 It was preceded by the Pragian stage and followed by the Eifelian stage.It is named after the Ems river in Germany.During this period, earliest known agoniatitid ammonoid fossils began appearing within this stage after first appearing in previous stage and began to evolutionarily radiate within this stage, in which a new ammonoid order Goniatitida rises in the end of Zlichovian stage (Siberian representation; corresponds to early Eifelian and after the end of Early Devonian, before 391.9 mya).