American biologist and historian of science (1941–2002)
Stephen Jay Gould ( ; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Gould spent most of his career teaching at Harvard University and working at the American Museum of Natural History in New York. In 1996, Gould was hired as the Vincent Astor Visiting Research Professor of Biology at New York University, after which he divided his time teaching between there and Harvard.
Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium developed with Niles Eldredge in 1972. The theory proposes that most evolution is characterized by long periods of evolutionary stability, infrequently punctuated by swift periods of branching speciation. The theory was contrasted against phyletic gradualism, the popular idea that evolutionary change is marked by a pattern of smooth and continuous change in the fossil record.
Most of Gould's empirical research was based on the land snail genera "Poecilozonites" and "Cerion". He also made important contributions to evolutionary developmental biology, receiving broad professional recognition for his book "Ontogeny and Phylogeny". In evolutionary theory he opposed strict selectionism, sociobiology as applied to humans, and evolutionary psychology. He campaigned against creationism and proposed that science and religion should be considered two distinct fields (or "non-overlapping magisteria") whose authorities do not overlap.
Gould was known by the general public mainly for his 300 popular essays in "Natural History" magazine, and his numerous books written for both the specialist and non-specialist.
In April 2000, the US Library of Congress named him a "Living Legend".
Biography.
Stephen Jay Gould was born in Queens, New York, on September 10, 1941. His father Leonard was a court stenographer and a World War II veteran of the United States Navy. His mother Eleanor was an artist, whose parents were Jewish immigrants living and working in the city's Garment District. Gould and his younger brother Peter were raised in Bayside, a middle-class neighborhood in the northeastern section of Queens. He attended elementary school and graduated from Jamaica High School.
When Gould was five years old his father took him to the Hall of Dinosaurs in the American Museum of Natural History, where he first encountered "Tyrannosaurus rex". "I had no idea there were such things—I was awestruck," Gould once recalled. It was in that moment that he decided to become a paleontologist.
Raised in a secular Jewish home, Gould did not formally practice religion and preferred to be called an agnostic. When asked directly if he was an agnostic in "Skeptic" magazine, he responded:
 ;templatestyles src="Template:Blockquote/styles.css" / ;If you absolutely forced me to bet on the existence of a conventional anthropomorphic deity, of course I'd bet no. But, basically, Huxley was right when he said that agnosticism is the only honorable position because we really cannot know. And that's right. I'd be real surprised if there turned out to be a conventional God.
Though he "had been brought up by a Marxist father" he stated that his father's politics were "very different" from his own. In describing his own political views, he has said they "tend to the left of center." According to Gould the most influential political books he read were C. Wright Mills' "The Power Elite" and the political writings of Noam Chomsky.
While attending Antioch College in Yellow Springs, Ohio, in the early 1960s, Gould was active in the civil rights movement and often campaigned for social justice. When he attended the University of Leeds as a visiting undergraduate, he organized weekly demonstrations outside a Bradford dance hall which refused to admit black people. Gould continued these demonstrations until the policy was revoked. Throughout his career and writings, he spoke out against cultural oppression in all its forms, especially what he saw as the pseudoscience used in the service of racism and sexism.
Interspersed throughout his scientific essays for "Natural History" magazine, Gould frequently referred to his non-scientific interests and pastimes. As a boy he collected baseball cards and remained an avid New York Yankees fan throughout his life. As an adult he was fond of science fiction movies, but often lamented their poor storytelling and presentation of science. His other interests included singing baritone in the Boston Cecilia, and he was a great aficionado of Gilbert and Sullivan operas. He collected rare antiquarian books, possessed an enthusiasm for architecture, and delighted in city walks. He often traveled to Europe, and spoke French, German, Russian, and Italian. He sometimes alluded ruefully to his tendency to put on weight.
Marriage and family.
Gould married artist Deborah Lee on October 3, 1965. Gould met Lee while they were students together at Antioch College. They had two sons, Jesse and Ethan, and were married for 30 years. His second marriage in 1995 was to artist and sculptor Rhonda Roland Shearer.
First bout of cancer.
In July 1982 Gould was diagnosed with peritoneal mesothelioma, a deadly form of cancer affecting the abdominal lining (the peritoneum). This cancer is frequently found in people who have ingested or inhaled asbestos fibers, a mineral which was used in the construction of Harvard's Museum of Comparative Zoology. After a difficult two-year recovery, Gould published a column for "Discover" magazine in 1985 titled "The Median Isn't the Message", which discusses his reaction to reading that "mesothelioma is incurable, with a median mortality of only eight months after discovery." In his essay, he describes the actual significance behind this fact, and his relief upon recognizing that statistical averages are useful abstractions, and by themselves do not encompass "our actual world of variation, shadings, and continua."
Gould was also an advocate of medical cannabis. When undergoing his cancer treatments he smoked marijuana to help alleviate the long periods of intense and uncontrollable nausea. According to Gould, the drug had a "most important effect" on his eventual recovery. He later complained that he could not understand how "any humane person would withhold such a beneficial substance from people in such great need simply because others use it for different purposes." On August 5, 1998, Gould's testimony assisted in the successful lawsuit of HIV activist Jim Wakeford, who sued the Government of Canada for the right to cultivate, possess, and use marijuana for medical purposes.
Final illness and death.
In February 2002, a lesion was found on Gould's chest radiograph, and oncologists diagnosed him with stage IV cancer. Gould died 10 weeks later on May 20, 2002, from a metastatic adenocarcinoma of the lung, an aggressive form of cancer which had already spread to his brain, liver, and spleen. This cancer was unrelated to his previous bout of abdominal cancer in 1982, though it is also associated with asbestos exposure. He died in his home "in a bed set up in the library of his SoHo loft, surrounded by his wife Rhonda, his mother Eleanor, and the many books he loved."
Scientific career.
Gould began his higher education at Antioch College, graduating with a double major in geology and philosophy in 1963. During this time, he also studied at the University of Leeds in the United Kingdom. After completing graduate work at Columbia University in 1967 under the guidance of Norman Newell, he was immediately hired by Harvard University, where he worked until the end of his life (1967–2002). In 1973, Harvard promoted him to professor of geology and curator of invertebrate paleontology at the institution's Museum of Comparative Zoology.
In 1982, Harvard awarded him the title of Alexander Agassiz Professor of Zoology. That same year, he received the Golden Plate Award of the American Academy of Achievement. In 1983, he was awarded a fellowship at the American Association for the Advancement of Science, where he later served as president (1999–2001). The AAAS news release cited his "numerous contributions to both scientific progress and the public understanding of science." He also served as president of the Paleontological Society (1985–1986) and of the Society for the Study of Evolution (1990–1991).
In 1989, Gould was elected into the body of the National Academy of Sciences. Through 1996–2002 Gould was Vincent Astor Visiting research professor of biology at New York University. In 2001, the American Humanist Association named him the Humanist of the Year for his lifetime of work. In 2008, he was posthumously awarded the Darwin–Wallace Medal, along with 12 other recipients. (Until 2008, this medal had been awarded every 50 years by the Linnean Society of London.)
Punctuated equilibrium.
Early in his career, Gould and his colleague Niles Eldredge developed the theory of punctuated equilibrium, which describes the rate of speciation in the fossil record as occurring relatively rapidly, which then alternates to a longer period of evolutionary stability. It was Gould who coined the term "punctuated equilibria" though the theory was originally presented by Eldredge in his doctoral dissertation on Devonian trilobites and his article published the previous year on allopatric speciation.
According to Gould, punctuated equilibrium revised a key pillar "in the central logic of Darwinian theory." Some evolutionary biologists have argued that while punctuated equilibrium was "of great interest to biology generally," it merely modified neo-Darwinism in a manner that was fully compatible with what had been known before.
Other biologists emphasize the theoretical novelty of punctuated equilibrium, and argued that evolutionary stasis had been "unexpected by most evolutionary biologists" and "had a major impact on paleontology and evolutionary biology."
Comparisons were made to George Gaylord Simpson's work in "Tempo and Mode in Evolution" (1941), in which he also illustrated relatively sudden changes along evolutionary lines. Simpson describes the paleontological record as being characterized by predominantly gradual change (which he termed horotely), although he also documented examples of slow (bradytely), and rapid (tachytely) rates of evolution. Punctuated equilibrium and phyletic gradualism are not mutually exclusive (as Simpson's work demonstrates), and examples of each have been documented in different lineages. The debate between these two models is often misunderstood by non-scientists, and according to Richard Dawkins has been oversold by the media.
Some critics jokingly referred to the theory of punctuated equilibrium as "evolution by jerks",
which prompted Gould to describe phyletic gradualism as "evolution by creeps."
Evolutionary developmental biology.
Gould made significant contributions to evolutionary developmental biology, especially in his work "Ontogeny and Phylogeny". In this book he emphasized the process of heterochrony, which encompasses two distinct processes: neoteny and terminal additions. Neoteny is the process where ontogeny is slowed down and the organism does not reach the end of its development. Terminal addition is the process by which an organism adds to its development by speeding and shortening earlier stages in the developmental process. Gould's influence in the field of evolutionary developmental biology continues to be seen today in areas such as the evolution of feathers.
Selectionism and sociobiology.
Gould was a champion of biological constraints, internal limitations upon developmental pathways, as well as other non-selectionist forces in evolution. Rather than direct adaptations, he considered many higher functions of the human brain to be the unintended side consequence of natural selection. To describe such co-opted features, he coined the term exaptation with paleontologist Elisabeth Vrba. Gould believed this feature of human mentality undermines an essential premise of human sociobiology and evolutionary psychology.
Against "Sociobiology".
In 1975, Gould's Harvard colleague E. O. Wilson introduced his analysis of animal behavior (including human behavior) based on a sociobiological framework that suggested that many social behaviors have a strong evolutionary basis. In response, Gould, Richard Lewontin, and others from the Boston area wrote the subsequently well-referenced letter to "The New York Review of Books" entitled, "Against 'Sociobiology'". This open letter criticized Wilson's notion of a "deterministic view of human society and human action."
But Gould did not rule out sociobiological explanations for many aspects of animal behavior, and later wrote: "Sociobiologists have broadened their range of selective stories by invoking concepts of inclusive fitness and kin selection to solve (successfully I think) the vexatious problem of altruism—previously the greatest stumbling block to a Darwinian theory of social behavior... Here sociobiology has had and will continue to have success. And here I wish it well. For it represents an extension of basic Darwinism to a realm where it should apply."
Spandrels and the Panglossian paradigm.
With Richard Lewontin, Gould wrote an influential 1979 paper entitled, "The Spandrels of San Marco and the Panglossian Paradigm",
which introduced the architectural term "spandrel" into evolutionary biology. In architecture, a spandrel is a triangular space which exists over the haunches of an arch. Spandrels—more