Soviet biologist (1900–1981)
Nikolay Vladimirovich Timofeev-Ressovsky (; 20 September [O.S. 7 September] 1900 – 28 March 1981) was a Soviet biologist who, in principle, was a senior scientist in Soviet programs of nuclear and, later in biological weapons. He conducted research in radiation genetics, experimental population genetics, and microevolution. His life was highlighted by scientific achievements in the face of severe personal hardship, including his imprisonment and working in secret scientific facilities of Soviet Gulag. 
Timofeev-Ressovsky was a descendant of the old Russian school of scientists, characterised by broad naturalistic views on the world, simultaneously combined with exact analysis of causes and consequences and establishment of elementary phenomena. He widely collaborated with physicists. Known for his influential personality, he was a talented story-teller and teacher. He is the author of the term 'genetic engineering'.
Education.
Nikolaj Vladimirovich Timofeev-Resovski, began his university education from 1916 to 1917 at the Moscow City People's University named after A. L. Shanyavskij. From 1917 to 1922, he studied at the First Moscow State University.
The First World War and the consequences of the Russian Revolution of 1917 interrupted his education for periods of time. At the outbreak of the Russian Civil War, Timofeev-Resovskij was a follower of the anarchist Peter Kropotkin. In 1918, he volunteered to serve in a small anarchist cavalry unit, which was part of the Green army, i.e., they were neither supporters of the Bolshevik Red army nor the White army of General Anton Ivanovich Denikin. Eventually, in 1919, the anarchists joined the Red army. As a private in the 12th Red Army, Timofeev-Resovskij took part in the last battle of the Civil War in the Crimea on the Polish front.
A paper trail of his education was swept away in the chaos of war. Timofeev-Resovskij abandoned his education without a scientific degree; he left formal education with only a gold medal. Even though he eventually became a recognized world-class scientist, he would later, as a requirement for employment, be required to complete his doctorate. He defended his doctoral dissertation in 1963 in Sverdlovsk, and he was awarded a Doctor of Science diploma in 1964.
Career.
In Russia.
During the period 1920 to 1925, Timofeev-Resovskij was an instructor and researcher in Moscow. It was in this period that he met and married Elena Aleksandrovana Fidler (1898 – 1973), who was a student of the cytologist and biologist Nikolaj Konstantinovich Kol’tsov in Moscow; Elena became not only Timofeev-Resovskij's lifelong devoted companion but also his lifelong partner in research. Their eldest son, Dmitrij, was born on 11 September 1923.
From 1920 to 1925, Timofeev-Resovskij was a biology instructor in the pre-Chekist district primary school in Moscow. From 1922 to 1925, he was a zoology instructor on the biotechnical faculty of the Moscow Practical (Applied) Institute. From 1921 to 1925, he did research at the Institute of Experimental Biology, of the "Gosudarstvennyj Institut Narodnogo Zdravookhraneniya" (GINZ, State Institute of Public Health), under the directorship of N. K. Kol’tsov. Some of his research at the Institute involved Drosophila. It was in a department headed by Sergei Sergeevich Chetverikov, the founder of population genetics and a colleague of Kol’tsov, that Timofeev-Resovskij started his genetics experiments. From 1924 to 1925, he was a research assistant in Kol’tsov's department of zoology at the Medical Pedagogical Institute in Moscow. In 1924, Timofeev-Resovskij began his publications in the field of phenogenetics.
During the period 1923 to 1925, Oskar Vogt, an eminent psychiatrist and neurophysiologist, was a visiting scientist in Moscow; he was director of the "Kaiser-Wilhelm Gesellschaft’s" "Institut für Hirnforschung" (KWIH, Kaiser Wilhelm Institute for Brain Research) in Berlin. In part, he was looking to recruit researchers for the KWIH, especially in the relatively new field of genetics. In 1924, the Soviet government made an exchange agreement with Germany. The KWIH was invited to establish a laboratory for brain research in Moscow, which it did: "Институт Мозга" (Institut Mozga, Institute of the Brain). To reciprocate, the Soviets promised to help set up an experimental laboratory for genetics at the KWIH. Vogt had good relations with Nikolaj Aleksandrovich Semashko, People's Commissioner (Minister) of Health. It was Semashko and Kol’tsov who suggested Timofeev-Resovskij to Vogt. Thus, in the summer of 1925, Timofeev-Resovskij, his wife Elena and his colleague Sergei Romanovich Tsarapkin (Zarapkin), left Russia for an unspecified period of time in Germany, which lasted 20 years.
In Germany (1925-1945).
Once at the "Kaiser-Wilhelm-Institut für Hirnforschung" (KWIH "Kaiser Wilhelm Institute for Brain Research" ), Vogt and Timofeev-Resovskij began implementing the other half of the scientific agreement with Russia. In 1929 Timofeev-Resovskij became director of the "Abteilung für Experimentelle Genetik" (Department of Experimental Genetics). In 1930, the KWIH moved to its new facility in Berlin-Buch, which was partly financed by a grant from the Rockefeller Foundation. The KWIH, with its departments of Neuroanatomy (Oskar and Cécile Vogt), neurohistology (Alois Kronmüller), Neurochemistry (Marthe Vogt), and genetics (Nikolaij and Elena Timofeev-Resovskij), and its hospital, the KWIH was one of the largest and most modern research facilities of its kind in the entire world. The research conducted in the genetics department, along with the permanent seminar on genetics organized by Timofeev-Resovskij, drew many young postdoctoral researchers who would ultimately become prominent scientists and the department gained the status of an institute; while Timofeev-Resovskij remained a citizen of the Soviet Union for his entire stay in Germany, his status and that of his department were recognized by the fact that he was appointed, in 1938, to the position of scientific member of the "Kaiser-Wilhelm Gesellschaft". Timofeev-Resovskij's colleagues in his department included his wife Elena Aleksandrovana Timofeeva-Resovskaja, his Russian colleague Sergei Tsarapkin, the physician and radiation biologist Alexander Catsch (Katsch), the radiochemist Hans-Joachim Born, and the physicist and radiation biologist Karl Zimmer; a technical assistant, Natasha (Natalie) Kromm, was from Russia. Timofeev-Resovskij also collaborated with other eminent scientists including the nuclear chemist Nikolaus Riehl, who was the scientific director of the industrial corporation "Auergesellschaft" and the biophysicist Max Delbrück, who had studied physics under Niels Bohr and Max Born. Others drawn to Timofeev-Resovskij's department who would go on to become prominent scientists included Hans Bauer and Andriano Buzzati-Traverso. Through 20 years of collaboration with these and many others, Timofeev-Resovskij authored or coauthored over 100 papers written in German, English, Russian, and French. In addition to those already mentioned, Timofeev-Resovskij's scientific circle included the physicists William Astbury, Niels Bohr, P. A. M. Dirac, Pascual Jordan, and Erwin Schrödinger, and the biologists Torbjörn Caspersson, C. D. Darlington, Theodosij Dobzhans’skij, Boris Ephrussi, Åke Gustafsson, J. B. S. Haldane, Hermann Joseph Muller, Nikolai Vavilov, and Vladimir Vernadsky.
Timofeev-Resovskij's second son was born on 9 April 1927.
Together with the French geneticist of Russian origin, Boris Ephrussi, and with the generous support of the Rockefeller Foundation, Timofeev-Resovskij organized an annual conference on biophysics, genetics, and radiation biology. This even continued right up to the start of World War II in 1939.
In 1932, Timofeev-Resovskij attended the 6th International Congress of Genetics, which was held in Ithaca, New York. While there, he formed a friendly relationship with the plant geneticist Nikolaj Vavilov, who was President of the Soviet Adcademy of Agricultural Sciences.
Timofeev-Resovskij stayed in Germany even after Adolf Hitler came to power in 1933. That the Gestapo or the "Schutzstaffel" (SS) did not give him trouble as a citizen of the Soviet Union is all the more remarkable since Vogt was harassed by the National Socialists for reasons which could have translated into trouble for Timofeev-Resovskij. On the evening of 15–16 March 1933, members of the "Sturmabteilung" (Storm Troopers) raided Vogt's villa on the grounds of the KWIH. The action had been taken in response to a denunciation by M. H. Fischer, a physiologist at the Institute, who was seeking to rise in the "Nationalsozialistische Deutsche Arbeiterpartei" (NSDAP, National Socialist Workers Party). The accusations against Vogt included making payments to the Communist Party, maintaining connections to Russia, tolerating Communist staff members, and dismissing National Socialist staff members. 
In 1935, Timofeev-Resovskij published the major work, "Über die Natur der Genmutation und der Genstruktur", with Karl Zimmer, and Max Delbrück; it was considered to be a major advance in understanding the nature of gene mutation and gene structure. The work was a keystone in the formation of molecular genetics. It was also an inspirational starting point for Erwin Schrödinger's thinking, a course of lectures in 1943, and the eventual writing of the book "What Is Life? The Physical Aspect of the Living Cell".
In 1937, Timofeev-Resovskij and Tsarapkin were ordered back to Russia by the Soviet government. Going back to Russia was deemed impossible, as they would have to abandon their equipment and work in progress, as well as their students. Also, once back in Russia and under the political distortions of science and agriculture due to Lysenkoism, they would, at best, have to disparage the work they had accomplished in the preceding 15 years. At the worst, they could be subject to the same fate as many geneticists under Lysenkoism who had been arrested, sent to the Gulag and eventually died or were executed. In 1929, for example, Timofeev-Resovskij's former teacher Sergei Chetverikov had been arrested and exiled. During the period 1936 to 1940, many prominent geneticists, including Vavilov, Georgi Dmitrievich Karpechenko, and Solomon G. Levit, were arrested. Many who were arrested later died in the GULAG or were executed. Timofeev-Resovskij’s younger brothers were arrested; the one in Leningrad was executed, but the one in Irkutsk survived. Many of Elena’s relatives were also arrested and perished. Others, such as Nikolai Koltsov and Boris Astaurov, were dismissed from their positions. In this light, they decided not to return to the USSR.
Also in 1937, while Oskar Vogt had been director of the KWIH since 1919, he was dismissed from his position by the Nazis; he then became director of the "Institut für Hirnforschung und allgemeine Biologie" (Institute of Brain Research and General Biology) in Neustadt/Schwarzwald. That same year, the Rockefeller Foundation extended an invitation to Timofeev-Resovskij to become head of the Cold Spring Harbor Laboratory of the Carnegie Institution for Science. Timofeev-Resovskij declined the offer, but he had used it to negotiate improved terms for himself and his department at the KWIH.
Timofeev-Resovskij's eldest son Dmitrij, a student at the Humboldt University of Berlin, was arrested in the spring of 1943 for being a member of the Berlin Committee of the All-Union Communist Party (Bolsheviks) and making contact with Russian prisoners of war during World War II. He was sent to the German concentration camp in Mauthausen and executed by the Gestapo on 1 May 1945.
Starting in 1944, in order to minimize the possibility of casualties due to Allied air raids and to also avoid falling into the hands of the Russians, all departments at the KWIH, except for Timofeev-Resovskij's were evacuated from Berlin-Buch to Dillenburg, and later on to Giessen and Göttingen. Timofeev-Resovskij chose to remain in Berlin and await the Russians. He, as well as other German scientists, speculated that the Russian need for scientists would make collaboration with them better than with the Americans, who had far less of a need.
In the USSR again.
The Battle of Berlin was one of the last major engagements of World War II. With a great majority of German scientific facilities in Berlin and its suburbs, this area was a major target of search teams (see Russian Alsos) sent into Germany by Russia to requisition equipment, materiel, and scientific personnel to aid the Soviet atomic bomb project. Haste was necessary, as the American military forces were rapidly approaching Berlin. Soviet troops broke the Berlin defense ring on 25 April 1945, and the Soviet Union announced the fall of Berlin on 2 May. The main search team, headed by Colonel General A. P. Zavenyagin, arrived in Berlin on 3 May; it included Colonel General V. A. Makhnjov, and nuclear physicists Yulij Borisovich Khariton, Isaak Konstantinovich Kikoin, and Lev Andreevich Artsimovich. Georgij Nikolaevich Flerov had arrived earlier, although Kikoin did not recall a vanguard group. Even the scientists were attired in the uniforms of NKVD officers. Targets on the top of their list were the "Kaiser-Wilhelm Institut für" Physik (KWIP, Kaiser Wilhelm Institute of Physics), the University of Berlin, and the "Technische Hochschule Berlin".
A search team soon rounded up Timofeev-Resovskij's colleague Nikolaus Riehl. In early July, Riehl was flown to Russia to head up a group at Plant No. 12 in Ehlektrostal’ (Электросталь) for the production of metallic uranium within