 Similar to spiny neurons, aspiny neuron dendrites "branch extensively in stratum lucidum and stratum radiatum of CA3", in contrast to spiny neurons, however, some dendrites "traversed stratum pyramidale and entered stratum oriens", the second deepest layer of the hippocampus.Additionally, what distinguishes aspiny neurons form spiny neurons is their higher maximal firing rates and narrower action potential half-widths than their spiny counterparts.In the hippocampus, the stratum lucidum contains many neurons that act locally in local pathways.