
tacubayae, Rhysodesmus, 63 


136 U.S. NATIONAL MUSEUM BULLETIN 266 


tajumulco, Amplinus, 26 
Cleidogona, 68 
Pseudamplinus, 26 

Tancitares, 41 

tancitarus, Eurelus, 93 

tapachulae, Amplinus, 26 
Pseudamplinus, 26 

Taphrodesmus, 46 

tarasca, Polydesmus, 39, 40 
Rhachidomorpha, 40 

Tarascolus, 81 

tarascus, Julus, 75 
Paraiulus, 75 

tardus, Oxobolus, 84 

targioni, Crypturodesmus, 18 

taureus, Platydesmus, 108 

taurinus, Aphelidesmus, 22 

teapensis, Orthoporus, 103 

tehuacanus, Orthoporus, 104 

Teinorhachis, 41 

tejerianus, Rhinocricus, 90 

telana, Siphonophora, 111 

Telauxus, 52 

tenuis, Teinorhachis, 41 

tepaneca, Fontaria, 58 

tepanecus, Julus, 83 
Messicobolus, 83 
Polydesmus, 63 
Rhysodesmus, 63 
Spirobolus, 83 

tepoztlanus, Hiltonius, 92 
Maderesmus, 30 
Rhysodesmus, 63 

teres, Prostemmiulus, 72 

tesselatum, Abacion, 70 

tessellatus, Eutyporhacis, 14 

texense, Abacion, 70 

texensis, Spirostrephon, 70 

texicolens, Orthoporus, 104 
Scaphiostreptus, 104 

Thriniulus, 76 

tiburona, Colactis, 70 

tiburonum, Lysiopetalum, 70 

Tichodesmus, 56 

Tidopterus, 51 

Tiphallus, 42 

Tirodesmus, 35 

tizamensis, Orthoporus, 104 

Toltecolus, 79 

toltecus, Julus, 90 
Polydesmus, 63 
Rhinocricus, 90 
Rhysodesmus, 63 
Spirobolus, 90 

torreonus, Orthoporus, 104 

torreyana, Leiodere, 95 

torreyanus, Phreatodesmus, 56 

totanacus, Polydesmus, 63 
Rhysodesmus, 63 


totonacus, Julus, 91 
Messicobolus, 83 
Rhinocricus, 9] 
Spirobolus, 91 

Trachelacanthe, 21 

Tracheloaspis, 52 

Trachelorachis, 21] 

Trachyphloeus, 29 

triangulifer, Platydesmus, 108 

Trichodesmiella, 17 

Trichomorpha, 16 

Trichopetalidae, 66, 69 

Trichopolydesmidae, 10, 53 

Tridontomidae, 10, 57 

Tridontomus, 57 

trifasciatus, Microspirobolus, 92 

trifidus, Peridysodesmus, 44 

trifini, Siphonophora, 111 

Trigoniulidae, 94 

Trigoniulidea, 76, 94 

Trigonostylus, 19 

trimaculatus, Polylepiscus, 23 

trinus, Augesmus, 46 
Cynedesmus, 46 

triramus, Amplinus, 26 
Colobodesmus, 42 
Orthomorpha, 26 
Pseudamplinus, 26 

tristani, Idiurodesmus, 49 
Nyssodesmus, 33 
Platyrhacus, 33 
Prostemmiulus, 72 
Rhinocricus, 9] 

trisulcatus, Orthoporus, 104 
Spirostreptus, 104 

tuberculifer, Chondrodesmus, 14 

tuberculosum, Rhaphonotum, 108 

tumida, Tracheloaspis, 52 

Tunodesmus, 28 

Tylobolus, 93 

tylopus, Allopocockia, 77, 78 
Spirobolellus, 77, 78 

Tynoma, 70 

Typophallus, 16 

typotopyge, Diaporus, 104 
Orthoporus, 104 
Spirostreptus, 104 

typotopyge palmensis, 
Spirostreptus, 102 

tzendalus, Julus, 83 
Messicobolus, 83 
Spirobolus, 83 


ugmalanus, Orthoporus, 104 
unciferens, Orthoporus, 104 
uncinatus, Habrodesmus, 15 
Polydesmus, 39 
Rhachis, 39 
uncus, Jrazunus, 55 


MILLIPEDS OF MEXICO AND CENTRAL AMERICA 137 


unicolor, Aceratophallus, 35, 36 

unicolor lamellifer, 
Aceratophallus, 36 

unicus, Orthoporus, 104 


vagans, Epistreptus, 97 
Gymnostreptus, 97 
valerii, Nyssodesmus, 33 
Siphonophora, 111 
vallaris, Prostemmiulus, 72 
vallicolens, Orthoporus, 102 
valvanus, Cyclothyrophorus, 80 
Spirobolus, 80 
varicolor, Oxypyge, 86 
varilobatus, Cynedesmus, 46 
velaripes, Irazunus, 55 
velutinus, Polydesmus, 19 
vera, Siphonophora, 111 
veracruzanus, Hiltonius, 93 
vergelanus, Amplinus, 26 
Pseudamplinus, 26 
vermiformis, Neoleptodesmus, 38 
Polydesmus, 38 
vertebratus, Pararhachistes, 39 
verus, Bonetesmus, 111] 
vestitus, Taphrodesmus, 46 
viabilis, Rhysodesmus, 63 
Yucodesmus, 53 
vicinus, Polydesmus, 63 
Rhachidomorpha, 40 
Rhysodesmus, 63 
victorianus, Orthoporus, 104 
viganus, Paraiulus, 76 
Pinesmus, 31] 
villicola, Isoporostreptus, 97 
violacea, Fontaria, 63 
violaceus, Rhysodesmus, 63 
viridis, Polydesmus, 40 
Rhachodesmus, 40 
Strongylodesmus, 41 
virilis, Oxobolus, 84 
volcanicola, Euplinus, 22 
vulcan, Aztecolus, 93 
Rhinocricus, 93 


vulcani, Neoleptodesmus, 38 
vulvanus, Cyclothyrophorus, 80 
Spirobolus, 80 


Watichelus, 81 

wheeleri, Rhinocricus, 85 

wichitanus, Orthoporus, 102 

williamsi, Eurhinocricus, 89 
Rhinocricus, 89 

woodianus, Peridontodesmus, 31] 
Polydesmus, 31 


xelitlus, Amplinus, 26 
Pseudamplinus, 26 
Xenoporus, 52 
Xerodesmus, 49 
Xystodesmidae, 10, 57 


Yucatobolus, 9] 
Yucodesmus, 53 


zacatecus, Arinolus, 80 
Saussurobolus, 80 
zapotecus, Julus, 91 
Polydesmus, 64 
Rhinocricus, 9] 
Rhysodesmus, 64 
Spirobolus, 91 
zempoala, Cleidogona, 68 
zempoalus, Paraiulus, 76 
zendalus, Polydesmus, 64 
Rhysodesmus, 64 
Zeuctodesmus, 28 
zilchi, Apomus, 30 
Hexodontia, 30 
zimapaniensis, Cleidogona, 68 
Zipyge, 91 
zizicolens, Gymnostreptus, 105 
Orthoporus, 105 
zonatus, Messicobolus, 83 
zuniulus, Amplinus, 27 
Pseudamplinus, 27 


U.S. GOVERNMENT PRINTING OFFICE : 1968 O - 273-687 







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