Genus of cartilaginous fishes
Acanthodes (from , 'provided with spines') is an extinct genus of acanthodian fish. Species have been found in Europe, North America, and Asia, spanning the Early Carboniferous to the Early Permian, making it one of the youngest known acanthodian genera.
Description.
The largest species of "Acanthodes" like "Acanthodes confusus" and "Acanthodes splendidus" grew to lengths of at least , while some species like "Acanthodes ultimus" were much smaller, reaching a total body length of only . The body was elongate and had a pair of pectoral fins, an unpaired dorsal fin far back on the body, with an unpaired long ventral/pelvic fin and an anal fin on the underside of the body, which like other acanthodians were supported by stiff spines at their front edges. The whole body was covered in scales, which varied in shape depending on their position. The vertebral column was typically unossified. Species of "Acanthodes" differ from each other in the degree of body elongation and the size and shape of the fins. "Acanthodes" had no teeth and had long gill rakers. Because of this, "Acanthodes" is presumed to have been a suspension feeder, filtering plankton from the water. A specimen of "Acanthodes bridgei "was so well-preserved that traces of its eye tissue were sufficient to establish that "Acanthodes" had both rod and cone photoreceptor cells, suggesting that it was capable of color vision.
Ecology.
The various species of "Acanthodes" are known to have inhabited freshwater lakes, as well as saline lagoons. "Acanthodes bronni," which lived in freshwater lakes in southern Germany during the Early Permian, is known to have been fed upon by the temnospondyl amphibians "Archegosaurus" and "Glanochthon". "Acanthodes" was likely capable of opening its jaws wide as an adaptation to suspension feeding, with the jaws possibly also rotating outwards during opening in order to increase gape.
Taxonomy.
The classification of acanthodians was historically contentious, however, in the 2010s based in part based on detailed studies of the skull of "Acanthodes", it became widely accepted that acanthodians represented a paraphyletic assemblage of stem-group Chondrichthyes. Within the "Acanthodii", "Acanthodes" is traditionally placed within the Acanthodiformes, which is now also considered to be paraphyletic.
Species.
After Beznosov, 2009 and Heidtke, 2011
Indeterminate or dubious species.
"Acanthodes tholeyi" Heidtke, 1990 from Early Permian, Germany is later assigned to genus "Westrichus".
References.
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