Family of amphibians
The European spadefoot toads are a family of frogs, the Pelobatidae, with only one extant genus "Pelobates", containing six species. They are native to Europe, the Mediterranean, northwestern Africa, and western Asia.
Description.
The European spadefoot toad grows up to in length and is often inconspicuously coloured. They have squat bodies with smooth skin and eyes with vertical pupils. They are predominantly fossorial (burrowing) frogs, which dig into sandy soils. Pelobatidae frogs burrow backwards and they spend much of their time in the ground. They prefer open areas with loose soil as opposed to dense compact soil to facilitate the burrowing and have hardened protrusions on their feet to aid in digging, which is the source of the common name. They emerge from the ground during periods of rain and breed in pools, which are usually temporary.
All of the species from this family have free-living, aquatic tadpoles. The eggs are laid in temporary ponds that may quickly evaporate, so the tadpole stage is unusually brief, with rapid development to the adult form in as little as two weeks. To further speed their growth, some of the tadpoles are cannibalistic, eating their brood-mates to increase their supply of protein.
Taxonomy.
The seven species of American spadefoot toads (genera "Scaphiopus" and "Spea") were previously also included in the family Pelobatidae, but are now generally regarded as the separate family Scaphiopodidae.
Family Pelobatidae
Fossils.
The earliest fossil genus of pelobatids, "Elkobatrachus", was described in 2006.
In the Jurassic Morrison Formation, pelobatids are represented by the ilium of an unnamed but indeterminate species. This ilium is larger than that of "Enneabatrachus", a contemporary discoglossid species. A specimen has been recovered from Quarry 9 of Como Bluff in Wyoming. Pelobatids are present in stratigraphic zones 5 and 6 of the formation.
The Oligocene site of Enspel in Germany preserves evidence of pelobatid tadpoles feeding on pollen.
Notes.
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