وكان ليون أول من وصف جيبون بورنيو الأبيض ذو اللحية (" Hylobates albibarbis '), the Gansu pika ("Ochotona cansus"), في النيص سومطرة ("Hystrix sumatrae"), the shadowy broad-nosed bat ("Platyrrhinus umbratus"), and two species of slow loris from Borneo, "Nycticebus bancanus" and "Nycticebus borneanus".
Lyon was the first to describe the Bornean white-bearded gibbon ("Hylobates albibarbis"), the Gansu pika ("Ochotona cansus"), the Sumatran porcupine ("Hystrix sumatrae"), the shadowy broad-nosed bat ("Platyrrhinus umbratus"), and two species of slow loris from Borneo, "Nycticebus bancanus" and "Nycticebus borneanus".