 Spiny neurons are of importance because of their "pivotal role in motor control, habit formation, and motivated behavior".They receive synaptic inputs from mossy fibers boutons, as well as multiple other synaptic terminals.The majority of axon collaterals of these neurons remain in the stratum lucidum and CA3 region, though in some cases axon collaterals of these neurons are observed migrating into the CA1 region of the hippocampus.