eth10 and these will be described in the next several examples these markers had been assigned to bovine chromosome 5 on the international bovine reference family panel described in barendse et al 1997 with a location about one third of the way down the chromosome using the wagyu family material described in example 1 above dna markers from chromosome 5 were genotyped on the wagyu sire and his offspring the dna marker from chromosome 5 with the best association in the 2 u00d7 2 contingency chi square is eth10 toldo et al 1993 as shown in table 10 m2 and m4 are marbling scores of 2 and 4 respectively allele is the allele of the sire and that the steer inherited alleles are ranked in mobility with the fastest migrating allele 1 allele 2 is 217 bp long and allele 5 is 223 bp long other lengths of alleles are expected at the eth10 dna marker the polymorphic dna marker eth10 showed an association to marbling score in the offspring of the wagyu sire with a probability of less than 0 005 of this occurring by chance and with a probability of less than 0 0001 of this occurring by chance if a dominant mode of inheritance is assumed this.